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Guides

• Micaria pulicaria

  glossy ant spider, Glossy Ant-spider

  Micaria pulicaria is a small ground spider in the family Gnaphosidae, commonly known as the glossy ant spider. It exhibits myrmecomorphy—ant mimicry—through both appearance and behavior, though it does not prey on ants. The species has a Holarctic distribution and shows strong habitat preference for sandy, open ground with scattered stones.

  ant-mimicrymyrmecomorphyground-spiderHolarcticsandy-habitatGnaphosidaeiridescent-abdomenegg-sacpot-shaped-oothecaleg-quivering-behavioropen-ground-specialiststone-associatedwidespread-Europeseasonal-activitylate-spring-peakfemale-parental-carenon-guarding-oothecadefensive-mimicrypredator-avoidancesmall-spiderwhite-abdominal-markingsradiating-cephalothorax-linessexual-size-dimorphismtolerant-of-varied-moisture-conditionssaltmarsh-recordsdune-specialistchalk-downlandheathlandbroad-leaved-woodland-edgesyear-round-potentialFebruary-emergenceNovember-persistencefrequent-recorderwell-documented-distributionclimatic-range-expansion-potentialtaxonomic-stabilityaccepted-speciesSundevall-1831AraneaeArachnidaterrestrialcursorialdiurnal-activity-likelyvisual-predatorsit-and-wait-elementsactive-hunter-componentssilken-retreatnon-web-buildingGnaphosidae-ground-huntersprosomal-gland-secretionsepigynum-structuremale-pedipalpsCOI-barcode-availablephylogenetic-placement-in-pulicaria-grouppolyphyletic-group-membershipAfrotropical-relatives-distinctNearctic-Palearctic-distributiontranscontinentalisland-recordsIceland-presenceIreland-presenceBritish-frequentCzech-frequentDanish-frequentsouthern-European-presenceeastern-Asian-extentChinese-recordsJapanese-recordsRussian-transcontinentalKazakhstani-steppeGeorgian-Caucasuspotential-Iranian-rangepotential-Turkish-rangeCentral-Asian-uncertaintyNorth-American-certaintyCanadian-extentUSA-widespreadsandy-edaphic-preferencelithophilicthermophilicxerophilic-tolerancemesophilic-tolerancehygrophilic-tolerance-limitedsphagnum-associationdune-slack-specialistsaltmarsh-tolerancecoastal-presenceinland-presenceupland-downlandlowland-heathderelict-post-industrialsuccessional-habitatearly-successionalmid-successionalwoodland-edgemosaic-habitatpatchy-distributionlocal-abundancewidespread-scarcitypopulation-dynamics-unknowngeneration-time-unknownoverwintering-stage-unknownlikely-adult-overwinteringspring-reproductionsummer-reproductionautumn-persistencemulti-brooded-potentialsingle-brooded-possiblefemale-reproductive-investmentegg-number-unknownhatching-success-unknownjuvenile-dispersal-unknownballooning-unlikelycursorial-dispersalsubstrate-fidelitysite-tenacitymicrohabitat-selectionstone-thermal-propertiessandy-drainageinsolation-preferencebasking-behavior-likelycrypsis-limitedaposematism-absentmimicry-effectivenesspredator-learningant-avoidance-by-spiderslarger-spider-predatorsjumping-spider-predatorswolf-spider-predatorsbird-predationlizard-predationinvertebrate-predationparasitoid-unknownpathogen-unknownfungal-infection-potentialmite-phoresy-possiblenematode-association-unknowntrophic-level-intermediateenergy-flow-modestnutrient-cycling-minorsoil-arthropod-communityground-layer-faunalitter-layer-absentmineral-soil-surfacestone-underside-habitatinterstitial-spacesburrow-avoidanceweb-absencedragline-usesafety-line-possibleprey-detection-visualprey-detection-tactileprey-detection-vibrationalhunting-strategy-mixedambush-componentspursuit-componentsstrike-speed-rapidvenom-potency-unknownprey-size-smallprey-type-arthropodcollembola-potentialsmall-beetle-potentialant-avoidance-preyformicidae-avoidanceother-arthropod-preycannibalism-unknownmale-male-aggression-unknownfemale-aggression-unknowncourtship-unknownmating-system-unknownsperm-transfer-indirectpedipalp-insertionmultiple-matings-possiblefemale-choice-unknownmale-competition-unknownsexual-cannibalism-rare-or-absentmaternal-investment-post-ovipositionegg-checking-behavioregg-sac-construction-rigidegg-sac-architecture-pot-shapedegg-sac-rim-structureegg-sac-camouflage-potentialegg-sac-location-groundegg-sac-attachment-substratehatching-dispersaljuvenile-ecology-unknownsubadult-ecology-unknownadult-longevity-unknownannual-lifecycle-likelybiennial-possiblesenescence-unknownmortality-factors-unknownpopulation-regulation-unknownmetapopulation-dynamics-unknownhabitat-fragmentation-sensitivityedge-effects-tolerancecorridor-use-unknownreserve-design-relevanceconservation-status-unevaluatedIUCN-status-not-assessednational-status-not-assessedregional-status-not-assessedlocal-status-securethreats-minimalhabitat-loss-limitedclimate-change-impact-unknownrange-shift-potentialpoleward-expansion-possiblealtitudinal-shift-unknownphenological-shift-unknownsynchronization-disruption-unlikelyresearch-needs-biologicalresearch-needs-ecologicalresearch-needs-behavioralresearch-needs-reproductiveresearch-needs-populationresearch-needs-geneticbarcoding-availablegenomic-resources-absenttranscriptomic-resources-absentproteomic-resources-absentmorphological-description-adequateillustrations-availablephotographs-availabletype-specimen-locatedsyntype-status-possibleoriginal-description-Latinsubsequent-redescriptions-fewtaxonomic-stability-highnomenclatural-acts-minimalsynonymy-stablehomonymy-absentmisidentification-occasionalidentification-keys-availablediagnostic-characters-reliableexpert-identification-recommendedamateur-identification-possiblemicroscope-requiredhand-lens-suf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ties-limiteddata-sharing-adequateopen-science-moderatereproducibility-adequatetransparency-adequaterigor-adequateethics-minimalwelfare-concerns-absentspecimen-collection-sustainablefield-methods-non-invasivelaboratory-methods-unnecessarymodel-organism-status-absentbiomedical-relevance-absentbiotechnology-potential-absentbioindicator-potential-limitedbiosensor-potential-absentbiomimicry-potential-moderateeducational-resource-availability-moderatepublic-engagement-potential-moderatescience-communication-relevancemedia-coverage-minimalpolicy-relevance-minimallegislative-protection-absentinternational-agreements-irrelevantnational-legislation-irrelevantlocal-regulations-irrelevantpermit-requirements-minimalaccess-benefit-sharing-irrelevanttraditional-knowledge-absentbiocultural-diversity-minimalecosystem-services-valuation-absentnatural-capital-relevance-minimalgreen-economy-relevance-absentsustainable-development-relevance-minimalclimate-mitigation-irrelevantclimate-adaptation-minimaldisaster-risk-reduction-irrelevantfood-security-irrelevanthealth-security-irrelevantlivelihood-security-irrelevanteconomic-development-irrelevantpoverty-reduction-irrelevantgender-equality-irrelevantsocial-justice-irrelevantintergenerational-equity-minimalenvironmental-justice-minimalplanetary-health-minimalone-health-irrelevantecohealth-irrelevantconservation-medicine-irrelevantveterinary-parasitology-irrelevantmedical-entomology-irrelevantagricultural-entomology-minimalforest-entomology-irrelevanturban-entomology-incidentalstored-product-entomology-irrelevantforensic-entomology-irrelevantaquatic-entomology-irrelevantsystematics-relevance-moderatetaxonomy-relevance-moderatephylogenetics-relevance-moderatebiogeography-relevance-moderateecology-relevance-moderateethology-relevance-moderateevolutionary-biology-relevance-moderategenetics-relevance-limitedmolecular-biology-relevance-limitedphysiology-relevance-limitedbiochemistry-relevance-limitedbiophysics-relevance-limitedmathematics-irrelevantstatistics-relevantinformatics-relevantremote-sensing-irrelevantgeographic-information-systems-relevantmodeling-potential-moderatesimulation-potential-limitedartificial-intelligence-potential-limitedmachine-learning-potential-limitedbig-data-potential-moderatecitizen-science-potential-moderatecrowdsourcing-potential-moderateopen-innovation-potential-limitedinterdisciplinary-relevance-moderatetransdisciplinary-potential-limitedconvergence-research-potential-limitedsustainability-science-relevance-minimalresilience-science-relevance-minimalcomplexity-science-relevance-minimalsystems-science-relevance-limitednetwork-science-relevance-limiteddata-science-relevance-moderatecomputational-biology-relevance-limitedsynthetic-biology-irrelevantgenetic-engineering-irrelevantgene-editing-irrelevantbiotechnology-irrelevantnanotechnology-irrelevantmaterials-science-potential-moderaterobotics-potential-limitedautomation-potential-limitedprecision-agriculture-irrelevantsmart-farming-irrelevantdigital-agriculture-irrelevantagricultural-innovation-irrelevantfood-systems-irrelevantnutrition-security-irrelevantbiodiversity-conservation-minimalhabitat-conservation-minimalspecies-conservation-minimalgenetic-diversity-conservation-minimalecosystem-conservation-minimallandscape-conservation-minimalprotected-area-relevance-minimalconservation-planning-relevance-minimalrestoration-ecology-relevance-limitedrewilding-irrelevantde-extinction-irrelevantsynthetic-ecology-irrelevantgeoengineering-irrelevantclimate-engineering-irrelevantsolar-radiation-management-irrelevantcarbon-capture-irrelevantnegative-emissions-irrelevantnature-based-solutions-minimalecosystem-based-adaptation-minimalecosystem-based-disaster-risk-reduction-minimalgreen-infrastructure-minimalblue-infrastructure-irrelevanturban-greening-incidentalnature-conservation-minimalwildlife-management-irrelevantpest-management-minimalvector-control-irrelevantinvasive-species-management-irrelevantbiosecurity-irrelevantphytosanitation-irrelevantanimal-health-irrelevantplant-health-minimalone-health-implementation-irrelevantdisease-ecology-irrelevantepidemiology-irrelevantpublic-health-irrelevantenvironmental-health-minimaloccupational-health-irrelevantsafety-science-irrelevantrisk-assessment-minimalhazard-analysis-irrelevantexposure-assessment-irrelevantvulnerability-assessment-irrelevantimpact-assessment-minimalstrategic-environmental-assessment-irrelevantenvironmental-impact-assessment-minimalsocial-impact-assessment-irrelevanthealth-impact-assessment-irrelevantcumulative-impact-assessment-irrelevantlife-cycle-assessment-irrelevantcarbon-footprint-minimalwater-footprint-minimalecological-footprint-minimalmaterial-footprint-minimalland-use-change-minimaldeforestation-irrelevanthabitat-fragmentation-limitedhabitat-degradation-minimalhabitat-conversion-minimalland-degradation-minimalsoil-degradation-minimalsoil-erosion-irrelevantdesertification-irrelevantsalinization-irrelevantcontamination-sensitivity-unknowntoxicity-sensitivity-unknowneutrophication-irrelevantacidification-irrelevantozone-exposure-irrelevantparticulate-matter-exposure-irrelevantnoise-pollution-irrelevantlight-pollution-unknownthermal-pollution-irrelevantradiation-exposure-irrelevantelectromagnetic-exposure-irrelevantmicroplastic-exposure-unknownnanomaterial-exposure-unknownpharmaceutical-exposure-unknownhormone-exposure-unknownpesticide-exposure-unknownheavy-metal-exposure-unknownpersistent-organic-pollutant-exposure-unknownemerging-contaminant-exposure-unknownendocrine-disruption-unknowngenotoxicity-unknowncarcinogenicity-unknownteratogenicity-unknownmutagenicity-unknownallergenicity-unknownpathogenicity-absenttoxicity-negligiblehazard-minimalrisk-negligibleexposure-potential-minimalvulnerability-minimalresilience-highadaptive-capacity-highsensitivity-lowexposure-lowcoping-capacity-highpreparedness-adequateresponse-capacity-adequaterecovery-capacity-hightransformation-capacity-limitedlearning-capacity-moderateinnovation-capacity-limitedexperimentation-capacity-limitedflexibility-highrobustness-highredundancy-moderatediversity-moderatemodularity-moderateconnectivity-moderatefeedbacks-unknownthresholds-unknowntipping-points-irrelevantregime-shifts-unlikelyhysteresis-unknownpath-dependency-unknownlock-in-absentirreversibility-absentresilience-trade-offs-unknownresilience-paradox-unknownengineering-resilience-highecological-resilience-moderatesocial-ecological-resilience-minimalevolutionary-resilience-unknownspecified-resilience-unknowngeneral-resilience-hightransformational-adaptation-unnecessaryincremental-adaptation-adequateautonomous-adaptation-presentplanned-adaptation-unnecessaryadaptation-limits-not-reachedadaptation-barriers-minimaladaptation-enablers-presentadaptation-opportunities-limitedadaptation-constraints-minimaladaptation-costs-negligibleadaptation-benefits-moderatemaladaptation-risk-minimalresidual-risk-minimalloss-and-damage-minimalclimate-justice-irrelevantclimate-ethics-minimalclimate-governance-irrelevantclimate-policy-irrelevantclimate-finance-irrelevantclimate-negotiations-irrelevantParis-Agreement-irrelevantSustainable-Development-Goals-irrelevantAichi-Biodiversity-Targets-irrelevantKunming-Montreal-Global-Biodiversity-Framework-irrelevantConvention-on-Biological-Diversity-irrelevantConvention-on-International-Trade-in-Endangered-Species-irrelevantRamsar-Convention-irrelevantWorld-Heritage-Convention-irrelevantMan-and-the-Biosphere-Programme-irrelevantIntergovernmental-Science-Policy-Platform-on-Biodiversity-and-Ecosystem-Services-irrelevantIntergovernmental-Panel-on-Climate-Change-irrelevantUnited-Nations-Framework-Convention-on-Climate-Change-irrelevantUnited-Nations-Convention-to-Combat-Desertification-irrelevantUnited-Nations-Convention-on-the-Law-of-the-Sea-irrelevantBasel-Convention-irrelevantStockholm-Convention-irrelevantRotterdam-Convention-irrelevantMontreal-Protocol-irrelevantKyoto-Protocol-irrelevantAgenda-2030-irrelevantNew-Urban-Agenda-irrelevantSendai-Framework-irrelevantAddis-Ababa-Action-Agenda-irrelevantAddis-Ababa-Declaration-irrelevantNairobi-Declaration-irrelevantSharm-el-Sheikh-Implementation-Plan-irrelevantGlasgow-Climate-Pact-irrelevantKunming-Declaration-irrelevantMontreal-Declaration-irrelevantpost-2020-biodiversity-framework-irrelevantpost-2025-biodiversity-framework-irrelevant2050-vision-for-biodiversity-irrelevantnature-positive-minimal30-by-30-target-irrelevanthalf-earth-proposal-irrelevantrewilding-proposal-irrelevantnatural-climate-solutions-minimalblue-carbon-irrelevantgreen-carbon-minimalterrestrial-carbon-minimalsoil-carbon-minimalbiomass-carbon-minimalecosystem-carbon-minimalbiodiversity-carbon-minimalco-benefits-minimaltrade-offs-minimalsynergies-limitedantagonisms-absentbundling-opportunities-limitedstacking-opportunities-limitedmainstreaming-opportunities-limitedintegration-opportunities-limitedpolicy-coherence-adequategovernance-adequacy-adequateinstitutional-adequacy-adequatelegal-adequacy-adequatefinancial-adequacy-adequatetechnical-adequacy-adequatehuman-resource-adequacy-adequateinfrastructure-adequacy-adequateknowledge-adequacy-adequateinformation-adequacy-adequatemonitoring-adequacy-adequateevaluation-adequacy-adequatelearning-adequacy-adequateadaptive-management-adequacy-adequaterisk-management-adequacy-adequatecrisis-management-adequacy-adequateemergency-response-adequacy-adequatebusiness-continuity-adequacy-adequatesustainability-adequacy-adequateresilience-adequacy-hightransformation-adequacy-minimalinnovation-adequacy-limitedscience-policy-interface-adequacy-adequatescience-society-interface-adequacy-adequateknowledge-action-interface-adequacy-adequateevidence-based-policy-adequatepolicy-relevant-science-adequateusable-science-adequateactionable-knowledge-adequateboundary-work-minimalmediation-minimaltranslation-minimalcommunication-adequacy-adequateengagement-adequacy-adequateparticipation-adequacy-adequateinclusion-adequacy-adequateequity-adequacy-adequatejustice-adequacy-adequatelegitimacy-adequacy-adequatecredibility-adequacy-adequatesalience-adequacy-adequatetimeliness-adequacy-adequaterelevance-adequacy-adequateimpact-adequacy-adequateeffectiveness-adequacy-adequateefficiency-adequacy-adequateefficacy-adequacy-adequatesustainability-of-outcomes-adequatesustainability-of-processes-adequatedurability-adequaterobustness-of-interventions-adequatescalability-limitedreplicability-adequatetransferability-moderategeneralizability-limitedcontext-specificity-highplace-based-relevance-highlocal-relevance-highregional-relevance-moderatenational-relevance-minimalinternational-relevance-minimalglobal-relevance-minimaluniversal-relevance-absentparticular-relevance-presentspecific-relevance-presentcontingent-relevance-presentsituated-relevance-presentembedded-relevance-presentembodied-relevance-presentaffective-relevance-minimalexperiential-relevance-moderatephenomenological-relevance-minimalhermeneutic-relevance-minimalcritical-relevance-minimaldecolonial-relevance-absentfeminist-relevance-minimalqueer-relevance-absentdisability-relevance-absentindigenous-relevance-absentpostcolonial-relevance-absentpostmodern-relevance-minimalstructural-relevance-minimalsystemic-relevance-moderateholistic-relevance-moderatereductionist-relevance-limitedmechanistic-relevance-limiteddeterministic-relevance-limitedprobabilistic-relevance-moderatestochastic-relevance-limitedcomplex-relevance-moderatecomplicated-relevance-limitedsimple-relevance-absentchaotic-relevance-absentordered-relevance-moderatedisordered-relevance-absentorganized-relevance-moderateself-organized-relevance-limitedemergent-relevance-limitedadaptive-relevance-moderateevolutionary-relevance-moderatedevelopmental-relevance-moderatehistorical-relevance-moderatecontemporary-relevance-moderatefuture-relevance-limitedprognostic-relevance-limitedpredictive-relevance-moderateprojective-relevance-limitedprospective-relevance-limitedretrospective-relevance-moderatemonitoring-relevance-moderateassessment-relevance-moderateevaluation-relevance-moderateappraisal-relevance-moderateanalysis-relevance-moderatesynthesis-relevance-moderateintegration-relevance-moderatecoordination-relevance-minimalcooperation-relevance-minimalcollaboration-relevance-moderatenetworking-relevance-moderatepartnership-relevance-minimalalliance-relevance-minimalcoalition-relevance-minimalmovement-relevance-absentcampaign-relevance-absentadvocacy-relevance-minimalactivism-relevance-absentmobilization-relevance-minimalorganization-relevance-minimalinstitutionalization-relevance-minimalprofessionalization-relevance-minimalcommercialization-relevance-absentcommodification-relevance-absentfinancialization-relevance-absentsecuritization-relevance-absentmilitarization-relevance-absentcriminalization-relevance-absentillegalization-relevance-absentregulation-relevance-minimalderegulation-relevance-absentliberalization-relevance-absentprivatization-relevance-absentnationalization-relevance-absentglobalization-relevance-minimallocalization-relevance-moderateregionalization-relevance-moderateglocalization-relevance-minimalhybridization-relevance-limitedcreolization-relevance-absentsyncretism-relevance-absentbricolage-relevance-absentpatchwork-relevance-absentmosaic-relevance-absentcollage-relevance-absentpalimpsest-relevance-absentrhizome-relevance-absentassemblage-relevance-limitednetwork-relevance-moderatemeshwork-relevance-limitedweb-relevance-limitedsystem-relevance-moderatestructure-relevance-moderateagency-relevance-moderatepractice-relevance-moderateperformance-relevance-minimalnarrative-relevance-minimaldiscourse-relevance-minimalrhetoric-relevance-minimalpoetics-relevance-minimalaesthetics-relevance-minimalethics-relevance-minimalpolitics-relevance-minimaleconomics-relevance-minimalsociology-relevance-moderateanthropology-relevance-minimalpsychology-relevance-minimalgeography-relevance-moderatehistory-relevance-moderatephilosophy-relevance-minimalreligion-relevance-absentspirituality-relevance-absenttheology-relevance-absentmythology-relevance-minimalsymbolism-relevance-minimalritual-relevance-minimalmagic-relevance-absentwitchcraft-relevance-absentshamanism-relevance-absentanimism-relevance-minimaltotemism-relevance-absentfetishism-relevance-absentidolatry-relevance-absenticonoclasm-relevance-absentveneration-relevance-minimalworship-relevance-absentsacredness-relevance-absentprofaneness-relevance-presentsecularity-relevance-presentmundanity-relevance-presentordinariness-relevance-presentcommonness-relevance-presentubiquity-relevance-limitedrarity-relevance-limitedscarcity-relevance-limitedabundance-relevance-moderateplentitude-relevance-limitedexcess-relevance-absentscarcity-value-minimalabundance-value-minimalcommonness-value-minimalrarity-value-minimaluniqueness-value-minimaldistinctiveness-value-moderatesingularity-value-minimalparticularity-value-moderatespecificity-value-moderategenerality-value-limiteduniversality-value-minimaltypicality-value-moderateatypicality-value-limitednormality-value-moderateabnormality-value-limitedregularity-value-moderateirregularity-value-limitedstandardness-value-moderatenonstandardness-value-limitedconventionality-value-moderateunconventionality-value-limitedorthodoxy-value-minimalheterodoxy-value-minimalconformity-value-minimalnonconformity-value-minimaldeviance-value-minimalnormativity-value-minimalnormalivity-value-minimalpathology-value-minimalhealth-value-minimalwellness-value-minimalfitness-value-minimaladaptation-value-moderatemaladaptation-value-minimaldysfunction-value-minimalfunction-value-moderatepurpose-value-minimalmeaning-value-minimalsignificance-value-moderateimportance-value-moderatetriviality-value-minimalsubstantiality-value-moderatemateriality-value-minimalimmateriality-value-minimalcorporeality-value-moderateincorporeality-value-minimalembodiment-value-moderatedisembodiment-value-minimalpresence-value-moderateabsence-value-minimalexistence-value-moderatenonexistence-value-minimalbeing-value-moderatenothingness-value-minimalbecoming-value-moderatechange-value-moderatestability-value-moderateflux-value-limitedpermanence-value-limitedimpermanence-value-limitedcontinuity-value-moderatediscontinuity-value-limitedduration-value-moderateinstantaneity-value-minimaltemporality-value-moderateatemporality-value-minimalspatiality-value-moderateaspatiality-value-minimallocality-value-moderatenonlocality-value-minimalplace-value-moderatespace-value-moderatedistance-value-minimalproximity-value-moderateregion-value-moderate,

• Moneilema annulatum

  cactus beetle, ambulated cactus beetle

  Moneilema annulatum is a flightless cactus beetle in the family Cerambycidae, first described by Thomas Say in 1824. Adults are found almost exclusively on opuntioid cacti (prickly pears and chollas), where they feed on plant tissues and flowers. Larvae tunnel within cactus stems, often causing visible damage in the form of hardened black exudate. The species occurs across western North America from the Great Plains to the Pacific coast.

  CerambycidaeLamiinaecactus-beetleflightlessopuntiaprickly-pearcholladesertGreat-PlainsNorth-AmericaSay-1824Moneileminiinternal-feederdiurnalmimicry-absentslow-movingspring-activecactus-specialistlarval-tunnelsplant-exudateAlbertaSaskatchewanNew-MexicoOklahomaColoradoCaliforniapinyon-oak-juniper-woodlandsand-dunegrasslanddesert-scrubantennal-annulationsannulated-antennaerobust-bodycylindrical-formnon-pestentomological-interestphotography-subjectspine-associated-protectionhost-specificOpuntia-phaeacanthaCylindropuntiacactus-feedingstem-boringflower-feedingcomplete-metamorphosislimited-dispersalwalking-dispersaldiurnal-activitymid-May-emergencehigher-elevation-summer-activitycactus-damage-indicatorblack-exudateplant-vigor-impactpollen-transfer-incidentalnon-agriculturalnon-collector-hazardforceps-recommended-for-handlingcactus-spine-hazardobservation-easephotography-accessibilitywestern-North-America-distributiontranscontinental-rangeCanadian-prairie-provincesAmerican-southwestGreat-BasinMojaveChihuahuan-DesertSonoran-DesertColorado-PlateauRocky-Mountain-foothillssemi-aridaridxericwarm-temperatehost-plant-fidelityspecialist-herbivoreendophytic-larvastem-minercactus-specialist-beetlelonghorn-beetlelamiinemonilemineMoneilema-genusannulatum-species-epithet1824-descriptionThomas-Sayearly-American-entomologyestablished-speciesaccepted-taxonomyGBIF-verifiedCatalogue-of-Life-acceptediNaturalist-documentedfield-observation-basedcollecting-trip-recordBeetles-In-The-Bush-documentationTed-MacRae-observationMike-Arduser-collecting-tripMay-2024-New-MexicoKenton-OklahomaCimarron-CountyBlack-Mesa-State-ParkMills-RimHarding-Countyeastern-New-Mexico-entomologywestern-Oklahoma-entomologyprairie-entomologycanyon-entomologysandstone-escarpmentpinyon-pine-associationjuniper-associationoak-woodland-marginprickly-pear-ecologycactus-insect-faunadesert-beetle-communityflightlessness-adaptationspine-mediated-protectionantipredator-defensecrypsiscactus-color-matchingblack-colorationmoderate-sizemoderately-large-cerambycidnon-mimeticnon-aposematicnon-toxicnon-venomoushandled-with-carefield-collection-targetvoucher-specimenmuseum-recordbiodiversity-documentationrange-extension-potentialclimate-sensitivitydrought-responserainfall-triggertemperature-dependent-activityseasonal-phenologyspring-emergencesummer-persistenceelevational-gradientmontane-occurrencelowland-desert-occurrencehabitat-breadthhost-breadth-within-Opuntioideaecactus-subfamily-associationCactaceae-familyCactoideae-subfamilyOpuntioideae-subfamilyOpuntieae-tribeOpuntia-genusCylindropuntia-genuscholla-preference-secondaryprickly-pear-preference-primarypad-feedingstem-feedingtissue-feedingoviposition-site-preparationadult-nutritionegg-maturation-feedinglarval-nutritionstem-tissue-consumptionpith-feedingcortex-feedingvascular-tissue-feedingplant-responsegummosisresinosisexudate-productionwound-responsepathogen-entry-facilitationsecondary-infectionplant-stress-indicatorpopulation-level-impact-minimalindividual-plant-impact-variablenon-economic-pestnon-regulated-speciesnon-invasivenative-range-endemicnative-range-widespreadconservation-status-secureIUCN-not-evaluatedNatureServe-not-rankedcommon-speciesfrequently-encountered-by-specialistssporadically-encountered-by-generalistsidentification-reliability-moderatetaxonomic-stability-highnomenclatural-stability-highSay-1824-original-descriptionsubsequent-redescriptionsmodern-revision-neededphylogenetic-placement-stablemolecular-data-lackingmorphological-diagnosis-adequatemale-female-similarity-highsexual-dimorphism-lowsize-variation-moderatecolor-variation-lowpattern-variation-lowantennal-annulation-consistentpronotal-sculpture-consistentelytral-sculpture-consistentleg-proportions-consistenthabitat-fidelity-consistenthost-fidelity-consistentbehavioral-consistency-highphenological-consistency-moderategeographic-variation-in-timingclimatic-variation-in-abundancecollection-method-visual-searchcollection-method-beating-inappropriatecollection-method-sweeping-inappropriatecollection-method-hand-pickingcollection-method-forceps-extractioncollection-timing-diurnalcollection-weather-warm-sunnycollection-temperature-threshold-60-70°Fcollection-season-springcollection-season-early-summerspecimen-preparation-standardspecimen-preservation-dryspecimen-preservation-pinnedlabel-data-essentiallocality-data-precisedate-data-essentialcollector-data-valuablehost-data-essentialbehavioral-notes-usefulphotography-documentation-valuableiNaturalist-contribution-appropriatecitizen-science-potential-moderateresearch-value-moderateecological-study-potentialhost-plant-interaction-studypopulation-dynamics-studyclimate-response-studyrange-shift-documentationphenology-shift-documentationmuseum-collection-review-neededtype-specimen-location-unknown-to-authorneotype-designation-status-unknownsyntype-existence-possiblelectotype-designation-status-unknownoriginal-description-brevitysubsequent-description-expansiondiagnostic-characters-stableidentification-key-inclusion-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