Hymenoptera-parasite
Guides
Anthrax irroratus
Spotted Bee Fly
Anthrax irroratus is a bee fly (family Bombyliidae) widespread across North America, ranging from Alaska and northern Canada to Central America and the Caribbean. Adults are pollinators that frequent flowers, while larvae are parasitoids of solitary bees and wasps in at least five families. The species is one of numerous bee flies that exploit ground-nesting Hymenoptera as hosts.
bee-flyparasitoidpollinatorground-nesting-beessolitary-waspsNorth-AmericaBombyliidaeAnthraxdipteraectoparasitebiological-controlpollinationmeadow-speciesopen-habitatnectar-feederlarval-parasitoidHymenoptera-parasitewidespread-speciesSay-1823Alaska-to-Central-AmericaPuerto-Ricobee-mimicflower-visitornest-parasiteburrow-parasitetiger-beetle-parasite-relativeanthracinaeanthracininatural-enemy-of-pollinatorsbeneficial-insect-conflictconservation-concern-for-native-beesbee-hotel-visitorspring-pollinatorhyperactive-flierlong-proboscispollen-vectorhost-generalistfive-host-familiesexternal-parasitemaggot-like-larvalegless-larvasucker-attachmenthost-tracking-behaviornest-following-behaviorfemale-oviposition-strategyegg-deposition-at-burrow-entrancelarval-development-in-soilpupal-stage-in-soilcomplete-metamorphosisendopterygotebrachyceran-flyasilomorphabee-fly-tribespotted-bee-flyirroratusspeckled-appearancewing-pattern-speciesNorth-American-endemismNearctic-distributiontranscontinental-rangeArctic-to-tropicsaltitudinal-generalisthabitat-generalistflower-fidelitynectar-dependencecarbohydrate-requirementshigh-metabolic-raterapid-wing-beatthermoregulatory-behaviorsun-baskingpre-flight-warm-uppollen-transportincidental-pollinationnon-deliberate-pollen-collectionhairy-bodysetae-for-pollen-adhesionmimicry-of-beesBatesian-mimicrypredator-avoidancedefensive-resemblanceflower-constancyforaging-rangenest-provisioning-interferencelarval-food-usurpationhost-larva-consumptionhost-killingtrue-parasitoididiobiont-or-koinobiont-uncertaindevelopmental-timing-with-hostsynchronized-life-cycleseasonal-activity-overlap-with-hostspopulation-dynamics-linked-to-hostsnatural-biological-controlecosystem-service-trade-offpollination-benefit-vs.-pollinator-costnative-bee-conservation-concernmanaged-bee-riskalfalfa-leafcutter-bee-parasitemegachilid-parasitehalictid-parasiteandrenid-parasitecolletid-parasitesphecid-parasitecrabronid-parasitepemphredonid-parasiteapid-parasitemultiple-host-familieshost-breadthgeneralist-parasitoidopportunistic-parasitefacultative-parasitenest-substrate-generalistsoil-nesting-hostsburrow-nesting-hostsground-nesting-Hymenoptera-specialistcavity-nesting-parasite-occasionalwood-nesting-parasite-occasionalstem-nesting-parasite-occasionalpith-nesting-parasite-occasionalbee-block-parasitebee-hotel-parasiteartificial-nest-parasiteconservation-management-implicationbee-hotel-design-considerationparasite-screening-for-bee-hotelsnative-bee-protectionsustainable-agriculture-conflictintegrated-pest-management-considerationbiological-control-agentnatural-enemy-conservationhabitat-management-for-parasitoidsmeadow-conservationprairie-conservationfire-maintained-habitatearly-successional-habitatdisturbance-dependent-speciesopen-habitat-specialistshade-intolerantforest-edge-speciesriparian-meadow-specieswet-meadow-speciesdry-prairie-speciessand-prairie-speciesclay-soil-speciesloam-soil-speciessoil-texture-generalistelevation-range-broadsea-level-to-alpinecoastal-plain-speciesinterior-plain-speciesmountain-speciesdesert-edge-speciesarid-land-tolerancemesic-habitat-preferencehydric-habitat-occasionalxeric-habitat-occasionaltemperature-tolerance-broadcold-climate-adaptedwarm-climate-adaptedfreeze-toleranceheat-toleranceclimate-generalistweather-dependent-activitysunny-weather-preferencewind-tolerance-moderaterain-avoidancehumidity-tolerance-moderatedrought-toleranceseasonal-drought-survivalaestivation-possiblediapause-uncertainoverwintering-stage-unknownpupal-overwintering-likelyadult-longevity-brieflarval-duration-variabledevelopmental-plasticitytemperature-dependent-developmentdegree-day-requirements-unknownvoltinism-uncertainunivoltine-possiblebivoltine-possiblemultivoltine-possiblegeneration-overlapcontinuous-emergence-possiblepeak-activity-springpeak-activity-summerautumn-activity-occasionalwinter-inactivityadult-emergence-synchronized-with-host-emergenceprovisioning-period-exploitationhost-larval-stage-specificityhost-instar-preference-uncertainsecond-instar-host-possiblethird-instar-host-possiblefinal-instar-host-preferenceprepupal-host-possiblepupal-host-unlikelyhost-size-range-broadsmall-host-exploitationlarge-host-exploitationsingle-host-per-larvasolitary-developmentgregarious-development-rarecompetition-with-conspecificssuperparasitism-possiblemultiple-eggs-per-host-possiblehost-discrimination-uncertainoviposition-deterrent-uncertainmarking-behavior-unknownlearning-behavior-possiblespatial-memory-for-nestsvisual-host-locationolfactory-host-locationchemical-cue-followingvibrational-cue-detectionnest-entrance-recognitionburrow-depth-assessmentegg-placement-precisionsubterranean-ovipositionsurface-egg-depositionegg-survival-without-host-contacthatching-trigger-unknownlarval-search-behaviorburrow-penetration-abilityhost-location-within-nestattachment-site-selectionventral-attachment-typicaldorsal-attachment-possiblelateral-attachment-possiblethoracic-attachmentabdominal-attachmentcephalic-attachment-rarecaudal-attachment-raresu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Thick-headed Flies
Conopidae, commonly known as thick-headed flies, is a family of flies within the Brachycera suborder of Diptera and the sole member of the superfamily Conopoidea. The family comprises approximately 800 species in 47 genera worldwide, with about 70 species found in North America. Adult conopids are frequent flower visitors, feeding on nectar with their often elongated proboscis. The larvae of all conopids are internal parasitoids, primarily of aculeate Hymenoptera (stinging wasps and bees), with adult females aggressively intercepting hosts in flight to deposit eggs.
Conopinae
thick-headed flies
Conopinae is a subfamily of flies within the family Conopidae, commonly known as thick-headed flies. Members are characterized by their distinctive head shape, with a broad, inflated frons that gives the group its common name. The subfamily includes two tribes: Conopini (genera Conops and Leopoldius) and Physocephalini (genera Physocephala and Physoconops). These flies are known for their parasitoid life history, with females attacking various hymenopteran hosts.
Dalmanniinae
Dalmanniinae is a subfamily of thick-headed flies (family Conopidae) within the order Diptera. Members of this subfamily are distinguished from other conopids by specific morphological features of the head and wing venation. The subfamily contains genera such as Dalmannia and Zodion, which are primarily found in the Holarctic region. These flies are parasitoids, with larvae developing within the bodies of Hymenoptera hosts.
Eupathocera auripedis
Eupathocera auripedis is a species of twisted-wing parasite in the order Strepsiptera, family Xenidae. As an obligate endoparasite, it infects host insects and completes its life cycle within their bodies. The species has been documented parasitizing the grass-carrying wasp Isodontia mexicana. Like other strepsipterans, it exhibits extreme sexual dimorphism and complex life history traits associated with its parasitic lifestyle.
Eupathocera lugubris
Eupathocera lugubris is a species of twisted-wing parasite (order Strepsiptera) in the family Xenidae. The species was originally described by Pierce in 1908 and is currently treated as a synonym of Paraxenos lugubris. Strepsipterans are obligate endoparasites of other insects, with this species likely parasitizing wasps or related hosts. The genus is part of the diverse and complex parasitic wasp-associated Strepsiptera fauna of North America.
Euphyto
Euphyto is a genus of flies in the family Sarcophagidae, subfamily Miltogramminae. Members of this genus are parasitoid flies that develop in the nests of solitary bees and wasps. The genus was established by Townsend in 1908. Miltogrammine flies are known for their kleptoparasitic or parasitoid lifestyle, laying eggs in or near host provisions.
Haltichella
Haltichella is a genus of chalcidid wasps in the family Chalcididae, established by Spinola in 1811. The genus contains at least 20 described species with a cosmopolitan distribution. Members are parasitoid wasps whose larvae develop in or on host organisms. Host associations include Lepidoptera (families Bucculatricidae, Gelechiidae, Momphidae, Notodontidae, Oecophoridae, Pyralidae, Tortricidae) and Hymenoptera (Braconidae, Ichneumonidae).
Metopia argyrocephala
satellite fly
Metopia argyrocephala is a kleptoparasitic satellite fly in the family Sarcophagidae. Unlike most kleptoparasitic flies that specialize on either bees or wasps, this species has been documented developing in nests of both hymenopteran groups. Adult females follow host wasps or bees to their nests and deposit larvae at the entrance, which then crawl inside to parasitize the host's brood. The species is native to Europe and has been recorded across northern Europe including Belgium, Denmark, Norway, and Sweden.
Myopa rubida
Myopa rubida is a species of thick-headed fly in the family Conopidae, first described by Bigot in 1887. Like all conopids, it is an internal parasite of bees and wasps. The genus Myopa is seldom recorded, partly due to identification difficulties. Species in this genus are sometimes known as 'beegrabbers' for their habit of intercepting host insects in flight to oviposit eggs between abdominal segments.
Phrosinella aldrichi
Phrosinella aldrichi is a species of satellite fly in the family Sarcophagidae, subfamily Miltogramminae. The species was described by Allen in 1926. Like other miltogrammine flies, it is known to parasitize the nests of solitary wasps. It has been specifically reported as a parasite of the nests of the sand wasp Clitemnestra bipunctata (formerly Ochleroptera bipunctata), where adult females enter burrows and deposit live larvae that feed on paralyzed prey stored by the host wasp.
Physocephala burgessi
thick-headed fly
Physocephala burgessi is a species of thick-headed fly in the family Conopidae. Like other members of its genus, it is a parasitoid of bees and wasps. The species exhibits wasp mimicry in appearance and behavior, including an elongated abdomen resembling a wasp waist and dark pigmentation along the front wing margin that mimics the folded wing appearance of potter wasps at rest.
Physoconops
thick-headed flies
Physoconops is a genus of thick-headed flies (family Conopidae) containing approximately 13 described species. Members are wasp mimics, resembling potter wasps in appearance and behavior. The genus occurs primarily in the Neotropical and Andean regions, with some species in North America. Like other conopids, Physoconops species are endoparasitoids of bees and aculeate wasps.
Physoconops weemsi
A species of thick-headed fly in the family Conopidae, described by Camras in 2007. As a member of the genus Physoconops, it likely exhibits wasp-mimicking morphology and parasitic behavior toward Hymenoptera, though species-specific documentation remains limited. The genus is characterized by elongated abdomens resembling potter wasps and heavily pigmented wing margins that mimic the folded wings of resting wasps.
Stylops
stylops
Stylops is a genus of obligately endoparasitic insects in the family Stylopidae, order Strepsiptera. Members are endoparasites of Hymenoptera, particularly bees in the family Andrenidae. The genus exhibits extreme sexual dimorphism: males are free-living with wings and functional limbs, while females are neotenic, legless, and remain permanently embedded within host abdomens with only the cephalothorax protruding. The genus has cosmopolitan distribution with numerous described species, though taxonomic status of many remains problematic due to reliance on poor female morphological characters.
Toxophora
bee flies
Toxophora is a genus of bee flies (family Bombyliidae) comprising approximately 47 described species with worldwide distribution. These flies are characterized by a distinctive hunchbacked, robust body form and are most abundant in the southwestern United States and western Mediterranean region. Larvae are parasitoids of solitary wasps and bees, particularly in the family Vespidae.