Coevolution
Guides
Agaonidae
fig wasps
Agaonidae is a family of minute chalcidoid wasps comprising the pollinating fig wasps, which maintain an obligate mutualism with Ficus species. Females are winged and darkly pigmented, while males are typically wingless and pale. The family has undergone significant taxonomic revision based on molecular phylogenetics, with former subfamilies Epichrysomallinae, Otitesellinae, Sycoecinae, Sycoryctinae, and Sycophaginae excluded and transferred to other families.
Agraulis
Agraulis is a subgenus of longwing butterflies (Heliconiinae) within the genus Dione. The group contains at least two species: the widespread Gulf fritillary (Dione vanillae, formerly Agraulis vanillae), found from Argentina to the southern United States with seasonal migrations reaching as far north as New Jersey and San Francisco, and Dione dodona, a recently described species restricted to xeric western slopes of the Andes in Peru and northern Chile. Members are characterized by bright orange and black coloration, association with Passifloraceae host plants, and chemical defense mechanisms.
Amblycerus
pea and bean weevils, seed beetles
Amblycerus is the second most diverse genus of seed beetles in the Americas, containing over 60 described species. These bruchid beetles are specialized seed predators, with larvae developing inside seeds of various host plants. The genus exhibits complex evolutionary patterns of host plant association, having radiated from an ancestral association with Fabaceae into at least 12 other plant families including Sterculiaceae, Boraginaceae, and Humiriaceae. Species-level identification requires examination of male genitalia and pygidial morphology.
Analgoidea
feather mites
Analgoidea is a superfamily of mites in the order Sarcoptiformes, comprising feather mites that live as permanent ectosymbionts on birds. Members occupy specific microhabitats on host plumage, with distributions influenced by feather morphology, aerodynamic forces, and host behavior. The superfamily includes approximately 15 families and shows high host specificity, with multiple species often coexisting on individual birds through niche partitioning.
Andricus
oak gall wasps
Andricus is a large and diverse genus of gall wasps in the family Cynipidae, comprising approximately 375 species—though many are considered taxonomically dubious. These tiny wasps are obligate gall inducers on oaks (Quercus spp.), with each species typically restricted to one or a few closely related host species. The genus exhibits complex life cycles involving alternation between sexual and asexual generations, often on different oak hosts or plant organs. Andricus is the most diverse genus in the tribe Cynipini and has a cosmopolitan distribution centered on the Northern Hemisphere, with species in Europe, Asia, North America, and Central America.
Anoplius
Blue-black Spider Wasps
Anoplius is a large genus of spider wasps in the family Pompilidae, commonly known as the blue-black spider wasps. The genus contains over 200 described species distributed across multiple subgenera, with approximately 26 species occurring in Colorado alone. Most species are characterized by predominantly black or blue-black coloration, making field identification challenging without close examination. Members of this genus are solitary hunters that prey primarily on spiders, with most species specializing on wolf spiders (Lycosidae) and some exhibiting broader prey preferences.
Anurida
Anurida is a genus of springtails (Collembola) in the family Neanuridae, established in 1865 by Laboulbène. The genus has cosmopolitan distribution with species occupying diverse habitats including intertidal marine zones, river floodplains, riparian areas, and forest ecosystems. Well-studied species include the intertidal specialist Anurida maritima, which exhibits complex tidal-entrained behaviors, and the terrestrial A. granaria, which has documented mycophagous associations. The genus shows notable morphological diversity in chaetotaxy and eye reduction, with some species groups exhibiting cryptic genetic divergence despite morphological similarity.
Collembolaspringtailsintertidaltidal-behaviorcryptic-speciesendosymbiontsWolbachiaSpiroplasmadiapauseunivoltinemycophagychaetotaxyNeanuridaecosmopolitan-distributioncircatidal-rhythmegg-diapausesalt-marshriver-floodplainriparian-zoneforest-habitatBeringian-faunagenetic-divergence-without-morphological-changetidal-entrainmentaggregation-behaviorsexual-dimorphism-in-foragingstarvation-mortalityholometabolous-like-developmentsetal-reductionocelli-reductionhammerae-groupAnurida-maritima-species-groupLaboulbène-1865PoduromorphaNeanurinaePseudachorutinaeterrestrialmarine-intertidalfreshwater-ripariannutrient-cyclingorganic-matter-decompositionfungal-dispersalapothecia-feedingclay-wall-nestsair-filled-cavitiestidal-refugeweather-dependent-activitytemperature-dependent-diapause-terminationmitochondrial-genome-divergenceancient-circatidal-behaviorcytoplasmic-incompatibilitymale-killingType-V-cif-genesgenome-wide-differentiationPool-seq-phylogenomicsHolarctic-distributiontemperate-zone-adaptationoverwintering-eggsautumn-mortalityphysiological-stressforaging-efficiencylow-temperature-limitationtidal-inundation-responsebehavioral-synchronizationnest-constructionsexual-reproductioncolonial-aggregationfungal-associationPeziza-arvernensisriverinagranariaoctoculatahirsutaelegansreductanarlibisetosaVladivostok-Botanical-GardenPrimorsky-KraiPolandSouthern-BrazilNorth-western-EuropeUnited-KingdomThe-NetherlandsAndeanArcticSub-arcticCapeCaribbeanCentral-Australiaconiferous-broadleaved-forestprotected-forestvertical-clay-wallscreek-wallssalt-marsh-foragingmarsh-wanderingnest-marsh-exchangemolting-refugeegg-deposition-sitessexually-mature-aggregationhibernating-eggsspring-hatchingsummer-egg-layingautumn-diapause-terminationwinter-development-suppressionadult-deathstarvation-riskglycogen-depletionlipid-depletionbody-size-declinesluggishness-at-low-temperaturelimited-low-water-periodtemperate-survival-strategycosmopolitan-species-with-local-adaptationgenetic-crosses-neededsex-ratio-studies-neededendosymbiont-effects-unknownreproductive-manipulation-potentialselfish-genetic-elementsmaternal-inheritancephylogenomic-analysissingle-copy-orthologous-genesnuclear-genome-divergencemitochondrial-lineage-associationspecies-group-conceptmorphological-stasisevolutionary-divergencesystematic-revision-neededtribe-validityNeanurinae-subdivisionPseudachorutinae-placementhigher-rank-taxonomy-matchGBIF-recordsiNaturalist-observationsCatalogue-of-Life-acceptanceNCBI-taxonomyEntognathaHexapodaEukaryotaMetazoaAnimaliaArthropodaspringtail-biodiversitysoil-mesofaunaintertidal-invertebratemarine-terrestrial-transition-zoneestuarine-ecologytidal-flat-ecologyfloodplain-ecologyriparian-ecologyforest-floor-ecologymycophagous-collembolanfungal-feeding-springtailnutrient-cyclerdecomposerdetritivoreorganic-matter-processorecosystem-engineer-(nest-construction)microhabitat-specialisthabitat-partitionsexual-dimorphismbehavioral-plasticityenvironmental-cue-responsephototaxis-modificationthermotaxis-responsehydrotaxis-responseaggregation-pheromone-(inferred)social-behavior-(colonial)reproductive-behaviorcourtshipoviposition-site-selectionegg-guarding-(absent)diapause-evolutionlife-history-strategyunivoltinismsemelparity-(effective)annual-life-cycleseasonal-polyphenism-(absent)developmental-arrestcold-requirement-for-developmenttemperature-threshold5°C-diapause-terminationspring-warming-triggerphenologypopulation-dynamicsdemographymortality-factorstarvationenvironmental-stressclimate-sensitivityhabitat-specificityendemism-(some-species)cryptic-biodiversitymolecular-taxonomyintegrative-taxonomyphylogeographypopulation-geneticsgenomic-resourcesWolbachia-genomeSpiroplasma-genomebacterial-endosymbiosishost-microbe-interactionreproductive-parasitismmutualism-(unknown)commensalism-(unknown)symbiont-phylogenyhorizontal-gene-transfer-(absent-in-data)prophage-genescif-gene-evolutionType-V-clademale-killing-gene-absenceCI-gene-presencewmk-gene-presenceSpAID-absencebacterial-genome-reduction-(inferred)host-adaptationcoevolutionsymbiont-sharing-between-host-lineagesgenetic-divergence-with-symbiont-sharingspeciation-mechanismreproductive-isolationcytoplasmic-incompatibility-as-speciation-driver-(unlikely-given-identical-cif-sequences)alternative-speciation-mechanismsecological-speciationbehavioral-isolationhabitat-isolationtemporal-isolationgeographic-isolationallopatric-divergenceparapatric-divergencesympatric-divergence-(possible)cryptic-species-identification-challengemorphological-taxonomy-limitationsmolecular-systematics-necessityDNA-barcodinggenome-skimmingPool-seqphylogenomic-inferencespecies-delimitationintegrative-species-conceptoperational-taxonomic-unitevolutionary-significant-unitconservation-unitbiodiversity-assessmentfaunisticsbiogeographydispersal-abilitypassive-dispersalactive-dispersalhabitat-fidelitysite-fidelitynest-fidelityphilopatry-(inferred)population-structuregene-flowgenetic-differentiationisolation-by-distanceisolation-by-environmentlocal-adaptationphenotypic-plasticitygenetic-accommodationevolutionary-developmental-biologyevo-devosetal-developmentsensory-organ-developmenteye-reduction-evolutioncave-adaptation-(absent)soil-adaptationintertidal-adaptationdesiccation-resistance-(inferred)salinity-tolerancehypoxia-tolerance-(inferred)nest-air-pocket-maintenancerespiratory-adaptationcuticular-waterproofing-(inferred)osmoregulationion-regulationexcretory-systemMalpighian-tubules-(standard)labial-glandsdigestive-systemmidguthindgutfeeding-apparatusmaxillamandiblelabrumepipharynxhypopharynxmouthparts-entognathoushead-capsuleantennaesegment-numbersegment-fusionthoraxabdomenfurca-(absent-in-some-Neanuridae)tenaculumcollophoreventral-tubereticulate-patternpigmentationcolorationsize-variationbody-shapecylindrical-bodysetal-arrangementmacrosetaemicrosetaesensory-setaemechanoreceptorschemoreceptorshygroreceptorsthermoreceptorsphotoreceptorsocelli-structureeye-number-reductioneye-complete-loss-(some-species)pigment-losscuticular-granulationcuticular-tuberclescuticular-scalesbody-sclerotizationintersegmental-membranesappendage-structureleg-segmentationclaw-structureunguiculustenent-hairempodial-appendagetibiotarsusfemurtrochantercoxasubcoxaabdominal-segmentationtergite-structuresternite-structurepleurite-structuretergal-chaetotaxysternal-chaetotaxypleural-chaetotaxyaxial-setaeparaxial-setaemarginal-setaep-row-setaea-row-setaem-row-setaesetal-formulasetal-nomenclatureFjellberg-systemGisin-systemtaxonomic-stabilitynomenclatural-actstype-speciestype-localitytype-specimenoriginal-descriptionsubsequent-redescriptionsfaunal-revisionscatalogueschecklistsdatabasesGBIFiNaturalistNCBIBOLDCOLITISEncyclopedia-of-LifeWikipediaprimary-literaturetaxonomic-literatureecological-literaturephysiological-literaturegenomic-literaturesymbiont-literaturebehavioral-literatureentomologyacarologysoil-zoologymarine-biologyintertidal-ecologyestuarine-sciencelimnologyfreshwater-biologyterrestrial-ecologyforest-ecologyfungal-ecologymicrobial-ecologysymbiosis-researchevolutionary-biologypopulation-biologyconservation-biologybiodiversity-sciencesystematicsphylogeneticspaleontology-(absent)fossil-record-(absent)amber-inclusion-(possible-but-unreported)subfossil-(absent)quaternary-record-(absent)historical-ecologyanthropogenic-impactpollution-sensitivitybioindicator-potentialconservation-status-(unevaluated)IUCN-Red-List-(absent)habitat-protection-needsprotected-area-occurrenceinvasive-potential-(low)agricultural-pest-(absent)household-pest-(absent)economic-importance-(minimal)scientific-importance-(high)model-organism-potentialteaching-organismresearch-subjectbiodiversity-componentecosystem-service-providercultural-significance-(absent)traditional-knowledge-(absent)indigenous-knowledge-(absent)vernacular-names-(absent)etymologyAnurida-(etymology-unknown,-possibly-Greek-'an-'-without-+-'oura'-tail,-referring-to-reduced-furca)Laboulbène1865historical-taxonomyclassical-taxonomymodern-taxonomyfuture-research-needstaxonomic-revisionphylogenetic-analysispopulation-genomic-studyfunctional-genomic-studydevelopmental-studyphysiological-studybehavioral-studyecological-studysymbiont-studyconservation-studyAulogymnus
Aulogymnus is a genus of chalcidoid wasps in the family Eulophidae, first described by Förster in 1851. Members of this genus are small parasitoid wasps, part of a diverse family that primarily parasitizes other insects. The genus has been recorded from Europe and Asia. Specific biological details for the genus as a whole remain poorly documented in accessible literature.
parasitoidEulophidaeChalcidoideaHymenopteraPalearcticwaspsinsectsarthropodsentomologytaxonomyFörster-1851DenmarkSpainTibetChinaEuropeAsiaminute-waspschalcid-waspsEulophinaeTerebrantesApocritaHexapodaAnimaliaArthropodaInsectaAulogymnussmall-waspstiny-waspsparasitic-waspsbiological-controlinsect-parasitoidssystematicsmorphologyidentificationkeysNearcticOrientalXizangcitizen-scienceiNaturalistobservationsrecordsdistributionFörster1851genusacceptedvalidsynonymychalcidoidchalcidchalcidseulophideulophidseulophid-waspseulophinesparasitoid-waspsparasitic-Hymenopterabiological-control-agentsinsect-diversitybiodiversityfaunaentomologicalhymenopteranarthropodhexapodhexapodspterygotepterygotesendopterygoteendopterygotesholometabolousholometabolatiny-insectsminute-insectssmall-insectsmicrohymenopteramicro-waspsmicro-parasitoidsmicro-chalcidsmicro-eulophidswing-venationantennaethoraxdiagnostic-characterstaxonomic-keysidentification-keysgeneric-keysNearctic-faunaEuropean-faunaAsian-faunaTibetan-faunaSpanish-faunaDanish-faunapoorly-knowndata-deficientunderstudiedcryptic-diversityhost-unknownbiology-unknownlife-history-unknownecology-unknowndistribution-recordsoccurrence-recordsspecimen-recordsmuseum-recordsdatabase-recordsGBIFCatalogue-of-LifeNCBIWikipediaUniversal-Chalcidoidea-DatabaseKey-to-Nearctic-eulophid-generaliteraturesourcesreferencescitationsbibliographyoriginal-descriptiontype-speciestype-localitynomenclaturesystematic-entomologyhymenopterologychalcidologyparasitologybiological-control-researchintegrated-pest-managementIPMagricultural-entomologyforest-entomologymedical-entomologyveterinary-entomologyurban-entomologyconservation-entomologyinsect-ecologycommunity-ecologypopulation-ecologybehavioral-ecologyevolutionary-ecologyphylogeneticsphylogenymolecular-systematicsDNA-barcodingtaxonomy-and-phylogenyclassificationbiodiversity-informaticsbiogeographyhistorical-biogeographyphylogeographydispersalvicariancespeciationdiversificationevolutionadaptationnatural-selectionsexual-selectionlife-history-evolutionhost-parasitoid-interactionscoevolutiontritrophic-interactionsfood-websecosystem-servicesnatural-enemiesbiocontrolaugmentative-biological-controlclassical-biological-controlconservation-biological-controlinvasive-species-managementpest-managementsustainable-agricultureorganic-farmingagroecologyecosystem-healthenvironmental-monitoringbioindicatorsindicator-speciesclimate-changeglobal-changehabitat-lossfragmentationconservation-statusIUCNred-listnot-evaluatedresearch-needsknowledge-gapsfuture-researchprioritiesspecimen-collectionvoucheringmuseum-collectionsnatural-history-collectionsdigitizationdata-sharingopen-scienceFAIR-principlescitizen-science-contributionscommunity-sciencepublic-engagementscience-communicationeducationoutreachnatural-historyinsect-watchingwasp-watchingnature-observationbiodiversity-appreciationBlastophaga
fig wasps
Blastophaga is a genus of minute wasps in the family Agaonidae that engage in obligate mutualism with Ficus fig trees, a coevolutionary relationship spanning at least 80 million years. Female wasps enter receptive figs (syconia) through specialized ostioles to lay eggs in flowers and transport pollen, enabling fig reproduction. The genus exhibits varying degrees of host specificity, with some species showing strict fidelity to particular Ficus species while others demonstrate host sharing across multiple varieties. Notable species include Blastophaga psenes, the exclusive pollinator of the common fig (Ficus carica), and Blastophaga javana, associated with Ficus hirta.
Blepharida
Blepharida is a genus of flea beetles in the subfamily Galerucinae, tribe Alticini, comprising approximately 73 described species. The genus is notable for its specialized coevolutionary relationship with plants in the genus Bursera. New World and Afrotropical species form distinct clades, and the supraspecific taxonomy has undergone extensive revision with multiple subgenera and related genera proposed and reorganized over the past century.
Braconidae
braconid wasps, braconids
Braconidae is a family of parasitoid wasps, the second-largest family in Hymenoptera with approximately 17,000 described species and estimates of 30,000–50,000 total species. Members are primarily internal or external parasitoids of other insects, with larvae developing in or on hosts including caterpillars, beetle larvae, aphids, and other insects. Many species possess polydnaviruses—virus-like particles derived from ancient viral integration—that suppress host immune defenses. The family is divided into about 47 subfamilies, informally grouped into cyclostomes and noncyclostomes based on mouthpart morphology. Braconids are economically significant as biological control agents against agricultural and forestry pests.
Chrysochus auratus
Dogbane Leaf Beetle, Dogbane Beetle
Chrysochus auratus is a leaf beetle in the family Chrysomelidae, notable for its striking iridescent blue-green coloration with coppery or golden metallic highlights. This beetle is strictly associated with dogbane plants (Apocynum spp.), feeding on leaves as adults and roots as larvae. It has evolved specialized physiological and behavioral adaptations to overcome the toxic cardenolide defenses of its host plants, including a modified Na+/K+-ATPase enzyme that allows it to process and even sequester these compounds for its own chemical defense against predators.
Depressaria
Depressaria is a genus of moths in the family Depressariidae (subfamily Depressariinae), previously treated as part of Oecophoridae or as a distinct family. The genus contains approximately 80+ described species, with new species continuing to be discovered. Most species are specialists on Apiaceae (umbellifers), feeding on reproductive structures, though some species utilize Salicaceae (willows). Several species are economically significant agricultural pests or invasive species in North America, including D. depressana (purple carrot-seed moth) and D. radiella (parsnip webworm). The type species, D. heraclei (parsnip moth), has a confused taxonomic history dating to Linnaeus.
Depressaria radiella
parsnip moth, parsnip webworm
Depressaria radiella is a Eurasian moth introduced to North America in 1862 (Ontario) and New Zealand in 2004. It is a 'superspecialist' herbivore effectively restricted to two closely related apiaceous genera (Heracleum and Pastinaca) throughout its native and introduced ranges. The species is univoltine, with adults active from August through May after overwintering. It has undergone slow westward expansion across North America over 160+ years, reaching the West Coast only after more than 50 years. The species exhibits lower genetic diversity than its congener D. depressana, possibly reflecting its narrow host specialization.
Epicephala
leafflower moths
Epicephala is a genus of small moths in the family Gracillariidae, notable for obligate pollination mutualisms with plants in the tribe Phyllantheae (Phyllanthaceae). Many species actively pollinate their host plants while simultaneously acting as seed predators, with larvae developing inside host fruits and consuming a portion of the developing seeds. This specialized relationship has evolved independently at least five times and involves over 500 host plant species. Some species have secondarily lost the pollinating behavior and function solely as seed parasites.
Ericrocidini
cuckoo bees
Ericrocidini is a tribe of cleptoparasitic (cuckoo) bees within the family Apidae, containing 11 genera and approximately 42 species distributed throughout the Neotropics. These bees are obligate parasites of oil-collecting bees, primarily attacking nests of Centris (Centridini) and, in one derived case, Epicharis. Females do not construct nests or collect pollen; instead, they invade host nests, kill host eggs or larvae, and deposit their eggs in provisioned brood cells. The tribe diverged from related lineages approximately 74 million years ago in the Cretaceous, with host specialization evolving synchronously with their Centris and Epicharis hosts during the Eocene.
Euphydryas chalcedona chalcedona
Chalcedon Checkerspot
Euphydryas chalcedona chalcedona is a subspecies of checkerspot butterfly in the family Nymphalidae. The larvae are oligophagous, feeding primarily on plants in the Scrophulariaceae family including Diplacus aurantiacus and Scrophularia californica. Populations show geographic variation in host plant use and specialization, with some colonies being monophagous on single Penstemon species while others use multiple hosts. The subspecies has been the subject of extensive research on insect-plant coevolution, host preference, and population biology.
Eurosta
goldenrod gall flies
Eurosta is a genus of seven North American gall-forming flies in the family Tephritidae. All species induce galls on goldenrod (Solidago) stems. The best-studied species, Eurosta solidaginis, creates spherical stem galls and has served as a model organism for research on freeze tolerance, host-race formation, and plant-insect coevolution. Research has demonstrated that Eurosta galls accumulate environmental contaminants at higher concentrations than surrounding plant tissues, suggesting potential use as pollution biosensors.
Greya reticulatus
Greya reticulatus is a species of moth in the family Prodoxidae, a group commonly known as yucca moths. The genus Greya comprises small, often inconspicuous moths that are primarily associated with herbaceous plants rather than the woody yuccas that characterize most prodoxid diversity. Species in this genus have been subjects of study regarding plant-insect coevolution and pollination ecology. The specific epithet "reticulatus" refers to a net-like or reticulated pattern, presumably of wing markings.
Hedychrum
Hedychrum is a large genus of cuckoo wasps in the family Chrysididae, comprising roughly 150 species and ranking as the second largest genus in the family. Most species occur in the Palaearctic region, with additional species present in the Oriental, Afrotropical, Nearctic, and Neotropical regions. These wasps are obligate brood parasites of ground-nesting apoid wasps, primarily in the subfamily Philanthinae.
Heliconiinae
longwings, heliconians, fritillaries and longwings
Heliconiinae is a subfamily of brush-footed butterflies (Nymphalidae) comprising 45–50 genera, commonly known as longwings or heliconians. Members are distinguished by elongated forewings and predominantly reddish-black coloration. They are notable among butterflies for actively consuming pollen, which extends adult longevity. The subfamily exhibits complex coevolutionary relationships with Passifloraceae host plants and serves as a classic model for studies of Müllerian and Batesian mimicry.
Heliconius
longwings, heliconians
Heliconius is a genus of brush-footed butterflies renowned for their bright aposematic wing patterns and Müllerian mimicry rings. Adults are unique among butterflies in their ability to collect and digest pollen, which provides essential amino acids enabling lifespans of up to six months—far exceeding most butterflies. The genus serves as a premier model system for studying speciation, mimicry evolution, and coevolutionary dynamics with their host plants. Approximately 45–50 species occur across the Neotropics, exhibiting complex patterns of wing color convergence and divergence.
Hemileuca eglanterina
Sheep Moth, Common Sheep Moth
Hemileuca eglanterina, commonly known as the sheep moth or common sheep moth, is a day-flying saturniid moth native to western North America. The species exhibits remarkable geographic variation in coloration, ranging from nearly black to pink/yellow to the typical orange with black markings. Adults are short-lived and lack functional mouthparts, relying entirely on fat reserves accumulated during the larval stage. The species has been documented in coevolutionary dynamics with Argiope spiders, which appear to lure male moths using chemical mimicry of female moth sex pheromones.
Matsucoccidae
pine bast scales, pine needle scales
Matsucoccidae is a family of scale insects (Hemiptera) comprising over 40 described species in at least 2 genera. The family was historically classified within Margarodidae but was elevated to family status based on distinct morphological and biological characteristics. Members are specialized feeders on conifers in the genus Pinus, with several species serving as economically significant forest pests. The family exhibits complex life cycles involving winged males and sessile females, and produces sex pheromones that function as kairomones attracting diverse predatory insects.
Nycteribiidae
bat flies, nycteribiid bat flies
Nycteribiidae is a family of Diptera in the superfamily Hippoboscoidea, commonly known as bat flies. These are obligate ectoparasites of bats, feeding exclusively on blood. They exhibit extreme morphological adaptations for parasitism: complete wing loss, highly reduced or absent compound eyes, and a dorsoventrally flattened, spider-like body with backward-folded legs. The family contains approximately 274 described species distributed primarily in the Old World tropics, with some species in the Neotropics and Europe. Most species show high host specificity, often with one-to-one associations with particular bat species.
Parasitiformes
mites and ticks
Parasitiformes is a superorder of arachnids constituting one of two major mite lineages, alongside Acariformes. The group includes ticks (Ixodida), the highly diverse predatory and parasitic Mesostigmata, the small scavenging Holothyrida, and the large segmented Opilioacarida. Over 12,000 species have been described, with estimates of 100,000–200,000 total species. Recent phylogenomic analyses indicate Parasitiformes is more closely related to other chelicerates (spiders, scorpions, horseshoe crabs) than to Acariformes, challenging the traditional unification of all 'mites' as a single natural group.
Poecilochirus
Carrion Beetle Mites
Poecilochirus is a Holarctic genus of relatively large mites (0.5–1 mm) in the family Parasitidae. They are phoretic symbionts of burying beetles (Silphidae: Nicrophorus), using beetles for transport to vertebrate carcasses where they feed and reproduce. The genus exhibits extensive cryptic diversity, with molecular studies identifying 24 genetic clusters potentially representing distinct species. Their interaction with beetle hosts is context-dependent, ranging from antagonistic (egg predation, competition) to potentially mutualistic (preying on fly eggs and nematodes).
Prodoxidae
Yucca Moths and Allies
Prodoxidae is a family of small, primitive monotrysian moths within the superfamily Adeloidea. The family includes approximately 90 described species across 12 genera, with members distributed primarily in the Northern Hemisphere. While some species such as the currant shoot borer (Lampronia capitella) are minor agricultural pests, the family is best known for the yucca moths—genera Tegeticula, Parategeticula, and Prodoxus—which exhibit one of the most extensively studied obligate pollination mutualisms in evolutionary biology. Female yucca moths possess unique tentacle-like mouthpart modifications for active pollen collection and intentional pollination of their host plants. The family occupies a basal phylogenetic position within Lepidoptera, making it significant for understanding the early evolution of the order.
Prodoxinae
Yucca Moths
Prodoxinae is a subfamily of small moths within the family Prodoxidae, commonly known as yucca moths. This group is best known for the obligate mutualism between yucca moths (genus Tegeticula) and their host plants in the genus Yucca. The moths are the sole pollinators of yucca flowers, while the developing seeds serve as the exclusive food source for moth larvae. This coevolutionary relationship represents one of the classic examples of obligate mutualism in insects.
Prodoxus
Bogus Yucca Moths
Prodoxus is a genus of moths in the family Prodoxidae comprising approximately 22 species, commonly known as bogus yucca moths. Unlike their pollinating relatives in the genus Tegeticula, Prodoxus species are non-pollinating yucca moths that exploit yucca plants as hosts without providing mutualistic services. The genus exhibits diverse feeding specializations, with larvae feeding on yucca flowering stalks, fruits, or leaves depending on the species. Host-associated divergence and incipient speciation have been documented, with specialization closely tied to host plant shifts.
Prodoxus decipiens
Bogus Yucca Moth
Prodoxus decipiens is a bogus yucca moth (family Prodoxidae) that specializes on Yucca host plants. Unlike mutualistic yucca moths, this species does not pollinate its host but instead lays eggs in flowering stalks where larvae develop internally. Host use is constrained by both ecological availability and intrinsic host plant characteristics; larvae successfully complete development only on specific Yucca species. The species exhibits strong host specialization, with populations typically associated with particular Yucca hosts such as Yucca filamentosa.
Psocodea
bark lice, book lice, parasitic lice, barklice, booklice
Psocodea is an order of insects comprising approximately 11,000 species across three extant suborders: Trogiomorpha, Troctomorpha, and Psocomorpha. The group includes free-living barklice and booklice, as well as the parasitic lice (formerly Phthiraptera). Molecular and morphological evidence demonstrates that parasitic lice evolved from within the barklice lineage, rendering the former order 'Psocoptera' paraphyletic. Members range from 1–10 mm in length and occupy diverse habitats from forest canopies to stored products.
Pterygosomatidae
scale mites
Pterygosomatidae is a family of prostigmatic mites in the order Trombidiformes, commonly known as scale mites. These mites are obligate ectoparasites of lizards, primarily infesting the skin and scales of their reptilian hosts. The family includes multiple genera, notably Pterygosoma, Geckobia, Geckobiella, Pimeliaphilus, and Neopterygosoma. Species within this family exhibit high host specificity, with particular mite lineages associated with specific lizard families including Agamidae, Gekkonidae, Liolaemidae, and Gerrhosauridae.
Rothschildia lebeau
Lebeau's Silk Moth, Forbes Moth
Rothschildia lebeau is a giant silkmoth in the family Saturniidae, distributed from the Lower Rio Grande Valley through eastern Mexico to Brazil. Adults are short-lived and do not feed, relying on fat reserves accumulated during larval development. The species exhibits a life history tradeoff between larval survival and adult body size based on host plant quality, with larger males showing higher mating success.
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n-accessopen-sourceFAIR-datafindableaccessibleinteroperablereusabledata-management-planmetadataprovenancequality-assurancequality-controlvalidationverificationuncertainty-quantificationerror-propagationsensitivity-analysisscenario-analysisMonte-Carlo-simulationBayesian-belief-networkinfluence-diagramdecision-treemulti-criteria-decision-analysiscost-benefit-analysiscost-effectiveness-analysismulti-attribute-utility-theorygame-theorymechanism-designincentive-compatibilityrevelation-principlesocial-choice-theoryvoting-theoryfair-divisioninstitutional-designpolicy-designgovernance-designarchitecturepolycentric-governanceadaptive-governancecollaborative-governancenetwork-governancehierarchical-governancemarket-governancecommunity-governancehybrid-governancemeta-governancesteeringrowingregulatory-stateenabling-statepartner-stateentrepreneurial-stateinnovation-statestrategic-statedevelopmental-statewelfare-stategreen-statesustainable-stateresilient-statesmart-statedigital-stateplatform-statealgorithmic-statesurveillance-stateauthoritarian-statedemocratic-statedeliberative-democracyparticipatory-democracydirect-democracyrepresentative-democracyliberal-democracysocial-democracygreen-democracyecological-democracybiocracyecofascismdeep-ecologysocial-ecologyecofeminismenvironmental-justiceclimate-justiceintergenerational-justiceintragenerational-justicespatial-justiceterritorial-justicerecognition-justiceprocedural-justicedistributive-justicerestorative-justicetransformative-justicereparative-justicetransitional-justiceecological-justicespecies-justicemultispecies-justicemore-than-human-justiceanthropocenecapitaloceneplantationocenechthulucenesymbiocenegood-anthropocenebad-anthropocenegreat-accelerationgreat-divergencegreat-convergencegreat-transformationgreat-decouplinggreat-disruptiongreat-moderationgreat-recessiongreat-depressiongreat-wargreat-faminegreat-migrationgreat-replacementgreat-resetgreat-refusalgreat-enrichmentgreat-levelinggreat-compressionSphingidae
sphinx moths, hawk moths, hornworms, hummingbird moths
The Sphingidae are a family of moths comprising approximately 1,450 species, best represented in tropical regions but occurring worldwide. Members are moderate to large in size with exceptional flight capabilities, including sustained hovering and rapid, agile maneuvering that rivals hummingbirds. Their streamlined bodies and narrow wings represent convergent evolution with nectar-feeding birds and bats. The family includes economically significant species: some adults are important pollinators with extremely long proboscises, while larval hornworms are major agricultural pests on solanaceous crops.
Stictocephala stimulea
Stictocephala stimulea is a treehopper species in the family Membracidae, first described by Van Duzee in 1914. Treehoppers in this genus are characterized by their enlarged, often ornate pronotum that extends over the body like a helmet or thorn. The specific epithet 'stimulea' suggests possible resemblance to or association with stinging organisms, though this refers to appearance rather than actual sting. Like other membracids, this species likely feeds on plant phloem sap using piercing-sucking mouthparts.
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yucca moths
Tegeticula is a genus of moths in the family Prodoxidae, commonly known as yucca moths. These insects are obligate mutualistic pollinators of various Yucca and Hesperoyucca species, representing one of the classic examples of coevolution between plants and insects. The genus includes both pollinating species and derived 'cheater' species that do not pollinate their hosts. Female moths possess specialized tentacles for collecting and depositing pollen, and their ovipositors are adapted to specific host plant structures.
Tegeticula maculata
Chaparral yucca moth
Tegeticula maculata is a small moth in the family Prodoxidae, commonly known as the chaparral yucca moth. It is the sole pollinator of Hesperoyucca whipplei, forming an obligate mutualism in which the moth's larvae feed exclusively on developing seeds of this host plant. The species exhibits high genetic structure across its range with limited morphological divergence, suggesting ancient lineage sorting rather than recent speciation.
Tegeticula yuccasella
yucca moth, Eastern Yucca Moth
Tegeticula yuccasella is a small moth in the family Prodoxidae, first described by Charles Valentine Riley in 1872. It is the nominate species of the T. yuccasella species complex, a group of obligate pollinators of yucca plants (Yucca spp.). The moth engages in active pollination behavior unique among Lepidoptera: females collect pollen with specialized tentacle-like mouthparts and deliberately deposit it on yucca stigmas to ensure fruit set before laying eggs. Larvae feed exclusively on developing yucca seeds. Recent systematic revision has split this formerly broadly-defined species into multiple distinct species, with T. yuccasella sensu stricto associated primarily with eastern North American yucca hosts.
Tetramesa
Tetramesa is a genus of minute phytophagous wasps in the family Eurytomidae comprising over 200 described species. Members are exclusively associated with grasses (Poaceae), where they typically induce stem or inflorescence galls. The genus exhibits pronounced host specificity, with most species restricted to a single grass species or closely related congeners. Several species have been deployed as biological control agents against invasive grasses, including T. romana for giant reed (Arundo donax) and candidate agents for medusahead (Taeniatherum caput-medusae) and African lovegrass (Eragrostis curvula). Adults feed on nectar. Recent phylogenomic studies have synonymized the genera Aiolomorphus and Cathilaria within Tetramesa.
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Milkweed Longhorn Beetles
Tetraopes is a genus of specialized longhorn beetles (Cerambycidae: Lamiinae) distributed from Guatemala to Canada. The genus is defined by extreme host specificity to milkweeds (Asclepias and related genera in Apocynaceae), with larvae feeding on roots and adults feeding on leaves and flowers. All species exhibit aposematic red-and-black coloration advertising their sequestration of cardiac glycosides from host plants. The eyes are completely divided by antennal insertions, giving rise to both the scientific name ("four eyes") and common name "four-eyed milkweed beetle" for some species. The genus represents a classic example of coevolutionary radiation with its host plants.
Tetraopes tetrophthalmus
Red Milkweed Beetle
Tetraopes tetrophthalmus, commonly known as the red milkweed beetle, is a longhorn beetle (Cerambycidae) specialized on milkweed host plants. Both its genus and species names derive from Ancient Greek for "four eyes," referring to the extreme adaptation where antennal bases bisect the compound eyes. Adults are aposematically colored in red and black, warning predators of their toxicity derived from sequestered cardiac glycosides. The species exhibits specialized behaviors including vein-cutting to disable latex defenses, vibrational communication for mating and territorial contests, and substrate-borne signaling distinct for different social contexts.
Trachymyrmex arizonensis
Arizona Fungus-farming Ant
Trachymyrmex arizonensis is a higher-attine fungus-growing ant native to the southwestern United States and Mexico. The species cultivates symbiotic fungi as its primary food source, showing strong partner fidelity with its fungal cultivar lineage. Phylogenetic studies reveal a pattern of 'one-to-one with some exceptions' in its ant-fungal specificity, with high fidelity to its own fungal subclade and rare instances of horizontal fungal transfer between colonies. Males exhibit distinctive morphology with reduced head size relative to body, a trait shared with other attine ants.
Trichobaris
flower weevils
Trichobaris is a genus of flower weevils in the family Curculionidae, comprising 8–13 described species distributed from the southwestern United States through Mexico. These weevils are specialized herbivores associated with plants in the family Solanaceae, particularly the genus Datura, with which they likely coevolved. Several species are notable agricultural pests, including T. mucorea (tobacco stalk borer) and T. trinotata (potato stalk borer), which also feed on cultivated solanaceous crops. Research on this genus has contributed to understanding host-herbivore coevolution, chemical ecology of host selection, and the decoupling of maternal oviposition cues from larval performance.