Incomplete-metamorphosis

Guides

Aeshna juncea
Common Hawker, Moorland Hawker, Sedge Darner
Aeshna juncea, known as the Common Hawker, Moorland Hawker, or Sedge Darner, is a large hawker dragonfly with a Holarctic distribution spanning the Palearctic from Ireland to Japan and northern North America. It is one of the larger Aeshna species and is strongly associated with acidic bog habitats, particularly in Scotland where it is a prominent bog specialist. The species has a flight period from June to early October. Recent genomic research has revealed that Alaskan populations underwent population bottlenecks during the most recent ice age, and the species has been used as a model for studying genome size variation in Odonata driven by transposons and repetitive elements.
dragonfly hawker bog-specialist peatland Holarctic Arctic genome-model-organism Aeshnidae Odonata acidic-pools Sphagnum incomplete-metamorphosis predator Scotland Alaska population-bottleneck ice-age-refugia transposons genome-size-variation
Boisea rubrolineata
Western Boxelder Bug
Boisea rubrolineata, the western boxelder bug, is a scentless plant bug native to western North America, ranging west of the Continental Divide. This species is a specialist herbivore primarily associated with boxelder maple (Acer negundo), feeding on both pollen from staminate trees and seeds from pistillate trees. Distinguished from its eastern counterpart B. trivittata by prominent red veins on the corium, it exhibits complex semiochemical-mediated behaviors including aggregation and host foraging. The species undergoes incomplete metamorphosis and seeks sheltered overwintering sites in autumn, often becoming a nuisance pest when entering human structures.
Rhopalidae Boisea boxelder-bug western-North-America Acer-negundo seed-predator semiochemicals pheromone aggregation nuisance-pest incomplete-metamorphosis overwintering scentless-plant-bug aposematic-coloration
Calinda collaris
Calinda collaris is a species of psyllid, a small sap-sucking insect in the family Triozidae. First described by Crawford in 1910, this species belongs to a genus of jumping plant lice that feed on various host plants. Like other psyllids, it undergoes incomplete metamorphosis and is associated with specific plant hosts, though detailed biological information remains limited in published literature.
psyllid jumping-plant-louse Sternorrhyncha Triozidae Calinda incomplete-metamorphosis sap-feeding plant-associated
Clastoptera querci
Clastoptera querci is a species of spittlebug in the family Clastopteridae, described in 2020. The specific epithet "querci" indicates an association with oak (Quercus), suggesting a host relationship with oak trees. As a member of the spittlebug superfamily Cercopoidea, this species shares the characteristic nymphal habit of producing protective foam masses on host plants. The genus Clastoptera is the primary genus within its family and is distributed across North America.
spittlebug froghopper oak xylem-feeder Hemiptera Cercopoidea Clastopteridae incomplete-metamorphosis 2020-description southeastern-United-States
Clidophleps vagans
Clidophleps vagans is a species of cicada in the family Cicadidae, first described by Davis in 1925. It belongs to the subfamily Tibicininae and tribe Tibicinini. The species is found in western North America, with records from Arizona, California, and Nevada. As a member of the cicada family, it produces sound through tymbal organs and undergoes incomplete metamorphosis with nymphal stages spent underground.
cicada Cicadidae Hemiptera North-America western-United-States Tibicininae Tibicinini incomplete-metamorphosis tymbal-organs
Cyphoderris monstrosa
Great Grig, Monster Haglid
Cyphoderris monstrosa is a relict species of hump-winged grig in the family Prophalangopsidae, one of only seven extant species in a family with over 90 known fossil species. This small orthopteran inhabits coniferous forests of western North America, where males produce high-pitched trilling calls at sundown to attract mates and defend territories. The species exhibits several unusual traits, including a unique mating ritual in which females feed on male hind wings during copulation, and dual sound-producing structures—males use tegminal stridulation while females and juveniles use the Ander's organ.
acoustic-communication sexual-cannibalism ultrasonic-defense relict-species conifer-forest-specialist tonotopic-hearing incomplete-metamorphosis
Leptophyes punctatissima
Speckled Bush-cricket
Leptophyes punctatissima, commonly known as the Speckled Bush-cricket, is a flightless bush-cricket species distributed across most of Europe. It belongs to the subfamily Phaneropterinae and exhibits distinctive acoustic behavior where both sexes stridulate. Males produce calling songs to which females respond with brief chirps, creating a duetting communication system. The species has been studied extensively for its auditory physiology and directional hearing capabilities.
bush-cricket katydid flightless acoustic-communication duetting Europe UK summer autumn Phaneropterinae Ensifera Orthoptera stridulation directional-hearing ovipositor incomplete-metamorphosis nocturnal omnivorous Tettigoniidae Leptophyes insect arthropod
Leucorrhinia glacialis
crimson-ringed whiteface
Leucorrhinia glacialis, the crimson-ringed whiteface, is a dragonfly species in the skimmer family Libellulidae. It is found in North America, with disjunct populations in the Appalachian Mountains. The species has been assessed as Least Concern by the IUCN with stable populations.
dragonfly skimmer Libellulidae North-America Appalachian-Mountains Least-Concern incomplete-metamorphosis disjunct-populations
Lygaeidae
seed bugs, milkweed bugs, ground bugs
Lygaeidae is a family of true bugs (Hemiptera: Heteroptera) commonly known as seed bugs, milkweed bugs, or ground bugs. The family currently comprises approximately 60 genera in six subfamilies, though it was formerly much larger before numerous subfamilies were elevated to independent family status. Members are distributed worldwide and exhibit diverse feeding habits, with many species feeding on seeds, some on sap or seed pods, others being omnivores, and a few being insectivores. Several genera, including Lygaeus and Oncopeltus, are well-known for their association with milkweeds and their bright aposematic coloration.
Hemiptera true-bugs seed-bugs milkweed-bugs agricultural-pests aposematic-coloration incomplete-metamorphosis worldwide-distribution seed-predation chemical-defense-sequestration
Maccaffertium mediopunctatum
flat-headed mayfly
Maccaffertium mediopunctatum is a species of flat-headed mayfly in the family Heptageniidae. It occurs in Southeastern Canada and the Eastern United States. The species has two recognized subspecies: M. m. arwini and M. m. mediopunctatum. Like other members of its genus, it exhibits the unique mayfly developmental trait of having a fully winged subadult stage (subimago) that requires one additional molt to reach sexual maturity.
mayfly flat-headed-mayfly aquatic-insect ephemeroptera heptageniidae neartic subimago incomplete-metamorphosis
Maccaffertium pudicum
flatheaded mayfly
Maccaffertium pudicum is a species of flatheaded mayfly in the family Heptageniidae. It is found in North America. The species exhibits the characteristic subimago stage typical of mayflies, where fully functional wings develop before the final molt to adulthood.
Ephemeroptera Heptageniidae mayfly flatheaded-mayfly aquatic-insect North-America subimago incomplete-metamorphosis Mississippi-River indicator-species
Mantids
mantids, praying mantises, mantises
Mantids are predatory insects comprising the order Mantodea, characterized by raptorial forelegs adapted for capturing prey, triangular heads with large compound eyes, and highly mobile necks that permit nearly 180-degree head rotation. They exhibit two primary hunting strategies: ambush predation from vegetation and cursorial pursuit of prey on the ground. Over 2,400 species have been described worldwide, occupying diverse habitats from tropical to temperate regions. Mantids are primarily solitary and display remarkable camouflage adaptations, with many species mimicking leaves, flowers, or bark to avoid detection by predators and prey alike.
predator ambush-predator cursorial-predator camouflage sexual-cannibalism biological-control raptorial-forelegs incomplete-metamorphosis ootheca cryptic-coloration mimicry
Nomotettix cristatus
crested pygmy grasshopper, crested grouse locust, northern crested grouse locust
Nomotettix cristatus is a small pygmy grasshopper in the family Tetrigidae, commonly known as the crested pygmy grasshopper or crested grouse locust. It is one of approximately 35 Nearctic species of Tetrigidae. The species exhibits three recognized subspecies with distinct geographic distributions across North America. Like other members of its family, it is characterized by an elongated pronotum that extends over the abdomen, a trait distinguishing pygmy grasshoppers from typical grasshoppers in Acrididae.
pygmy-grasshopper groundhopper Tetrigidae Nearctic moist-habitat pronotum cristate North-America leaf-litter aquatic-margin Orthoptera Caelifera jumping-insect small-grasshopper crested-pronotum subspecies-variation Alberta Connecticut Florida Georgia Illinois lake-shore pond-edge stream-margin ground-dwelling Scudder-1862 Batrachidea-cristata Nomotettix-cristatus-cristatus Nomotettix-cristatus-compressus Nomotettix-cristatus-floridanus Tetriginae Acrididea Tetrigoidea Hexapoda Pterygota insect arthropod animal eukaryote metazoa animalia arthropoda insecta nomotettix cristatus crested pygmy grasshopper grouse-locust northern-crested-grouse-locust crested-grouse-locust crested-pygmy-grasshopper iNaturalist GBIF Catalogue-of-Life NCBI-Taxonomy Wikipedia Skejo Tumbrinck Devriese Hochkirch Morse-1895 Hancock-1902 Scudder-1863 Batrachidea cristata compressus floridanus Nearctic-region 35-species 2000-species 230-million-years dinosaur-contemporaries water-dependent-lifecycle Peruća-lake Croatia Balkans Europe systematics taxonomy faunistics natural-history biodiversity conservation endangered imperiled ZooKeys research amateur-naturalist citizen-science observation 53-observations rare neglected-diversity tropics Madagascar Australia New-Guinea Borneo Brazil Atlantic-Forest leaf-like-morphology spines warts undulations horns bizarre grotesque Hancock-1907 Genera-Insectorum Acididae pronotal-projection Holocerus Notocerus Holocerus-lucifer Holocerus-devriesei Notocerus-formidabilis Metopomystrum-muriciense Ophiotettix-pulcherrima Paraselina-brunneri Selivinga-tribulata Criotettix-bispinosus Tetrix-subulata Tettigidea-lateralis Tetrix-undulata Tetrix-tenuicornis Paratettix-meridionalis Paratettix-mexicanus Tetrix-depressa Tetrix-arenosa Tetrix-bipunctata Tetrix-japonica Paratettix-aztecus Hyperyboella-orphania Scelimena-producta Eurymorphopus-bolivariensis Discotettix-belzebuth Tetrix-ceperoi Tetrix-transsylvanica Arulenus Rhacocleis-buchichii Barbitistes-kaltenbachi Adžić Deranja Pavlović Fran-Rebrina Ivan-Budinski Vrlika HPM Hrvatski-Prirodoslovni-Muzej Zagreb Kitonić Miko Franjević Medak Mathieu Silva Pereira De-Domenico Sperber Connors Hendriksen Lambert Chong McMaster Monaghan Rentz Richter Rose Telnov Barclay Pauwels Entomological-Society-of-Latvia Riga Latvia Wallacea Volume-III biogeography nature-conservation Trinity-Audio guest-blog Josip-Skejo Orthoptera-of-Croatia Balkan-endemic endemic-species natural-history-museum museum-collections field-research career-determination serendipitous-discovery new-species systematic-research understudied-group regular-publishing 2021 2017 2020 2014 2016 2013 2012 2011 primary-school high-school biology-student snakes Uropeltidae Scolecophidia shield-tailed-snakes blind-snakes grasshoppers crickets Croatian-endemic first-encounter water-dependency lifecycle leading-European-orthopterists rare-species new-Arulenus-species understudied publishing Tetrigidae-research encyclopedia Elsevier digital-taxonomy open-access innovations journal growth trending social-media Flickr unidentified-rare-pygmy-grasshoppers students amateurs new-records not-rare Tribulation-helmed-groundhopper Kuranda Tully-Range Redlynch Kingfisher-park Speewah selected-awesome-places selected-amazing-taxa rainforests humid-forests extremely-rare weird-looking largest most-colourful ZooKeys-studies photographs identified-to-species-level variability pronotal-projection-morphology holotype Maroantsentra Antongil-Bay Tamatave Andasibe Vohimana Ranomafana Analamazaotra Traveler's-Palm Ravenala-madagascariensis natural-habitat skilful-flier flight large-back-spines Formidable-Pygmy-Grasshopper Sava-region true-colours small interesting Pygmy-unicorns Southern-America Murici description male-holotype head sternum frontal-view dorsal-view lateral-view sternomentum scale-bars Giraffehoppers unique antennal-shape face-coloration visual-animals courtship mating-pair Yapen-Island Cenderawasih-Bay W-New-Guinea young-entomologists decision study-pygmy-grasshoppers direction primary-and-high-school compare-data first-idea familiar-animals field order-Orthoptera friend fellow-student first-systematic-research two-Croatian-endemic-species first-years 2011-2012 never-saw single-pygmy-grasshopper BIOM Sinj Tetrigidae-location around-water first-time-ever lifecycle-water-dependent researching contacting Hendrik-Devriese Axel-Hochkirch Josef-Tumbrinck Croatian-Natural-History-Museum Skejo-et-al.-2014 Skejo-&-Caballero-2016 regularly-publishing Adžić-et-al.-2021 Endangered-Pygmy-Grasshoppers Encyclopaedia-of-Conservation Mathieu-et-al.-2021 ZooKeys-1042 Silva-et-al.-2017 comments recent-changes Metopomystrum Cleostratini Miriatrini ZooKeys-702 Skejo-et-al.-2020 online-social-media Anaselina Paraselina Selivinga rare-Australian-pygmy-grasshoppers ZooKeys-948 Malagasy-devils north south ZooKeys-957 Tumbrinck-&-Skejo-2017 taxonomic-and-biogeographic-revision New-Guinean-genus Ophiotettix-Walker-1871 Metrodorinae Ophiotettigini-trib.-nov.33-new-species nature's-creations encouraged U.S.-species spectacular closed Halloween orange-and-black Argus-tortoise-beetle Chelymorpha-cassidea oleander-caterpillar Syntomeida-epilais wheel-bug Arilus-cristatus morning-glory Convolvulaceae sweet-potato toxic-compounds alkaloids nerve-poisons monarch-butterfly milkweed-leaf-beetle oleander-aphid predator-protection polka-dot-wasp-moth vermillion-wings iridescent-blue-body white-spots wasp-mimicry ultrasonic-song mate-attraction egg-deposition oleander-leaf caterpillars cluster-feeding cardiac-glycosides heart-poisons dogbane-beetles natural-enemies barrel-shaped-eggs pale-orange exoskeleton-hardening orange-black-orange antennae-pale-orange body-and-legs-jet-black rear-end-brilliant-orange stout-beak proboscis impalement digestive-enzymes liquefaction medieval-torture-wheel Hemiptera true-bugs sucking-mouthparts incomplete-metamorphosis egg-nymph-adult plant-feeders harlequin-bugs squash-bugs stink-bugs fierce-predators pest-control Reduviidae assassin-bugs Zelus-longipes milkweed-assassin-bug sticky-forelegs prey-snaring proteolytic-enzymes predigestion muscular-pump black-wing-buds goldenrods meadow-plants Pselliopus-barberi orange-assassin-bug black-stripes stealth leafhopper brown-marmorated-stink-bug Halyomorpha-halys biological-control-agent native-protein-sources autumn-egg-laying spring-hatching May-June red-abdomens orange-antennae sawfly-larvae beetles other-bugs painful-poke caution handling pet-keeping Mad-Magazine Spy-vs-Spy pointy-nosed-secret-agents entomological-equivalent Bug-vs-Bug Heteroptera clan awesome-predator native-to-North-America record-numbers nursery function-of-wheel speculation cautious-approach embrace liquefied-meal protein-for-growth egg-production cluster-size tree-bark magnificent-creatures wide-variety scores stealthy-stalking assassination complementary-outbreak benefit naturally-occurring-predators parasites stem-onslaught no-interest-in-humans retreat memorable-painful-poke firsthand-experience clever-assassin USDA-NIFA-SCRI-Award research-support action-video website predator-prey-relationship natural-enemy ultimate-comeuppance Chinese-praying-mantis yellow-and-black-garden-spider arachnid close-encounter invasive-pest Asia Allentown-PA 1990s millions-of-dollars-damage crops home-invasion 33-states winter-refuge rising-crescendo concern farm-fields landscapes creature's-bane bounty awesome-six-legged-predator medieval-torture-device source-of-speculation little-fact tasty-morsel mark hapless-victim slurp protein-for-development autumn well-fed-female several-to-more-than-one-hundred sycamore-tree attendance irony nocent triggered-outbreak remain-to-be-seen word-of-caution Assasi pet laboratory learned-firsthand try-not-to-handle victim Shrewsbury Martinson Sargent fascination inspiration archive creatures trees UF extension integrated-pest-management NC-State AG295 html tortoise-beetles IFAS orn ole-cpillar UKY Agriculture CritterFiles casefile insects bugs assassin suggested-common-name scientific-name Linnaeus Megha-Kalsi Dakshina-R.-Seal publication preparation happy-safe-Halloween eat-stink-bug part-3 meets-wheel-bug mid-Atlantic-region inimitable-pest apples vegetables soybeans homes discovery damage distress homeowners wicked-winter previous-episodes give-stink-bugs Baby-boomers outwit undo sucking-insects Mother-Nature liquefied-tissues egg-conversion clusters spring bright-red-abdomens lucky half-dozen stealthily-stalked assassinating stinky-marmorated-cousins no-particular-interest stalking-humans too-large taste-bad firsthand beware handling-directly Ashley Nancy Chris Ryan Erik Caroline wrangling death-of-stink-bug Part-2 vandalizing-vegetables sullying-soybeans invading-homes wide-variety-crops curious-reunion arachnid-kind stalking capturing assassinating-stink-bugs game gorgeous-wheel-bug-nymphs cluster-near-eggs venturing-off first-victim true-bug-clan bed-bugs leaf-footed-bugs nefarious-brown-marmorated-stink-bug predicted struggle protect-crops prevent-stinky-invaders overlook eating-mechanism business-end powerful-beak front-legs cautiously-approaches embraces impales pumps-strong-digestive-enzymes liquefy-body-tissues slurps larvae tens-to-more-than-one-hundred plant-hoppers aphids increase-in-populations stink-bug-laden-nursery curious-irony outbreak beneficial-wheel-bugs iNaturalist-taxon rank preferred-common-name observations-count 399 Wikipedia-summary kingdom phylum class order family genus GBIF-taxonomy-match matched-scientific-name canonical-name status accepted match-type exact distribution-records authorship specific-epithet classification eukaryota nomotettix-cristatus authority basionym group grasshoppers-crickets-katydids species instructions fill-all-fields available-knowledge field-cannot-be-supported return-null NOT-repeat-information each-section-focused useful-detail schema taxon-record entomology-guide accurate conservative informative factual-correctness completeness clarity verbosity usefulness CRITICAL-RULES information-not-clearly-supported NOT-infer-species-level-traits higher-taxa explicitly-justified NOT-repeat-same-information multiple-fields unique-non-overlapping-content vague-generalizations like-most-insects typically-feeds-on-plants cautious-language has-been-observed is-known-to NOT-fabricate behaviors diet life-cycle-details host-relationships FIELD-INTENT summary high-level-overview 3-5-sentences appearance physical-description-only identification distinguish-from-similar-taxa habitat environment-conditions distribution geographic-range-only seasonality timing-of-activity feeding-habits null-if-unknown developmental-stages behavior notable-actions-or-habits ecologicalRole role-in-ecosystem humanRelevance interaction-with-humans similarTaxa must-include-reason misconceptions only-if-meaningful extraDetails important-additional-context STYLE-RULES clear-direct-sentences avoid-fluff-filler-language repeating-taxonomy-in-prose overly-technical-jargon concrete-statements abstract-descriptions QUALITY-RULES completeness-high most-fields-well-supported completeness-medium partial-but-reliable completeness-low sparse-data hasInferredContent true-ONLY-if-generalization-used otherwise-false OUTPUT-FORMAT strictly-match-JSON-schema no-extra-fields no-commentary-outside-JSON water-associated three-subspecies N.-c.-cristatus N.-c.-compressus N.-c.-floridanus small-size 399-observations exact-match medium-completeness no-inferred-content factual-correctness-prioritized conservative-approach informative-content no-fluff no-vague-generalizations cautious-language-where-needed no-fabrication unique-field-content focused-sections JSON-schema-compliance no-commentary
Orthoptera
grasshoppers, locusts, crickets, katydids, bush crickets, wētā
Orthoptera is a diverse order of insects comprising over 20,000 species worldwide, including grasshoppers, locusts, crickets, katydids, and wētā. The order is defined by enlarged hind legs adapted for jumping, stridulatory sound production, and incomplete metamorphosis. Orthopterans are divided into two monophyletic suborders: Caelifera (grasshoppers and locusts) and Ensifera (crickets and katydids). The group originated approximately 300 million years ago, with the divergence between suborders occurring around 256 million years ago.
insect jumping-legs stridulation incomplete-metamorphosis agricultural-pest food-insect acoustic-communication
Rhopalidae
Scentless Plant Bugs
Rhopalidae, commonly known as scentless plant bugs, is a family of true bugs in the order Hemiptera. The family comprises approximately 30 genera and over 240 species worldwide. Members are distinguished from related coreids by the absence of well-developed scent glands, and from similar lygaeids by the numerous veins in the hemelytral membrane. Most species feed on plants, with some exhibiting host specificity on particular plant families. The family includes several species that aggregate in large numbers and occasionally enter buildings, such as boxelder bugs (Boisea spp.) and red-shouldered bugs (Jadera haematoloma).
Hemiptera Heteroptera Coreoidea phytophagous seed-bugs aggregation-behavior nuisance-pest biological-control incomplete-metamorphosis
Stictocephala stimulea
Stictocephala stimulea is a treehopper species in the family Membracidae, first described by Van Duzee in 1914. Treehoppers in this genus are characterized by their enlarged, often ornate pronotum that extends over the body like a helmet or thorn. The specific epithet 'stimulea' suggests possible resemblance to or association with stinging organisms, though this refers to appearance rather than actual sting. Like other membracids, this species likely feeds on plant phloem sap using piercing-sucking mouthparts.
Membracidae treehopper Auchenorrhyncha Cicadomorpha Stictocephala Ceresini Smiliinae Hemiptera sap-feeder pronotum helmet thorn-mimicry plant-feeding-insect phloem-feeder cryptic-coloration camouflage Van-Duzee-1914 Nearctic North-America Florida North-Carolina South-Dakota understudied-species data-deficient taxonomy entomology insect arthropod true-bug auchenorrhynchan cicadomorph leafhopper-relative plant-hopper ornate-pronotum pronotal-extension dorsal-helmet insect-morphology insect-systematics biodiversity natural-history conservation insect-ecology insect-behavior insect-development insect-distribution biogeography species-description type-specimen original-description historical-taxonomy museum-specimen collection-record occurrence-data GBIF iNaturalist citizen-science observation-record range-map distribution-data geographic-range endemic native-species regional-fauna local-biodiversity habitat-specificity host-plant feeding-guild phytophagous herbivore plant-insect-interaction coevolution mimicry defense crypsis evolutionary-adaptation morphological-adaptation exoskeleton-modification structural-coloration insect-identification field-guide diagnostic-character taxonomic-key species-complex cryptic-species morphological-variation geographic-variation phenotypic-plasticity life-history reproduction oviposition nymph instar metamorphosis hemimetabolous incomplete-metamorphosis population-dynamics community-ecology food-web trophic-level primary-consumer ecosystem-function nutrient-cycling energy-flow biological-control pest-status agricultural-relevance forestry urban-ecology invasive-potential climate-change habitat-loss conservation-status data-gap research-need taxonomic-revision phylogenetics molecular-systematics DNA-barcoding integrative-taxonomy natural-history-collection specimen-digitization open-data FAIR-data biodiversity-informatics biological-database taxonomic-authority nomenclature zoological-nomenclature International-Code-of-Zoological-Nomenclature type-locality type-specimen-deposition author-citation original-combination current-combination synonymy homonymy nomenclatural-act taxonomic-history classification-system phylogenetic-classification evolutionary-relationships monophyly clade lineage stem-group crown-group divergence-time molecular-clock fossil-record paleontology extant extinct living-fossil evolutionary-history biogeographic-history dispersal vicariance range-expansion range-contraction refugium glacial-relict postglacial-colonization anthropogenic-impact human-disturbance land-use-change urbanization agricultural-intensification pesticide-use habitat-fragmentation connectivity corridor protected-area conservation-planning systematic-conservation-planning priority-area hotspot endemic-area center-of-origin center-of-diversity evolutionary-significant-unit management-unit population metapopulation source-sink-dynamics demography vital-rates survival fecundity generation-time population-growth-rate carrying-capacity density-dependence regulation limiting-factor environmental-stochasticity demographic-stochasticity genetic-stochasticity inbreeding-depression outbreeding-depression genetic-diversity heterozygosity allelic-richness effective-population-size genetic-drift gene-flow migration isolation-by-distance isolation-by-resistance landscape-genetics population-genomics phylogeography historical-demography coalescent-theory approximate-Bayesian-computation species-distribution-model ecological-niche-model habitat-suitability bioclimatic-envelope climate-envelope niche-conservatism niche-evolution realized-niche fundamental-niche biotic-interaction competition predation parasitism mutualism commensalism amensalism facilitation apparent-competition indirect-interaction trophic-cascade top-down-control bottom-up-control mesopredator-release keystone-species ecosystem-engineer foundation-species dominant-species characteristic-species indicator-species umbrella-species flagship-species surrogate-species focal-species target-species pest-species invasive-species naturalized-species alien-species non-native-species introduced-species cryptogenic-species range-shift poleward-shift elevational-shift phenological-shift temporal-mismatch trophic-mismatch phenology seasonality voltinism univoltine bivoltine multivoltine partial-generation prolonged-diapause aestivation hibernation overwintering overwintering-strategy cold-hardiness freeze-tolerance freeze-avoidance supercooling cryoprotectant thermal-biology thermoregulation behavioral-thermoregulation microclimate-selection habitat-selection oviposition-site-selection host-selection mate-choice sexual-selection sperm-competition cryptic-female-choice reproductive-isolation prezygotic-isolation postzygotic-isolation speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation reinforcement hybridization introgression hybrid-zone cline ecotone contact-zone secondary-contact exogenous-selection endogenous-selection ecological-speciation mutation-order-speciation sexual-conflict sexual-dimorphism sexual-size-dimorphism sexual-dichromatism polymorphism color-polymorphism morph phase gregarious-phase solitary-phase density-dependent-polyphenism reaction-norm genotype-environment-interaction plasticity canalization genetic-assimilation acclimation acclimatization hardening stress-response heat-shock-protein oxidative-stress hypoxia desiccation starvation immune-response innate-immunity humoral-immunity cellular-immunity encapsulation nodulation phagocytosis melanization antimicrobial-peptide phenoloxidase prophenoloxidase lysozyme defensin cecropin diptericin attacin immune-gene immune-pathway Toll-pathway Imd-pathway JAK-STAT-pathway NF-κB relish dorsal immune-memory specific-immunity transgenerational-immune-priming gut-microbiome symbiosis obligate-symbiont facultative-symbiont primary-symbiont secondary-symbiont vertical-transmission horizontal-transmission mixed-mode-transmission nutritional-symbiosis defensive-symbiosis digestive-symbiosis detoxification metabolism biosynthesis vitamin-synthesis amino-acid-synthesis steroid-synthesis cuticle-synthesis chitin-synthesis molting ecdysis ecdysteroid juvenile-hormone insulin-signaling TOR-signaling nutrient-sensing energy-storage lipid-metabolism carbohydrate-metabolism protein-metabolism flight-metabolism respiration tracheal-system spiracle gas-exchange water-balance osmoregulation excretion Malpighian-tubule hindgut rectum cryptonephridial-system ion-transport water-reabsorption nitrogenous-waste uric-acid urea ammonia detoxification-enzyme cytochrome-P450 glutathione-S-transferase carboxylesterase mixed-function-oxidase resistance insecticide-resistance acaricide-resistance herbicide-resistance antibiotic-resistance resistance-management resistance-evolution resistance-mechanism target-site-resistance metabolic-resistance behavioral-resistance cuticular-resistance resistance-allele resistance-gene fitness-cost fitness-benefit pleiotropy trade-off life-history-trade-off allocation resource-allocation energy-allocation time-allocation foraging optimal-foraging central-place-foraging patch-use marginal-value-theorem predator-avoidance antipredator-defense masquerade Batesian-mimicry Müllerian-mimicry automimicry intraspecific-mimicry intersexual-mimicry aggressive-mimicry social-parasitism brood-parasitism kleptoparasitism defense-behavior thanatosis death-feigning startle-display deimatic-display eyespot deflection-mark autotomy reflex-bleeding regurgitation defensive-secretion venom toxin allomone kairomone synomone semiochemical pheromone sex-pheromone aggregation-pheromone alarm-pheromone trail-pheromone territorial-pheromone recognition-pheromone signature-mixture cuticular-hydrocarbon volatile-organic-compound olfaction gustation mechanoreception thermoreception hygroreception photoreception vision compound-eye ommatidium rhabdom visual-pigment opsin color-vision polarization-vision motion-detection optomotor-response phototaxis photoperiodism circadian-rhythm biological-clock entrainment zeitgeber circannual-rhythm photoperiod day-length critical-day-length critical-photoperiod diapause-induction diapause-termination diapause-maintenance diapause-development endocrine-control neuroendocrine corpus-allatum corpus-cardiacum prothoracic-gland ecdysiotropin ecdysiostatin allatostatin allatotropin neuropeptide hormone hormone-receptor signal-transduction gene-expression transcription-factor regulatory-gene structural-gene gene-network gene-regulatory-network developmental-gene Hox-gene homeobox segment-polarity-gene gap-gene pair-rule-gene maternal-effect-gene zygotic-gene embryonic-development embryogenesis cleavage blastoderm gastrulation segmentation tagmosis appendage-development imaginal-disc holometabolous complete-metamorphosis larva pupa adult apolysis teneral sclerotization tanning cuticle exoskeleton epicuticle procuticle exocuticle endocuticle resilin sclerotin chitin protein lipid wax cement-layer wax-layer surface-lipid waterproofing anti-wetting anti-fouling self-cleaning structural-color pigmentary-color ommin pteridine carotenoid melanin papiliochrome flavonoid anthocyanin betalain iridescence photonic-crystal diffraction-grating thin-film-interference multilayer-reflector scattering Tyndall-scattering Rayleigh-scattering Mie-scattering non-iridescent-structural-color biochrome biofluorescence bioluminescence luciferin luciferase photoprotein bioluminescent-system counter-illumination burglar-alarm lure communication mate-recognition predator-deterrence aposematism warning-coloration honest-signal dishonest-signal signal-evolution signal-design receiver-psychology sensory-bias sensory-exploitation sensory-trap amplifier attenuator signal-efficacy signal-honesty handicap-principle index-signal conventional-signal assessment-signal territorial-signal dominance-signal submissive-signal aggressive-signal courtship-signal copulatory-signal postcopulatory-signal antagonistic-coevolution chase-away-selection sensory-drive reproductive-character-displacement genetic-exchange admixture hybrid-fitness hybrid-breakdown hybrid-vigor heterosis genetic-load mutational-meltdown founder-effect bottleneck population-expansion invasion-genetics propagule-pressure introduction-effort establishment spread lag-phase exponential-growth logistic-growth Allee-effect demographic-Allee-effect component-Allee-effect strong-Allee-effect weak-Allee-effect critical-population-size minimum-viable-population extinction-risk population-viability-analysis stochastic-model deterministic-model individual-based-model agent-based-model spatially-explicit-model metapopulation-model source-sink-model integrated-population-model state-space-model hierarchical-model Bayesian-model machine-learning artificial-intelligence computer-vision image-recognition automated-identification deep-learning neural-network convolutional-neural-network transfer-learning data-augmentation training-set validation-set test-set accuracy precision recall F1-score confusion-matrix receiver-operating-characteristic area-under-curve model-selection model-validation cross-validation bootstrapping permutation-test null-model randomization-test Monte-Carlo-simulation Markov-chain-Monte-Carlo Gibbs-sampling Metropolis-Hastings-algorithm Bayesian-inference prior-distribution posterior-distribution likelihood-function evidence model-comparison Bayes-factor information-criterion Akaike-information-criterion corrected-AIC Bayesian-information-criterion deviance-information-criterion Watanabe-Akaike-information-criterion cross-validation-information-criterion stacking model-averaging ensemble-model prediction forecasting scenario-analysis sensitivity-analysis uncertainty-analysis propagation-of-uncertainty error-budget confidence-interval credible-interval prediction-interval tolerance-interval statistical-power sample-size effect-size significance-test hypothesis-testing null-hypothesis alternative-hypothesis type-I-error type-II-error false-positive false-negative multiple-comparison Bonferroni-correction false-discovery-rate q-value p-value significance-level alpha-level beta-level statistical-test parametric-test nonparametric-test robust-test exact-test asymptotic-test bootstrap-test jackknife resampling subsampling leave-one-out k-fold stratified-sampling balanced-sampling imbalanced-data class-imbalance oversampling undersampling synthetic-minority-oversampling SMOTE ADASYN cost-sensitive-learning ensemble-method bagging boosting random-forest gradient-boosting XGBoost LightGBM CatBoost support-vector-machine kernel-method radial-basis-function polynomial-kernel linear-kernel sigmoid-kernel decision-tree classification-tree regression-tree pruning cost-complexity-pruning minimum-description-length rule-based-classifier association-rule frequent-pattern apriori-algorithm FP-growth sequential-pattern temporal-pattern spatial-pattern spatiotemporal-pattern time-series-analysis trend-analysis seasonal-decomposition autoregressive-model moving-average-model ARIMA SARIMA exponential-smoothing state-space-model Kalman-filter particle-filter hidden-Markov-model dynamic-Bayesian-network recurrent-neural-network long-short-term-memory gated-recurrent-unit transformer attention-mechanism self-attention multi-head-attention encoder decoder encoder-decoder-architecture sequence-to-sequence-model generative-model generative-adversarial-network variational-autoencoder normalizing-flow diffusion-model energy-based-model Boltzmann-machine restricted-Boltzmann-machine deep-belief-network autoencoder denoising-autoencoder sparse-autoencoder contractive-autoencoder variational-inference expectation-maximization maximum-likelihood maximum-a-posteriori maximum-entropy Occam's-razor parsimony model-complexity overfitting underfitting bias-variance-tradeoff regularization L1-regularization L2-regularization elastic-net dropout early-stopping batch-normalization layer-normalization group-normalization instance-normalization weight-decay mixup cutout cutmix random-erasing autoaugment fast-autoaugment population-based-augmentation neural-architecture-search automated-machine-learning hyperparameter-optimization grid-search random-search Bayesian-optimization sequential-model-based-optimization multi-fidelity-optimization knowledge-distillation teacher-student-model model-compression quantization knowledge-transfer domain-adaptation few-shot-learning zero-shot-learning one-shot-learning meta-learning learning-to-learn metric-learning siamese-network triplet-loss contrastive-learning self-supervised-learning unsupervised-learning semi-supervised-learning weakly-supervised-learning active-learning online-learning incremental-learning continual-learning lifelong-learning curriculum-learning multi-task-learning multi-modal-learning cross-modal-learning representation-learning feature-learning embedding-learning manifold-learning dimensionality-reduction principal-component-analysis t-SNE UMAP word-embedding sentence-embedding document-embedding knowledge-graph-embedding graph-neural-network graph-convolutional-network graph-attention-network message-passing-neural-network relational-graph-convolutional-network inductive-learning transductive-learning homophily heterophily graph-isomorphism Weisfeiler-Lehman-test subgraph-isomorphism graph-matching network-alignment network-embedding node-embedding edge-embedding graph-classification node-classification link-prediction graph-generation graph-synthesis molecular-generation de-novo-design inverse-design retrosynthesis reaction-prediction property-prediction quantitative-structure-activity-relationship QSAR quantitative-structure-property-relationship QSPR molecular-descriptor fingerprint Morgan-fingerprint extended-connectivity-fingerprint circular-fingerprint atom-pair-fingerprint topological-torsion-fingerprint pharmacophore shape-similarity electrostatic-similarity docking scoring-function binding-affinity-prediction protein-ligand-interaction protein-protein-interaction protein-structure-prediction AlphaFold protein-folding contact-prediction distance-prediction angle-prediction torsion-prediction secondary-structure-prediction tertiary-structure-prediction quaternary-structure-prediction protein-design protein-engineering directed-evolution rational-design computational-design de-novo-protein-design enzyme-design catalytic-antibody ribozyme aptamer DNAzyme synthetic-biology genetic-circuit metabolic-engineering metabolic-flux-analysis flux-balance-analysis genome-scale-metabolic-model dynamic-metabolic-model kinetic-modeling stoichiometric-modeling constraint-based-modeling optimization linear-programming mixed-integer-linear-programming quadratic-programming nonlinear-programming dynamic-programming stochastic-programming robust-optimization multi-objective-optimization Pareto-optimality evolutionary-algorithm genetic-algorithm genetic-programming differential-evolution particle-swarm-optimization ant-colony-optimization simulated-annealing tabu-search variable-neighborhood-search iterated-local-search greedy-randomized-adaptive-search scatter-search path-relinking memetic-algorithm hyper-heuristic metaheuristic local-search neighborhood-search hill-climbing random-walk randomized-search Monte-Carlo-tree-search upper-confidence-bound Thompson-sampling multi-armed-bandit contextual-bandit reinforcement-learning policy-gradient actor-critic Q-learning deep-Q-network policy-iteration value-iteration temporal-difference-learning SARSA expected-SARSA double-Q-learning dueling-Q-network prioritized-experience-replay distributed-reinforcement-learning impala A3C asynchronous-advantage-actor-critic PPO proximal-policy-optimization TRPO trust-region-policy-optimization DDPG deep-deterministic-policy-gradient TD3 twin-delayed-deep-deterministic-policy-gradient SAC soft-actor-critic model-based-reinforcement-learning model-predictive-control guided-policy-search imitation-learning inverse-reinforcement-learning apprenticeship-learning behavior-cloning dataset-aggregation DAgger generative-adversarial-imitation-learning third-person-imitation-learning one-shot-imitation-learning meta-reinforcement-learning hierarchical-reinforcement-learning option skill subgoal intrinsic-motivation curiosity-driven-exploration empowerment information-gain prediction-error novelty-seeking diversity-seeking multi-agent-reinforcement-learning mean-field fictitious-play correlated-equilibrium Nash-equilibrium game-theory mechanism-design auction-theory incentive-design social-choice-theory voting-theory fair-division matching-theory market-design algorithmic-game-theory computational-social-choice computational-complexity NP-completeness NP-hardness approximation-algorithm approximation-ratio inapproximability fixed-parameter-tractability parameterized-complexity kernelization treewidth graph-minor forbidden-minor-characterization well-quasi-ordering Robertson-Seymour-theorem graph-structure-theorem bidimensionality-theory meta-theorem Courcelle's-theorem automata-theory finite-automaton pushdown-automaton Turing-machine computability decidability halting-problem Rice's-theorem Gödel's-incompleteness-theorems lambda-calculus combinatory-logic type-theory proof-theory model-theory set-theory forcing large-cardinal axiom-of-choice continuum-hypothesis independence-result consistency-strength reverse-mathematics constructive-mathematics intuitionistic-logic modal-logic temporal-logic epistemic-logic deontic-logic description-logic ontology knowledge-representation semantic-web linked-data RDF SPARQL OWL knowledge-graph entity-linking relation-extraction event-extraction coreference-resolution named-entity-recognition part-of-speech-tagging dependency-parsing constituency-parsing semantic-parsing natural-language-inference textual-entailment natural-language-generation machine-translation summarization question-answering dialogue-system conversational-AI chatbot virtual-assistant speech-recognition speech-synthesis speaker-recognition emotion-recognition sentiment-analysis opinion-mining aspect-based-sentiment-analysis target-dependent-sentiment-analysis stance-detection rumor-detection fake-news-detection fact-checking claim-verification evidence-retrieval argument-mining persuasion-detection deception-detection personality-prediction mental-health-detection suicide-prevention crisis-intervention social-media-mining network-analysis community-detection influence-maximization information-diffusion cascade-modeling epidemic-modeling SEIR-model SIR-model agent-based-epidemic-model compartmental-model network-epidemic-model metapopulation-epidemic-model spatial-epidemic-model stochastic-epidemic-model branching-process renewal-equation generation-interval serial-interval incubation-period latent-period infectious-period basic-reproduction-number effective-reproduction-number herd-immunity-threshold vaccination-coverage vaccine-efficacy vaccine-effectiveness breakthrough-infection waning-immunity immune-escape variant-of-concern variant-of-interest variant-under-monitoring phylogenetic-analysis molecular-clock-analysis phylodynamics viral-evolution antigenic-drift antigenic-shift zoonotic-spillover reservoir-host intermediate-host amplifying-host dead-end-host vector-borne-disease mosquito-borne-disease tick-borne-disease flea-borne-disease lice-borne-disease midge-borne-disease sandfly-borne-disease tsetse-borne-disease blackfly-borne-disease triatomine-borne-disease kissing-bug Chagas-disease trypanosomiasis leishmaniasis malaria dengue yellow-fever Zika-virus chikungunya West-Nile-virus Japanese-encephalitis tick-borne-encephalitis Lyme-disease Rocky-Mountain-spotted-fever ehlichiosis anaplasmosis babesiosis tularemia plague murine-typhus epidemic-typhus endemic-typhus scrub-typhus Q-fever trench-fever bartonellosis cat-scratch-disease rat-bite-fever leptospirosis brucellosis anthrax melioidosis glanders tularaemia francisellosis coxiellosis rickettsiosis orientia-infection ehrlichiosis neoehrlichiosis neorickettsiosis anaplasmataceae rickettsiaceae coxiellaceae bartonellaceae brucellaceae francisellaceae pasteurellaceae legionellaceae chlamydiaceae mycoplasmataceae ureaplasmataceae spiroplasmataceae achaenoplasmataceae entomoplasmataceae haloplasmataceae tenericutes mollicutes bacteroidetes fusobacteria spirochaetes synergistetes deferribacteres thermotogae aquificae caldiserica caldithrix deferrisoma desulfurobacterium thermovibrio thermocrinis hydrogenobacter sulfurihydrogenibium persephonella hydrogenobaculum hydrothermus thermothrix sulfuricurvum sulfurimonas sulfurovum thiovulum thiomicrospira pseudomonas vibrio aeromonas photobacterium aliivibrio gracilibacillus halobacillus halomonas chromohalobacter cobetia salinicola kushneria halotalea halovibrio salinivibrio thioalkalivibrio thiohalophilus ectothiorhodospira halorhodospira thiorhodospira halochromatium thioflavicoccus marichromatium thiocapsa lamprocystis thiodictyon thiopedia amoebobacter thiocystis chromatium allo-chromatium thermochromatium rhabdochromatium thiobaca thiohalocapsa halocapsa thiolamprovum thioploca thiomargarita beggiatoa methylococcus methylomonas methylomicrobium methylosinus methylocystis methylocella methylocapsa methylacidiphilum verrucomicrobia planctomycetes chlamydiae lentisphaerae kiritimatiellaeota calditrichaeota fervidibacteria
Tibicinoides pallidula
Tibicinoides pallidula is a species of cicada in the family Cicadidae, originally described as Okanagana pallidula by Davis in 1917 before being transferred to the genus Tibicinoides. It belongs to the subfamily Tibicininae and tribe Tibicinini. As a member of the Auchenorrhyncha, it is a true bug with piercing-sucking mouthparts and undergoes incomplete metamorphosis. The species epithet 'pallidula' suggests pale coloration.
cicada Hemiptera Cicadidae Tibicinoides Auchenorrhyncha incomplete-metamorphosis