Aggregation
Guides
Andrognathus
Andrognathus is a genus of small, thin-bodied millipedes in the order Platydesmida, family Andrognathidae. The genus contains three extant species—A. corticarius, A. grubbsi, and A. hoffmani—distributed in deciduous forests of eastern North America, New Mexico, and Mexico respectively. A fossil species, A. burmiticus, is known from 99-million-year-old Burmese amber, indicating a formerly broader distribution. These millipedes are euanamorphic, adding segments throughout their lifespan, and exhibit paternal care behaviors.
Anisolabis maritima
maritime earwig, seaside earwig
A cosmopolitan, wingless earwig species found on coastal beaches worldwide. Males possess asymmetrical forceps used for mating, prey capture, and defense. Exhibits notable maternal care behavior, with females guarding eggs and nymphs for up to 28 days and progressively provisioning offspring. Displays complex social dynamics including sex-specific aggression patterns and size-based dominance in shelter competition.
Anisomorpha
two-striped walkingsticks, twostriped walkingsticks
Anisomorpha is a genus of large wingless walking stick insects (Phasmida) known for their potent chemical defense. Adults possess paired metathoracic glands that discharge an irritating secretion capable of causing intense burning pain and temporary blindness in predators, including humans. Females are substantially larger than males, with some individuals reaching nearly four inches in length. The genus contains four accepted species distributed across the southeastern United States, Central America, and northern South America.
Anurida maritima
seashore springtail, rock springtail
Anurida maritima is a cosmopolitan springtail confined to the intertidal zone, where it forms dense aggregations of up to several hundred individuals on rock-pool surfaces. This wingless collembolan exhibits unique adaptations to marine conditions, including a vestigial furcula that prevents jumping and hydrophobic body hairs that enable it to remain on water surfaces. The species demonstrates complex behavioral rhythms synchronized with tidal cycles and survives winter through egg diapause. Recent genetic studies have revealed cryptic speciation within this morphospecies, with distinct mitochondrial and nuclear lineages now recognized.
Aphis varians
Aphis varians is an aphid species described by Patch in 1914. Research has demonstrated that this species employs aggregation as an effective anti-predator strategy, with individuals benefiting from reduced per capita predation risk when grouped together. The species serves as prey for the convergent lady beetle Hippodamia convergens, with which it has been studied in predator-prey interaction research.
Armadillidium vulgare
common pill-bug, common pill woodlouse, roly-poly, potato bug, doodle bug, carpenter
Armadillidium vulgare is a terrestrial isopod native to Mediterranean Europe that has become one of the most widespread woodlouse species globally through human-mediated dispersal. It is the most extensively studied terrestrial isopod and serves as a model organism for research on mitochondrial genome evolution, desiccation resistance, and conglobation behavior. The species exhibits remarkable morphological plasticity, including numerous color morphs maintained through selective breeding in the pet trade.
Bembidion petrosum
A ground beetle species with a broad Eurosiberian distribution spanning North America, Europe, and temperate Asia. Breeds during summer and hibernates as an overwintering strategy. Frequently aggregates beneath stones in riparian habitats. Four subspecies are recognized, including the nominate form and three regional variants.
Boisea rubrolineata
Western Boxelder Bug
Boisea rubrolineata, the western boxelder bug, is a scentless plant bug native to western North America, ranging west of the Continental Divide. This species is a specialist herbivore primarily associated with boxelder maple (Acer negundo), feeding on both pollen from staminate trees and seeds from pistillate trees. Distinguished from its eastern counterpart B. trivittata by prominent red veins on the corium, it exhibits complex semiochemical-mediated behaviors including aggregation and host foraging. The species undergoes incomplete metamorphosis and seeks sheltered overwintering sites in autumn, often becoming a nuisance pest when entering human structures.
Boisea trivittata
eastern boxelder bug, box bug, maple bug, adobe bug
Boisea trivittata, commonly known as the eastern boxelder bug, is a North American true bug in the family Rhopalidae. Despite being classified as 'scentless plant bugs,' this species produces defensive monoterpene hydrocarbons that deter predators. Adults are approximately 12-13 mm long with dark brown or black coloration marked by distinctive red wing veins and abdominal markings. The species undergoes incomplete metamorphosis, with bright red nymphs maturing to winged adults before winter. It is primarily associated with boxelder trees (Acer negundo), though it also occurs on other maples and ash trees.
Corythucha juglandis
Walnut Lace Bug
Corythucha juglandis, the walnut lace bug, is a univoltine or bivoltine lace bug (Hemiptera: Tingidae) native to North America and specialized on walnut (Juglans) host plants. Adults overwinter in leaf litter and become active in April or May, synchronizing with host leaf phenology. The species aggregates on the undersides of leaflets for feeding and reproduction. Both adults and nymphs are gregarious. It has been studied in Ontario, Québec, Canada, and southern Illinois, USA, with some geographic variation in voltinism reported.
Dendroctonus brevicomis
Western Pine Beetle
Dendroctonus brevicomis, commonly known as the western pine beetle, is a destructive bark beetle species in the family Curculionidae. It is a major pest of ponderosa pine (Pinus ponderosa) and Coulter pine (Pinus coulteri) in western North America. The species is known for causing extensive tree mortality during outbreaks, with documented losses of 60–90% of host trees in affected landscapes. A notable outbreak occurred in the central and southern Sierra Nevada Range from 2014 to 2017, killing millions of ponderosa pines. The beetle plays a significant role in forest dynamics but also poses substantial economic and ecological challenges.
Epicauta costata
Four-ribbed Blister Beetle
Epicauta costata is a blister beetle in the family Meloidae, commonly known as the Four-ribbed Blister Beetle. The species was described by LeConte in 1854. It is distributed across Central America and North America. Like other meloids, it produces cantharidin, a defensive compound that causes blistering upon contact with skin.
Exenterus amictorius
Exenterus amictorius is an ichneumonid parasitoid wasp introduced to North America that attacks sawfly larvae (Hymenoptera: Diprionidae), particularly during the pre-spinning eonymph stage. The species exhibits adaptive host discrimination behavior that changes during the host's spinning period, lacks discrimination initially but rapidly acquires and maintains this ability. It shows strong positive density-dependent responses to host abundance and has established successfully on multiple diprionid hosts across North America. In multiparasitism situations with the native E. diprionis, E. amictorius consistently survives due to faster larval development.
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affinis
Ixodes affinis is a hard-bodied tick species distributed across the southeastern United States, Mexico, Central America, and South America. Unlike its congener Ixodes scapularis, this species does not bite humans but serves as a competent vector for Borrelia burgdorferi sensu stricto, the causative agent of Lyme disease. Immature stages feed on birds and small mammals, while adults parasitize larger mammals including deer and dogs. Its expanding range into southeastern Virginia and association with migratory birds has raised concerns about its role in pathogen dispersal.
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pulchrissimus
desert bee fly
Lordotus pulchrissimus is a bee fly in the family Bombyliidae, commonly known as the desert bee fly. Males and females exhibit strong sexual dimorphism in size, fur density, and coloration—females are smaller (2–14 mm), more densely furred, and display brighter orange-yellow tones that fade rapidly with age, while males are larger (8–16 mm), less hairy, and possess black markings on the femora. The species is notable for the daily aerial swarming behavior of males, which form aggregations over stabilized dunes for reasons that remain unclear; this behavior is energetically costly and occurs independently of female presence or resource density. Adults feed primarily on nectar from desert brush, particularly rabbitbrush (Chrysothamnus nauseosus), and serve as pollinators. Larvae are parasitoids, though specific host insects remain unidentified.
Lygus hesperus
Western Tarnished Plant Bug
Lygus hesperus is a significant agricultural pest in western North America, causing extensive damage to cotton, strawberries, alfalfa seed, and other crops. In California alone, annual losses exceed $30 million in cotton and $40 million in strawberries. Adults overwinter in reproductive diapause triggered by short day lengths, resuming activity when conditions improve. The species has been the subject of extensive research on sampling methods, biological control, and insecticide resistance.
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admirabilis
Admirable Seed Bug
Melacoryphus admirabilis is a seed bug in the family Lygaeidae, native to the Nearctic region of North America. The species is closely related to other soapberry bugs and shares ecological associations with plants in the family Sapindaceae. It occurs in the southwestern United States and Mexico, typically in warm, arid habitats. Like other members of its genus, it feeds on seeds and plant fluids. The species is occasionally encountered in large aggregations.
Melacoryphus lateralis
Charcoal Seed Bug, Black-and-red Seed Bug
Melacoryphus lateralis is a species of true bug in the family Lygaeidae, commonly known as the charcoal seed bug or black-and-red seed bug. Native to desert regions of western North America, this species is recognized by its distinctive black body fringed with red and gray coloration, resembling a dying ember. Populations are known to appear in large aggregations during late summer. The species is closely related to the small milkweed bug (Lygaeus kalmii) and shares similar coloration patterns with other lygaeid seed bugs.
Palaemon vulgaris
Marsh Grass Shrimp, Common American Prawn, Common Grass Shrimp, Marsh Shrimp
Palaemon vulgaris is a small, transparent caridean shrimp native to the western Atlantic Ocean. Adults reach less than 5 cm in length and display distinctive orange pigmentation on the eyestalks. The species has been studied for its specialized swimming mechanics, employing metachronal beating of pleopods and drag-reducing mechanisms during locomotion. It occupies coastal and estuarine habitats from Cape Cod Bay to the Gulf of Mexico.
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rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetlesDynastinaeScarabaeidaeColeopteratribeglobal-distributionmorphological-diversitysexual-dimorphismgeneric-diversitymouthpart-morphologysecondary-sexual-charactershornsbiogeographic-restrictiontaxonomic-revisiondichotomous-keysnew-species-descriptionnew-genus-descriptionlectotype-designationsynonymynew-combinationdistribution-mappingfemale-descriptionhabitat-databehavioral-observationsAustraliaColombiaBoliviaIndiaWestern-AustraliaNew-South-WalesNeotropicalAustralianAfrotropicalOrientalPalaearcticCheiroplatinaDipelicinaPentodontinaPseudoryctinaBothynusHeteronychusEpironastesPhilcarneumConstricticollisCarneiolaAnomalomorphaEnraciusErbmahcediusCavonusPericoptusPentodonCalicnemisMetanastesNeometanastesPimelopusPodalgusPseudoryctesCheiroplatysDipelicusDenheziaEuetheolaHylobothynusOxyligyrusParapucayaPucayaTomarusAdoryphorusCarneoryctesTeinogenysLigyrusAllsoppHutchinsonArrowCarneEndrödiDechambrePrellOhausBatesHopeLaporte-de-CastelnauErichsonBurmeisterSharpMulsantBlackburnDupuisÖzdikmenYamayaFairmaireRedtenbacherSteinheilRatcliffeCaveFabriciusDejeaniNaturalistWikipediaCatalogue-of-LifeZootaxaJournal-of-Insect-BiodiversityRecords-of-the-Zoological-Survey-of-IndiaThe-Coleopterists-BulletinBioLib.czWikimedia-CommonsDOI10.11646/zootaxa.4048.4.110.11646/zootaxa.4852.4.210.11646/zootaxa.5351.3.210.26515/rzsi/v125/i2s/2025/17296410.11646/zootaxa.5716.4.710.11646/zootaxa.5072.5.210.11646/zootaxa.4852.4.310.12976/jib/2024.54.2.210.1649/1186.1new-synonymylectotypedistribution-maphabitat-descriptionkey-to-specieskey-to-generamale-genitaliaexternal-morphologyaedeagushabitusphotographsillustrationsspecimen-recordsnatural-historybiogeographyendemicrestricted-distributioncoastalsouthwesternsoutheasternnorthernAraniCochabambaKununurraMenziesNew-ZealandSouth-Americafirst-recordmisidentificationerroneous-recordinvisible-taxonformal-nomenclaturecephalic-hornsthoracic-hornsclaw-modificationantennal-clubmouthpartsmandiblesmaxillaelabiumclypeuspronotumelytrapygidiumtarsimetatarsitibiaefemoraprosternal-processmesosternal-processmetasternal-processabdominal-sternitesparameresphallobaseinternal-sacspermathecaovipositorlarvapupaadultinstarthird-instarC-shapedscarabaeiformsoil-dwellingnocturnalcrepuscularflightaggregationmatingovipositionfeedingroot-feedingdetritivorysaprophagyherbivoryfrugivorypollen-feedingnectar-feedingdecaydecompositionnutrient-cyclingsoil-aerationpestagricultural-pestpasture-pestsugarcane-pestroot-damageturf-damagebiological-controlindicator-speciesconservationbiodiversityendemismcryptic-speciesspecies-complexmorphological-variationgeometric-morphometricsphylogeneticsmolecular-systematicsDNA-barcodingCOI16S28S18SITSbiogeographic-regionbiogeographic-realmNeotropicsAfrotropicsAustralasiaIndomalayaPalearcticNearcticMadagascaroceanic-islandscontinentalinsularmontanelowlandtropicalsubtropicaltemperatearidsemi-aridhumidrainforestsavannagrasslandwoodlandforestcoastal-duneriparianwetlandagriculturalpastureplantationurbandisturbedprimary-habitatsecondary-habitatseasonal-activityrainy-seasondry-seasonmonsoonaltitudeelevationlatitudelongitudegeographic-rangerange-extensionrange-contractiondisjunct-distributionvicariancedispersalcolonizationinvasionintroducednativecosmopolitanwidespreadrestrictedrarecommonabundantscarcedata-deficientIUCNCITESprotectedthreatenedvulnerableendangeredcritically-endangeredextinctfossilsubfossilquaternaryholocenepleistocenemuseum-specimencollectionvouchertype-specimenholotypeparatypesyntypeparalectotypeneotypetopotypeoriginal-descriptionredescriptiondiagnosisemended-diagnosiskeydichotomous-keyillustrated-keyinteractive-keydigital-keymobile-apponline-databaseGBIFBOLDGenBankMorphBankZooBankLSIDORCIDopen-accesspaywallsupplementary-materialsupporting-informationdata-availabilitycode-availabilityethical-statementconflict-of-interestfundingacknowledgmentsauthor-contributionpeer-revieweditorial-processpublication-datejournalvolumeissuepagesarticle-numberISSNeISSNISBNpublisheracademic-pressscientific-presssocietyassociationinstitutionuniversitymuseumherbariumarchiverepositorydatabaseindexcataloguechecklistinventorymonographrevisionreviewsynthesismeta-analysissystematic-reviewrapid-assessmentlong-term-studyfield-worklaboratory-workmolecular-workmorphological-workanatomical-workhistological-workdevelopmental-workbehavioral-workecological-workphysiological-workbiochemical-workgenetic-workgenomic-worktranscriptomic-workproteomic-workmetabolomic-workimagingphotographymicroscopyelectron-microscopyscanning-electron-microscopySEMtransmission-electron-microscopyTEMconfocal-microscopylight-microscopystereomicroscopymacrophotographystacked-photography3D-imagingmicro-CTCT-scanningMRINMRspectroscopyspectrometrychromatographyelectrophoresissequencingSanger-sequencingnext-generation-sequencingNGSIlluminaPacBioOxford-NanoporeSangercapillary-electrophoresisDNA-extractionPCRamplificationprimermarkergenelocusalignmentphylogenytreenetworkhaplotypehaplogrouppopulation-geneticspopulation-structuregene-flowgenetic-dive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hconnectivity-conservationcorridorbuffer-zoneprotected-areanational-parknature-reservewildlife-refugewilderness-areaworld-heritage-sitebiosphere-reserveRamsar-siteImportant-Bird-AreaKey-Biodiversity-AreaAlliance-for-Zero-Extinction-siteconservation-priorityhotspotcrisis-ecoregionglobal-200last-of-the-wildhuman-footprintcumulative-impactthreat-indexvulnerability-indexadaptive-capacityexposuresensitivityresilienceresistancerecoveryrestorationrehabilitationreintroductiontranslocationex-situin-situcaptive-breedingbotanic-gardenzoogene-bankseed-banktissue-banksperm-bankoocyte-bankembryo-bankDNA-bankfrozen-zooarkinsurancesafety-netde-extinctiongenetic-rescuegenetic-restorationgenetic-augmentationgenetic-managementpopulation-managementmetapopulationsource-sinkpatchmatrixlandscapeseascapeecosystembiomeecoregionprovincezoneregiondistrictsitelocalityhabitatmicrohabitatnicheecological-nichefundamental-nicherealized-nichetrophic-nichespatial-nichetemporal-nichebiotic-nicheabiotic-nichemultidimensional-nichen-dimensional-nicheHutchinsonian-nicheGrinnellian-nicheEltonian-nicheresourcerequirementlimitationstressdisturbanceperturbationfluctuationvariabilityheterogeneitycomplexitydiversityredundancystabilitypersistenceadaptationacclimationplasticityevolvabilityheritabilityselectiondriftflowmutationrecombinationspeciationcoalescencedivergenceconvergenceparallelismhomoplasyanalogyhomologysynapomorphysymplesiomorphyautapomorphyapomorphyplesiomorphyderivedancestralprimitiveadvancedbasalcrownstemnodebranchcladegradesubfamilyfamilysuperfamilyinfraordersuborderordersuperorderinfraclasssubclassclasssuperclasssubphylumphylumsuperphylumkingdomdomainlifeorganismindividualetc.Petrolisthes
porcelain crabs
Petrolisthes is a genus of marine porcelain crabs in the family Porcellanidae, containing approximately 50 extant species distributed across temperate and tropical coastal regions of the Pacific Rim and warm Western Atlantic. These small, flattened crabs occupy intertidal and subtidal zones, often forming dense aggregations under rocks and within crevices. Multiple species have been subject to intensive behavioral and ecological study, revealing complex patterns of intraspecific competition, thermal adaptation, and settlement behavior. Two species (P. cinctipes and P. manimaculis) have been sequenced, providing genomic resources for studying physiological responses to climate change.
Petrolisthes eriomerus
flattop crab
Petrolisthes eriomerus, commonly called the flattop crab, is a small porcelain crab inhabiting the eastern Pacific coast of North America. It reaches 20 mm in carapace width and exhibits a distinctly flattened, rounded body form adapted for life under rocks and in crevices. The species employs filter feeding and deposit sweeping to consume diatoms and organic material. It displays notable social behavior, including aggregations and ritualized agonistic interactions between individuals.
Pyrrhocoris apterus
firebug, European firebug
Pyrrhocoris apterus, commonly known as the firebug or European firebug, is a striking true bug recognized by its vivid red and black coloration. Native to the Palaearctic region, it has expanded its range to North America, Central America, India, and Australia. The species exhibits pronounced gregarious behavior, with larvae and adults frequently forming aggregations of tens to hundreds of individuals. It is primarily seed-feeding, specializing on lime tree and mallow seeds. A notable feature is its non-functional wing polymorphism: populations contain rare long-winged (macropterous) individuals alongside the typical short-winged (brachypterous) form, though the species is effectively flightless.
Spirostreptidae
Flatplate Millipedes
Spirostreptidae is a family of large millipedes in the order Spirostreptida, commonly known as flatplate millipedes. The family comprises approximately 100 genera distributed across tropical and subtropical regions of the Americas, Africa, Madagascar, Seychelles, and the eastern Mediterranean. Members are characterized by their elongated cylindrical bodies and are primarily soil-dwelling detritivores, though some species exhibit arboreal habits. The family includes both synanthropic species that can become urban pests and species with specialized thermoregulatory and social behaviors.