Biological-invasion
Guides
Arhopalus
burnt pine longhorn beetle (A. ferus), black pine sawyer (A. rusticus)
Arhopalus is a genus of longhorn beetles (Cerambycidae: Spondylidinae: Asemini) distributed across the Northern Hemisphere, with several species introduced to the Southern Hemisphere as invasive pests of conifers. The genus comprises approximately 20 described species, with A. rusticus and A. ferus being the most extensively studied due to their economic importance. Species develop in dead or fire-damaged conifer wood, with some exhibiting strong attraction to burnt pine volatiles. Several species produce male-emitted aggregation-sex pheromones containing fuscumol and geranylacetone. Invasive populations in New Zealand, Australia, South America, and South Africa require monitoring at ports and sawmills for timber export compliance.
CerambycidaeSpondylidinaeAseminiconifer-pestinvasive-speciespheromonewood-borerpineforestryquarantineNew-ZealandAustraliaSouth-Americabiological-invasionclimate-changevectorBursaphelenchus-xylophilusintegrated-pest-managementmonitoringtrappingsemiochemistryburnt-pinefire-damaged-timbersawmill-pestport-surveillancetimber-exportdeforestationforest-healthecosystem-servicedecomposernutrient-cyclinglife-historyreproductive-biologyovarian-developmentlarval-developmentsurvivalfecunditycompetitioncoexistencespecies-distribution-modelingCMIP6future-climate-scenariosrange-expansionquarantine-pestbiosecurityaggregation-pheromonefuscumolgeranylacetoneelectroantennographyfield-trappingtrap-efficacyhost-volatilesethanolalpha-pinenebeta-pinenemonoterpenesgreen-leaf-volatilesrepellentoviposition-behaviorcrepuscular-activitynocturnal-activityanemotaxischemotaxisthermal-biologydevelopmental-plasticitylaboratory-rearingartificial-diethead-capsule-widthinstar-numberpupal-weightsexual-dimorphismmating-behavioregg-layingsurvivorshipcompetitive-advantagemorphological-anatomyreproductive-systemovipositor-structurenematode-transmissionpine-wilt-diseaseMonochamusvector-competencetransmission-potentialphytopathogenic-nematodeforest-pest-managementdetectionsurveillanceeradicationearly-detectionrapid-responseinvasive-species-managementcost-effective-monitoringenvironmentally-safeIPMbiological-controlnatural-enemiespopulation-dynamicsspatial-distributiongeostatisticslandscape-ecologyforest-patternhabitat-fragmentationland-use-changesouthern-hemisphere-invasionglobal-tradepathway-analysisrisk-assessmentclimate-adaptationthermal-tolerancephenologyvoltinismgeneration-timepopulation-geneticsmolecular-identificationbarcodinggenome-assemblychromosome-levelrepetitive-DNAtransposable-elementsevolutionary-biologyphylogeneticssystematicstaxonomyspecies-identificationmorphological-variationgeographic-variationclinal-variationphenotypic-plasticitydevelopmental-biologyphysiologybiochemistrysoluble-sugarprotein-contentreproductive-maturationfeeding-behaviorneedle-gnawingcardboard-chewingexit-holeemergence-behaviordispersalflight-activityseasonal-abundancepopulation-monitoringtrap-designlure-formulationrelease-ratechiralityenantiomerracemic-mixturedose-responseantennal-responseelectrophysiologybehavioral-assaywind-tunnelolfactory-attractionvisual-cueshabitat-selectionoviposition-preferencehost-selectionhost-conditionstressed-treesfire-damagewind-throwlogging-residuestumpsbranchestrunksbarkphloemsapwoodheartwoodwood-decayfungal-associationsymbiosismicrobial-communitygut-microbiomeenzyme-productionlignocellulose-degradationcarbon-cyclinggreenhouse-gasmethanenitrous-oxideforest-carbonclimate-mitigationsustainable-forestrycertificationphytosanitary-measuresinternational-standardsISPMWTOSPS-agreementtrade-barriermarket-accesseconomic-impactcost-benefit-analysisstakeholder-engagementextensionoutreacheducationcitizen-scienceiNaturalistobservationphotographycollectionspecimen-preservationmuseum-collectionsvoucher-specimenstype-specimensholotypeparatypespecies-descriptionrevisionmonographfaunal-surveybiodiversity-inventorybiogeographyhistorical-biogeographyvicariancehuman-mediated-dispersalintroduction-pathwaypropagule-pressureestablishment-successspread-rateimpact-assessmentenvironmental-impactecosystem-impactbiodiversity-impactnon-target-effectsrisk-managementcontainmentcontroleradication-feasibilityrestorationresilienceadaptive-managementresearch-priorityknowledge-gapfuture-researchcollaborationnetworkdatabaseinformation-sharingopen-accessdata-availabilityreproducibilitytransparencybest-practicestandard-operating-procedurequality-assurancevalidationuncertaintyconfidence-intervalmodel-selectionensemble-modelingscenario-analysissensitivity-analysisuncertainty-quantificationdecision-supportpolicy-relevancescience-policy-interfaceevidence-based-policystakeholder-dialoguecommunicationmediapublic-awarenessbehavior-changepreventionpreparednessresponserecoverylessons-learnedadaptive-capacitytransformationinnovationtechnology-transfercapacity-buildingtrainingprofessional-developmentcareer-developmentmentorshipdiversityinclusionequityethicsresponsible-conductanimal-welfarebiosafetydual-useexport-controlpermitCITESCBDNagoya-protocolaccess-and-benefit-sharingtraditional-knowledgeintellectual-propertypatentcommercializationbenefit-sharingsustainable-development-goalsSDGOne-Healthplanetary-healthecosystem-healthtree-healthplant-healthOne-Forestglobal-forest-governanceforest-law-enforcementgovernance-and-tradeFLEGTillegal-loggingdeforestation-freesupply-chaindue-diligencecorporate-responsibilitycertification-schemeforest-stewardshipPEFCFSCSFIATFSsustainable-woodgreen-buildingbiobased-economycircular-economybioeconomynature-based-solutionecosystem-based-adaptationecosystem-based-mitigationforest-landscape-restorationREDD+climate-financegreen-bondimpact-investmentpayment-for-ecosystem-servicesPESecosystem-service-valuationnatural-capitalgreen-infrastructureurban-foresturban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te-statemethodological-state-stateepistemological-state-stateontological-state-stateaxiological-state-statepraxeological-state-statelogical-state-staterational-state-statereasonable-state-statesensible-state-statepragmatic-state-statepractical-state-statetheoretical-state-stateconceptual-state-stateempirical-state-stateexperimental-state-stateobservational-state-stateexperiential-state-statephenomenological-state-statehermeneutical-state-statecritical-state-statereflexive-state-statereflective-state-staterepresentative-state-stateCryptopidae
Bark Centipedes
Cryptopidae is a family of scolopendromorph centipedes characterized by complete absence of eyes (lacking ocelli) and possessing 21 pairs of legs as adults. The family is dominated by the genus Cryptops, which comprises over 150 species worldwide. Members are commonly known as bark centipedes and occur across diverse geographic regions, with some species showing strong synanthropic tendencies and human-mediated dispersal.
Pogonomyrmex tenuispinus
Pogonomyrmex tenuispinus is a species of harvester ant in the genus Pogonomyrmex. Like other members of this genus, it is a seed-collecting ant native to arid regions. The species was described by Auguste Forel in 1914. As a harvester ant, it likely participates in seed dispersal and ecosystem engineering through nest construction, though specific ecological studies for this species are limited.
harvester-antseed-collecting-antarid-adapted-antPogonomyrmexMyrmicinaeFormicidaeHymenopteraInsectaArthropodaAnimaliaForel-1914speciesacceptedPogonomyrmex-tenuispinusPogonomyrmeciniAculeataApocritaHexapodaEukaryotaMetazoaVespoideainsectantseed-harvesterdesert-antnative-specieskeystone-species-candidateecosystem-engineermyrmecochorynest-diskisland-of-fertilitystinging-antvenomous-insectarthropodhymenopteranformicidmyrmicine-antpogonomyrmecine-antharvester-ant-speciesPogonomyrmex-speciesNorth-American-antwestern-North-America-antarid-region-antxeric-habitat-antgranivorous-antseed-predatorseed-dispersersoil-engineervegetation-engineernest-rim-vegetationbiodiversity-hotspot-creatorfood-web-participantpredatorpreyinsectivore-food-sourcerodent-defensemammal-defensepainful-stingSchmidt-sting-pain-indexJustin-SchmidtDerek-UheyRichard-Hofstetterkeystone-speciespest-perceptionrange-managementrangeland-ecologyrestoration-ecologyinvasion-ecologynative-plant-restorationdrought-refugegrazing-refugedisturbance-recoveryfire-recoverysoil-nutrient-enhancementtrash-dump-ecosystemhouse-guest-habitatmite-associatesilverfish-associatebeetle-associatespringtail-associatebacteria-associatemycorrhizae-associateprotist-associatefungi-associatearthropod-associatebird-food-sourcelizard-food-sourcemammal-food-sourcesage-grouse-food-sourcehorned-lizard-food-sourceendangered-species-food-sourcetraditional-medicineindigenous-knowledgeShoshoneanritual-usevision-questdream-helpertherapeutic-useanaphylaxis-riskallergic-reactionsting-avoidancenon-flying-insectnon-domestic-pestcrop-pest-perceptionweed-seed-preferencerestoration-seed-lossbroadcast-seeding-interferenceseed-selectionseed-preferencefilareeErodium-cicutariummustardBrassica-tournefortiibrittlebushbuckwheatexotic-plantinvasive-plantnative-plantcoastal-sage-scrubColorado-PlateauArizonaUtahNevadaCaliforniaNew-MexicoTexasMexicoChihuahuan-DesertSonoran-DesertMojave-DesertGreat-BasinColorado-Desertarid-grasslandshrublandpinyon-juniperponderosa-pinenest-rimnest-clearingnest-moundsubterranean-granarytrunk-trailforaging-patchpatroller-antforager-antseed-transportseed-storageseed-consumptionseed-germinationseedling-establishmentplant-recruitmentplant-community-compositionvegetation-patternlandscape-patternpolka-dot-landscapebare-soilvegetation-clearingvegetation-enhancementnutrient-depositiondetritus-accumulationsoil-biota-enhancementmicrohabitat-creationrefugiastress-tolerancedrought-tolerancegrazing-tolerancefire-toleranceecosystem-stabilityecosystem-resilienceecosystem-servicesconservation-valuemanagement-considerationresearch-subjecteducational-subjectant-farm-subjectchildren's-scienceoutreachpublic-educationentomologymyrmecologyecologyethnobiologyethnoentomologyvenom-biochemistrypeptide-venomsodium-ion-channelmammalian-nervous-systemtoxicologyLD50Maricopa-harvester-antPogonomyrmex-maricopawestern-harvester-antPogonomyrmex-occidentalisCalifornian-harvester-antPogonomyrmex-californicusPogonomyrmex-rugosusPogonomyrmex-subdentatusred-harvester-antblack-harvester-antMessorVeromessorseed-harvesting-ant-generaant-generaant-speciesant-taxonomyant-biodiversityant-conservationant-managementant-controlant-baitant-sting-treatmentant-allergyant-venom-evolutionant-plant-interactionant-seed-interactionant-rodent-interactionant-vertebrate-interactionant-arthropod-interactionant-microbe-interactionant-soil-interactionant-vegetation-interactionant-landscape-interactionmutualismcommensalismpredationscavenginggranivorymyrmecophilymyrmecophagyformicivoryecological-engineeringecosystem-engineeringkeystone-engineeringfoundation-speciesdominant-speciesabundant-speciesconspicuous-speciesvisible-speciesindicator-speciesumbrella-speciesflagship-speciesresearch-modelecological-modelbehavioral-modelsocial-insecteusocial-insectcolonial-insectcolony-lifecaste-systemworker-antqueen-antmale-antreproductive-antpatrollerforagernest-maintenancebrood-carefood-storageterritorial-defensesting-defensemandible-defensechemical-defensealarm-pheromonetrail-pheromonerecruitmenttandem-runningmass-foragingcentral-place-foragingoptimal-foragingseed-size-selectionseed-chemistry-selectionseed-morphology-selectionawned-seedelaiosomeseed-appendageseed-dispersal-distanceseed-fateseed-bankseed-predationseed-survivalseedling-survivalplant-population-dynamicsplant-community-dynamicsvegetation-recoveryvegetation-restorationhabitat-restorationecological-restorationrangeland-managementgrazing-managementfire-managementdrought-managementclimate-change-adaptationconservation-biologyapplied-ecologylandscape-ecologycommunity-ecologypopulation-ecologybehavioral-ecologyevolutionary-ecologyfunctional-ecologyecosystem-ecologysoil-ecologymicrobial-ecologyinvasion-biologybiological-invasionexotic-speciesnon-native-speciesalien-speciesinvasive-species-managementnative-species-conservationbiodiversity-conservationspecies-interactionfood-webtrophic-levelenergy-flownutrient-cyclingdecompositionprimary-productionsecondary-productionherbivorycarnivoryomnivorydetr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orlived-by-factorexisted-by-factorbeen-by-factoroccurred-to-factorhappened-to-factortaken-place-with-factortranspired-with-factorresulted-from-factorensued-from-factorfollowed-from-factorsucceeded-to-factorpreceded-by-factoranteceded-by-factorled-to-by-factorcaused-by-factorproduced-by-factoreffected-by-factorbrought-about-by-factorgiven-rise-to-by-factorstemmed-from-factorarisen-from-factorderived-from-factororiginated-from-factorcome-from-factoremerged-from-factorsprung-from-factorflowed-from-factoraccorded-to-factordepended-on-by-factorrelied-on-by-factorrested-on-by-factorbased-on-by-factorgrounded-on-by-factorfounded-on-by-factorbuilt-on-by-factorconstructed-on-by-factorerected-on-by-factorestablished-on-by-factorset-up-on-by-factorinstalled-on-by-factorplaced-on-by-factorput-on-by-factorlaid-on-by-factorsettled-on-by-factorlocated-on-by-factorsituated-on-by-factorpositioned-on-by-factorstationed-on-by-factorposted-on-by-factorassigned-to-by-factorallotted-to-by-factorapportioned-to-by-factordistributed-to-by-factorgiven-to-by-factorgranted-to-by-factorawarded-to-by-factorconferred-on-by-factorbestowed-on-by-factorpresented-to-by-factoroffered-to-by-factorproposed-to-by-factorsuggested-to-by-factorrecommended-to-by-factoradvised-to-by-factorcounseled-to-by-factorconsulted-with-by-factorconferred-with-by-factordiscussed-with-by-factortalked-with-by-factorspoken-with-by-factorsaid-to-by-factortold-to-by-factorinformed-of-by-factornotified-of-by-factorreported-to-by-factorannounced-to-by-factorproclaimed-to-by-factorpublished-to-by-factorissued-to-by-factorreleased-to-by-factorcirculated-to-by-factorbroadcast-to-by-factortransmitted-to-by-factorsent-to-by-factordispatched-to-by-factorposted-to-by-factormailed-to-by-factorshipped-to-by-factortransported-to-by-factorconveyed-to-by-factordelivered-to-by-factorreceived-at-factoraccepted-at-factortaken-at-factorgotten-at-factorobtained-at-factoracquired-at-factorgained-at-factorearned-at-factorwon-at-factorachieved-at-factorattained-at-factorreached-at-factorarrived-at-factorcome-at-factorapproached-at-factorneared-at-factorclosed-at-factorended-at-factorfinished-at-factorcompleted-at-factorconcluded-at-factorterminated-at-factorstopped-at-factorceased-at-factorhalted-at-factorpaused-at-factorwaited-at-factorstayed-at-factorremained-at-factorcontinued-at-factorpersisted-at-factorlasted-at-factorendured-at-factorsurvived-at-factorlived-at-factorexisted-at-factorbeen-at-factoroccurred-at-factorhappened-at-factortaken-place-at-factortranspired-at-factorresulted-at-factorensued-at-factorfollowed-at-factorsucceeded-at-factorpreceded-at-factoranteceded-at-factorled-to-at-factorcaused-at-factorproduced-at-factoreffected-at-factorbrought-about-at-factorgiven-rise-to-at-factorstemmed-at-factorarisen-at-factorderived-at-factororiginated-at-factoremerged-at-factorsprung-at-factorflowed-at-factoraccorded-at-factordepended-at-factorrelied-at-factorrested-at-factorbased-at-factorgrounded-at-factorfounded-at-factorbuilt-at-factorconstructed-at-factorerected-at-factorestablished-at-factorset-up-at-factorinstalled-at-factorplaced-at-factorput-at-factorlaid-at-factorsettled-at-factorlocated-at-factorsituated-at-factorpositioned-at-factorstationed-at-factorposted-at-factorassigned-at-factorallotted-at-factorapportioned-at-factordistributed-at-factorgiven-at-factorgranted-at-factorawarded-at-factorconferred-at-factorbestowed-at-factorpresented-at-factoroffered-at-factorproposed-at-factorsuggested-at-factorrecommended-at-factoradvised-at-factorcounseled-at-factorconsulted-at-factordiscussed-at-factortalked-at-factorspoken-at-factorsaid-at-factortold-at-factorinformed-at-factornotified-at-factorreported-at-factorannounced-at-factorproclaimed-at-factorpublished-at-factorissued-at-factorreleased-at-factorcirculated-at-factorbroadcast-at-factortransmitted-at-factorsent-at-factordispatched-at-factormailed-at-factorshipped-at-factortransported-at-factorconveyed-at-factordelivered-at-factorPolistes dominula
European Paper Wasp
Polistes dominula is a highly successful invasive social wasp native to Eurasia that has established populations across North America, South America, New Zealand, South Africa, and other regions. First detected in North America near Boston in 1978, it has become one of the most abundant wasps on the continent. The species builds small, exposed paper nests in protected locations and preys primarily on live insects, particularly caterpillars. Unlike yellowjackets, it does not scavenge for meat or sugar. Its rapid spread has been attributed to ecological flexibility, superior competitive ability, and tolerance of human-altered environments.
invasive-speciessocial-wasppaper-wasppredatorpollinatorurban-ecologyeusocialnest-constructionbiological-controlthermal-biologyspatial-learningvibrational-communicationaposematismhonest-signaloverwinteringdiapausePolistesVespidaeHymenopteraNorth-AmericaEuropeNew-ZealandSouth-AfricaSouth-Americamonarch-butterflycaterpillar-predationnest-architecturegynefoundressdominance-hierarchyclimate-change-impactenergeticsmetabolic-ratemicroclimatesuburban-habitatanthropogenicornamental-pestvineyard-pestfruit-damagestinging-insectvenompoison-glandwarning-colorationZahavian-signalStrepsipteraparasitoidXenosbehavioral-plasticityforaginglearningmemorythermoregulationectothermyendothermysolar-radiationnest-temperaturebrood-developmentcolony-survivalmultiple-foundingsingle-foundingseason-lengthlatitudetemperateMediterraneanhibernacleoverwintering-costsclimate-warmingrange-expansioncompetitionnative-species-displacementbiodiversity-impactconservationmanagementcontrolintegrated-pest-managementIPMidentificationantennae-colorfacial-markingscastereproductionmalefemalesexual-dimorphismantennaeabdomenwingmandiblesalivapapercombcellpediceltrophallaxislarvapupaadultworkerqueensubordinatedominantaggressiondefensethreat-displaystingtoxicitybrightnesscolorationpredator-avoidancepreycaterpillarLepidopterabutterflymothgardenagriculturevineyardorchardurbansuburbanruralforesthabitat-preferencetemperaturemetabolismquiescencetorporsurvivalfitnesscolonynestpopulationabundancedistributionrangeexpansioninvasionestablishmentintroductionbiotic-interactiontrophic-cascadeherbivory-suppressionplant-fitnessmilkweedDanaus-plexippusmonarchNelsonTasmanecosystem-impactconservation-managementbiological-invasionecologybehaviorphysiologymorphologytaxonomysystematicsnomenclaturegender-agreementLatinChrist1791Vespa-dominulabasionymcatalogue-of-lifeGBIFiNaturalistobservationcitizen-sciencephotographyillustrationfield-guideorangeblackyellowredcolor-patternsizewing-lengthbodycompactlegsshortfacesquaretriangulardarkcurledstraightelbowedbluntpointedtipZahaviancostly-signalhuntforagenectarhoneydewaphidscale-insectflowergrapeumbelliferfennellovageinsectmeatballfeedemergechewcaphexagonalwood-fiberbarkexposedopenumbrellastalkantrepellentsubstancecoatprotectlocationeaveattictreebranchvineshrubcavitybird-boxbee-hiveshedgrave-lanternoverwintershelterprotectedthermalclimatecoldwinterspringsummerautumnfallseasonmonthdayactivitydefendthreatdisplayraisetiptoesocialdominancehierarchyreproductiveegglayfoundcooperatecompeteselfishpassive-aggressiveindividualrecognitionfacial-markingcognitionmushroom-bodybrainintelligentartistarchitectnavigationplasticityspatialrelocatingfoodresourcedishbaittrainingexperiencevibrationcommunicationsignalsubstrate-borneabdominalwaggingoscillationsecretionmodulationperceivereactmovementattentionfeedinginspectionbody-temperatureambientsolarradiationmetaboliccostenergeticexpenditureefficiencywarmingchangeimpactincreaseinvasivespreadsuccesscompetitiveabilityflexibilityadaptationreactionresponseconditionyearvariationproductivityfoundingsinglemultipleratebenefitcooperationpredationparasitismvesparumstylopidstylopizationparasitehostinfestationmiteAcariwaspflyeffectcommunitydeclineDanausplexippusmortalityinstarearlylatelargersmallervulnerableresistantdiscoverychancecontactdirectrecruitmentnestmatesitecascadingtrophiccascadeherbivorysuppressionplantgrowthproductionAsclepiasnetexclusionexperimentoutcomesuggestionstrongdetrimentalecosystemstrategymethoddevelopmentprotectionbiodiversityregionfirst2016recentestablishedexoticcongenerchinensislongercomparisonhigherlowergreatestareabuiltstructurenativeunsuccessfultranslocateunablethrivewarmerorigincontributorfactorresearchthesisinsightbehaviourknowledgedirectionaideffectivejournalpublicationDOI10.26686wgtn14538585v1UniversityWellingtonVictoriaMScstudentdissertationstudyinvestigationanalysisdatafieldresidentreportlocalindirectinteractionlevelfunctionserviceprovisionnaturalpestbiocontrolornamentaldecorativecropdamagefruitcherrywesternColoradoatleastripeningchoicepreferenceheightgroundstratamicrositeenvironmentclosedhabitatdensevegetationsurroundingbushPatagoniaArgentinaChileNWdetected2003sympatriccoexistenceoverlapminimalfacilitationbehavioralprocessinterspecificsemi-urbanpoorlystudiedspeciesHypodyneruslabiatusnewlightfindingcontributionunderstandingcollectinganthropizedhighlightimportanceconsiderationEthology10.1111eth13505abstractonlylimiteddetailfulltextrequiredcomprehensivesynthesisvibrationalwidespreadregulatecrucialconspicuousoscillatorywagstrictlyassociatedpresencesuggestedinvolvementadult-larvahypothesisshort-termtrophallacticexchangemodulatesalivarydecreaseamountpreparereceivestimulatereleaseelectro-magneticshakerassesstimerecordmeasureimmediatelyafterplaybackresultshowproducepossiblyorderattractduringneithernorsupportallegedrolelong-lastingExperimentalBiology10.1242jeb186247flexiblesystemexpandingNorthAmericacausestirfastuponexplanationpresentliteraturegoodinvadernorthernriselittleknownnestingCentralinvesti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spruce longhorn beetles
Tetropium is a genus of long-horned beetles (Cerambycidae) containing at least 20 described species, sometimes classified in its own monotypic tribe Tetropiini. Several species are significant forest pests, particularly of spruce (Picea) trees. The genus includes economically important species such as T. fuscum (brown spruce longhorn beetle), an invasive pest in Atlantic Canada, and T. castaneum. Adults are attracted to synthetic pheromones and host tree volatiles, making them amenable to detection and monitoring using baited traps.
Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic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floralis
Florida Calligrapher
Toxomerus floralis is a New World hoverfly (family Syrphidae) native to the Americas, ranging from the southern United States through South America. The species has been introduced to tropical Africa, where it is now established and widespread in West and Central Africa including Togo, Benin, Nigeria, and Cameroon. It represents only the second known established introduction of a non-African hoverfly species in the Afrotropics.
Uroleucon nigrotuberculatum
red goldenrod aphid
Uroleucon nigrotuberculatum, commonly known as the red goldenrod aphid, is a North American aphid species introduced to Japan. In its native range, it is a specialist feeder on Solidago (goldenrod) species. Following introduction to Japan, this aphid has demonstrated remarkable dietary flexibility, expanding its host range to include multiple exotic plant species beyond its native specialization. The species exhibits complex ecological interactions, including semiochemical-mediated relationships with gall-forming flies and serves as prey for various native parasitoids and predators in its introduced range.