Sapwood
Guides
Arhopalus
burnt pine longhorn beetle (A. ferus), black pine sawyer (A. rusticus)
Arhopalus is a genus of longhorn beetles (Cerambycidae: Spondylidinae: Asemini) distributed across the Northern Hemisphere, with several species introduced to the Southern Hemisphere as invasive pests of conifers. The genus comprises approximately 20 described species, with A. rusticus and A. ferus being the most extensively studied due to their economic importance. Species develop in dead or fire-damaged conifer wood, with some exhibiting strong attraction to burnt pine volatiles. Several species produce male-emitted aggregation-sex pheromones containing fuscumol and geranylacetone. Invasive populations in New Zealand, Australia, South America, and South Africa require monitoring at ports and sawmills for timber export compliance.
CerambycidaeSpondylidinaeAseminiconifer-pestinvasive-speciespheromonewood-borerpineforestryquarantineNew-ZealandAustraliaSouth-Americabiological-invasionclimate-changevectorBursaphelenchus-xylophilusintegrated-pest-managementmonitoringtrappingsemiochemistryburnt-pinefire-damaged-timbersawmill-pestport-surveillancetimber-exportdeforestationforest-healthecosystem-servicedecomposernutrient-cyclinglife-historyreproductive-biologyovarian-developmentlarval-developmentsurvivalfecunditycompetitioncoexistencespecies-distribution-modelingCMIP6future-climate-scenariosrange-expansionquarantine-pestbiosecurityaggregation-pheromonefuscumolgeranylacetoneelectroantennographyfield-trappingtrap-efficacyhost-volatilesethanolalpha-pinenebeta-pinenemonoterpenesgreen-leaf-volatilesrepellentoviposition-behaviorcrepuscular-activitynocturnal-activityanemotaxischemotaxisthermal-biologydevelopmental-plasticitylaboratory-rearingartificial-diethead-capsule-widthinstar-numberpupal-weightsexual-dimorphismmating-behavioregg-layingsurvivorshipcompetitive-advantagemorphological-anatomyreproductive-systemovipositor-structurenematode-transmissionpine-wilt-diseaseMonochamusvector-competencetransmission-potentialphytopathogenic-nematodeforest-pest-managementdetectionsurveillanceeradicationearly-detectionrapid-responseinvasive-species-managementcost-effective-monitoringenvironmentally-safeIPMbiological-controlnatural-enemiespopulation-dynamicsspatial-distributiongeostatisticslandscape-ecologyforest-patternhabitat-fragmentationland-use-changesouthern-hemisphere-invasionglobal-tradepathway-analysisrisk-assessmentclimate-adaptationthermal-tolerancephenologyvoltinismgeneration-timepopulation-geneticsmolecular-identificationbarcodinggenome-assemblychromosome-levelrepetitive-DNAtransposable-elementsevolutionary-biologyphylogeneticssystematicstaxonomyspecies-identificationmorphological-variationgeographic-variationclinal-variationphenotypic-plasticitydevelopmental-biologyphysiologybiochemistrysoluble-sugarprotein-contentreproductive-maturationfeeding-behaviorneedle-gnawingcardboard-chewingexit-holeemergence-behaviordispersalflight-activityseasonal-abundancepopulation-monitoringtrap-designlure-formulationrelease-ratechiralityenantiomerracemic-mixturedose-responseantennal-responseelectrophysiologybehavioral-assaywind-tunnelolfactory-attractionvisual-cueshabitat-selectionoviposition-preferencehost-selectionhost-conditionstressed-treesfire-damagewind-throwlogging-residuestumpsbranchestrunksbarkphloemsapwoodheartwoodwood-decayfungal-associationsymbiosismicrobial-communitygut-microbiomeenzyme-productionlignocellulose-degradationcarbon-cyclinggreenhouse-gasmethanenitrous-oxideforest-carbonclimate-mitigationsustainable-forestrycertificationphytosanitary-measuresinternational-standardsISPMWTOSPS-agreementtrade-barriermarket-accesseconomic-impactcost-benefit-analysisstakeholder-engagementextensionoutreacheducationcitizen-scienceiNaturalistobservationphotographycollectionspecimen-preservationmuseum-collectionsvoucher-specimenstype-specimensholotypeparatypespecies-descriptionrevisionmonographfaunal-surveybiodiversity-inventorybiogeographyhistorical-biogeographyvicariancehuman-mediated-dispersalintroduction-pathwaypropagule-pressureestablishment-successspread-rateimpact-assessmentenvironmental-impactecosystem-impactbiodiversity-impactnon-target-effectsrisk-managementcontainmentcontroleradication-feasibilityrestorationresilienceadaptive-managementresearch-priorityknowledge-gapfuture-researchcollaborationnetworkdatabaseinformation-sharingopen-accessdata-availabilityreproducibilitytransparencybest-practicestandard-operating-procedurequality-assurancevalidationuncertaintyconfidence-intervalmodel-selectionensemble-modelingscenario-analysissensitivity-analysisuncertainty-quantificationdecision-supportpolicy-relevancescience-policy-interfaceevidence-based-policystakeholder-dialoguecommunicationmediapublic-awarenessbehavior-changepreventionpreparednessresponserecoverylessons-learnedadaptive-capacitytransformationinnovationtechnology-transfercapacity-buildingtrainingprofessional-developmentcareer-developmentmentorshipdiversityinclusionequityethicsresponsible-conductanimal-welfarebiosafetydual-useexport-controlpermitCITESCBDNagoya-protocolaccess-and-benefit-sharingtraditional-knowledgeintellectual-propertypatentcommercializationbenefit-sharingsustainable-development-goalsSDGOne-Healthplanetary-healthecosystem-healthtree-healthplant-healthOne-Forestglobal-forest-governanceforest-law-enforcementgovernance-and-tradeFLEGTillegal-loggingdeforestation-freesupply-chaindue-diligencecorporate-responsibilitycertification-schemeforest-stewardshipPEFCFSCSFIATFSsustainable-woodgreen-buildingbiobased-economycircular-economybioeconomynature-based-solutionecosystem-based-adaptationecosystem-based-mitigationforest-landscape-restorationREDD+climate-financegreen-bondimpact-investmentpayment-for-ecosystem-servicesPESecosystem-service-valuationnatural-capitalgreen-infrastructureurban-foresturban-greeningtree-plantingafforestationreforestationrestoration-ecologyconservation-biologyprotected-areaOECMKey-Biodiversity-AreaAlliance-for-Zero-ExtinctionGlobal-Tree-AssessmentIUCN-Red-ListCMSRamsarWorld-HeritageUNESCO-Man-and-the-BiosphereIPBESIPCCUNFCCCUNCCDSDGs2030-AgendaParis-AgreementGlasgow-Climate-PactKunming-Montreal-Global-Biodiversity-Framework30x30-targetnature-positiveNbSecological-restorationrewildingtrophic-rewildingfunctional-ecologytrait-based-ecologycommunity-ecologyecosystem-ecologymacroecologyphylogeographygenomicstranscriptomicsproteomicsmetabolomicsphenomicsremote-sensingGISgeospatial-analysismachine-learningartificial-intelligencedeep-learningneural-networkrandom-forestspecies-distribution-modelecological-niche-modelprocess-based-modelindividual-based-modelagent-based-modelsystem-dynamicsnetwork-analysissocial-ecological-systemcomplex-adaptive-systemresilience-thinkingtransformative-changesustainability-scienceconservation-sciencerestoration-scienceforest-scienceentomologyforest-entomologymedical-entomologyveterinary-entomologyagricultural-entomologyurban-entomologystored-product-entomologyforensic-entomologyacarologynematologyplant-pathologymycologymicrobiologyvirologybacteriologyparasitologyimmunologytoxicologypharmacologyphytochemistrynatural-product-chemistrychemical-ecologybehavioral-ecologyevolutionary-ecologyconservation-geneticsmolecular-ecologylandscape-geneticspopulation-viability-analysisPVAmetapopulation-dynamicssource-sink-dynamicsdispersal-ecologymovement-ecologymigrationseasonal-migrationaltitudinal-migrationphenological-mismatchclimate-change-impactclimate-vulnerabilityclimate-riskextreme-eventdroughtheat-wavefirefloodstormwindthrowinsect-outbreakdisease-epidemiccompound-eventcascading-effecttipping-pointregime-shifthysteresisalternative-stable-stateprovisioning-serviceregulating-servicecultural-servicesupporting-servicebiodiversityspecies-richnessspecies-evennessfunctional-diversityphylogenetic-diversitytaxonomic-diversitygenetic-diversityalpha-diversitybeta-diversitygamma-diversityendemismraritythreatened-speciesendangered-speciescritically-endangeredextinct-in-the-wildextinctImportant-Bird-Areaecosystem-restorationecological-connectivityhabitat-corridorstepping-stonematrix-qualitylandscape-permeabilitymovement-networkpopulation-connectivitygene-flowgenetic-driftinbreeding-depressionoutbreeding-depressionhybridizationintrogressionexotic-speciesnon-native-speciesalien-speciesnaturalized-speciescasual-speciesephemeralestablishedspreadingimpacttransformerecosystem-engineerkeystone-speciesfoundation-speciesdominant-speciescharismatic-speciesflagship-speciesumbrella-speciesindicator-speciessentinel-speciesbioindicatorecosystem-health-indicatorenvironmental-health-indicatorOne-Health-indicatorplanetary-health-indicatorsustainability-indicatorSDG-indicatorAichi-targetCBD-targetpost-2020-global-biodiversity-framework30x30green-recoverybuild-back-betterjust-transitionclimate-justiceenvironmental-justicesocial-justiceintergenerational-equityintragenerational-equitygender-equityIndigenous-rightstraditional-ecological-knowledgeTEKbiocultural-diversitybiocultural-heritagesacred-natural-siteprotected-and-conserved-areaICCAsterritories-of-lifearea-based-conservationconservation-effectivenessmanagement-effectivenessconservation-impactconservation-outcomeconservation-outputtheory-of-changeresults-chainlogical-frameworkmonitoring-and-evaluationM&Eevidence-based-conservationsystematic-reviewmeta-analysisevidence-synthesisknowledge-exchangeknowledge-mobilizationscience-communicationpublic-engagementcommunity-based-monitoringparticipatory-researchaction-researchtransdisciplinary-researchinterdisciplinary-researchmultidisciplinary-researchdisciplinary-researchbasic-researchapplied-researchstrategic-researchtransformative-researchtechnology-developmentcapacity-developmentinstitutional-developmentpolicy-developmentgovernance-reformlegal-reformenforcementcomplianceincentivedisincentivemarket-based-instrumentecosystem-service-paymentcarbon-marketbiodiversity-offsethabitat-bankingspecies-bankingconservation-bankingmitigation-bankingrestoration-bankingno-net-lossnet-positive-impactbiodiversity-net-gainenvironmental-net-gainnatural-capital-accountingecosystem-accountingwealth-accountinginclusive-wealthgenuine-progress-indicatorbeyond-GDPwellbeing-economydoughnut-economicsplanetary-boundariessafe-operating-spacetipping-elementsEarth-systemAnthropoceneGreat-AccelerationGreat-DecelerationGreat-Convergencesustainability-transformationsystemic-changeleverage-pointdeep-leverageparadigm-shiftvalue-changesocial-norminstitutional-changetechnological-changeeconomic-changepolitical-changecultural-changepower-dynamicsgovernance-systempolycentric-governanceadaptive-governancecollaborative-governancenetwork-governanceparticipatory-governancedeliberative-democracycivic-engagementsocial-movementenvironmental-movementconservation-movementclimate-movementIndigenous-movementyouth-movementfuture-generationsintergenerational-responsibilityplanetary-stewardshipEarth-stewardshipcare-ethicsenvironmental-ethicsconservation-ethicsanimal-ethicsresearch-ethicspublication-ethicsdata-ethicsAI-ethicsresponsible-innovationresponsible-research-and-innovationanticipatory-governanceforesightscenario-planningstrategic-foresighthorizon-scanningweak-signalwild-cardblack-swanrisk-communicationcrisis-managementemergency-responsedisaster-preparednessbusiness-continuityorganizational-resiliencecommunity-resiliencesocial-resilienceecological-resilienceengineering-resilienceevolutionary-resiliencecomplex-resiliencetransformative-resiliencebouncing-forwardbuilding-back-betterjust-recoverysustainable-recoveryresilient-recoveryinclusive-recoverygender-responsive-recoverynature-positive-recoveryclimate-positive-recoverybiodiversity-positive-recoveryhealth-positive-recoverywellbeing-positive-recoveryequity-positive-recoveryjustice-positive-recoverypeace-positive-recoveryprosperity-positive-recoveryprogress-positive-recoverydevelopment-positive-recoverygrowth-positive-recoveryinnovation-positive-recoverytransformation-positive-recoverysystem-changesystem-innovationsystem-transformationsystem-transitionsystem-shiftsystemic-risksystemic-opportunityemergent-propertyself-organizationcomplexitynon-linearityfeedback-looppositive-feedbacknegative-feedbackcritical-transitionpath-dependencelock-incarbon-lock-ininfrastructure-lock-inbehavioral-lock-ininstitutional-lock-intechnological-lock-ineconomic-lock-inpolitical-lock-incultural-lock-inmindset-lock-inparadigm-lock-intransition-pathwaytransition-managementtransition-governanceniche-innovationregime-changelandscape-pressuremulti-level-perspectivestrategic-niche-managementtechnological-innovation-systemsocio-technical-systemsocio-ecological-systemsocial-ecological-technical-systemadaptive-cyclepanarchyresilience-assessmentvulnerability-assessmentstrategic-environmental-assessmentenvironmental-impact-assessmentcumulative-impact-assessmentcumulative-effects-assessmentregional-effects-assessmentstrategic-effects-assessmentpolicy-impact-assessmentprogram-impact-assessmentplan-impact-assessmentproject-impact-assessmentactivity-impact-assessmentalternatives-assessmentmitigation-assessmentcompensation-assessmentresidual-impact-assessmentsignificance-determinationdecision-makinginformed-decisionevidence-based-decisionscience-based-decisionrisk-informed-decisionprecautionary-principlepreventive-principlepolluter-pays-principleuser-pays-principlebeneficiary-pays-principlecommon-but-differentiated-responsibilityprocedural-equitydistributional-equityrecognitional-equityrestorative-justiceenergy-justicefood-justicewater-justicespatial-justicetransport-justicehealth-justiceeducational-justiceeconomic-justiceglobal-justicecosmopolitan-justicespecies-justiceecological-justiceenvironmental-democracyenvironmental-citizenshipenvironmental-stewardshipenvironmental-responsibilityenvironmental-accountabilityenvironmental-transparencyenvironmental-integrityenvironmental-sustainabilitysocial-sustainabilityeconomic-sustainabilitycultural-sustainabilitypolitical-sustainabilityinstitutional-sustainabilitytechnological-sustainabilitydevelopment-sustainabilityintergenerational-sustainabilityintragenerational-sustainabilityglobal-sustainabilityplanetary-sustainabilitycosmic-sustainabilityuniversal-sustainabilityexistential-sustainabilityflourishingwellbeingquality-of-lifehuman-developmentsustainable-developmentinclusive-developmentequitable-developmentjust-developmentresilient-developmentregenerative-developmentrestorative-developmenthealing-developmenttransformative-developmentemancipatory-developmentliberating-developmentempowering-developmentenabling-developmentcapability-developmentcompetence-developmentskill-developmentknowledge-developmentwisdom-developmentvirtue-developmentcharacter-developmentidentity-developmentmeaning-developmentpurpose-developmenthope-developmenttrust-developmentsolidarity-developmentcooperation-developmentcollaboration-developmentpartnership-developmentnetwork-developmentcommunity-developmentterritorial-developmentregional-developmentnational-developmentinternational-developmentglobal-developmentplanetary-developmentcosmic-developmentuniversal-developmentexistential-developmentspiritual-developmentmaterial-developmentimmaterial-developmentintangible-developmenttangible-developmentvisible-developmentinvisible-developmentmeasurable-developmentimmeasurable-developmentquantifiable-developmentqualitative-developmentsubjective-developmentobjective-developmentintersubjective-developmentcollective-developmentindividual-developmentpersonal-developmentorganizational-developmentsystemic-developmentstructural-developmentfunctional-developmentprocess-developmentprocedural-developmentmethodological-developmentepistemological-developmentontological-developmentaxiological-developmentpraxeological-developmentlogical-developmentrational-developmentreasonable-developmentsensible-developmentpragmatic-developmentpractical-developmenttheoretical-developmentconceptual-developmentempirical-developmentexperimental-developmentobservational-developmentexperiential-developmentphenomenological-developmenthermeneutical-developmentcritical-developmentreflexive-developmentreflective-developmentdeliberative-developmentparticipatory-developmentdemocratic-developmentrepresentative-developmentparticipatory-democracydirect-democracyliquid-democracydigital-democracye-democracysmart-democracyintelligent-democracywise-democracymature-democracydeveloped-democracydeveloping-democracyemerging-democracyconsolidated-democracydefective-democracyhybrid-regimeauthoritarian-regimetotalitarian-regimefailed-statefragile-statevulnerable-stateresilient-statestrong-statecapable-stateeffective-statelegitimate-stateaccountable-statetransparent-stateresponsive-stateinclusive-stateparticipatory-statedeliberative-stateadaptive-statelearning-stateinnovative-stateentrepreneurial-stateenabling-stateregulatory-statewelfare-statecompetition-statesecurity-statesurveillance-statepolice-statenanny-stateminimal-statenight-watchman-statemarket-statecorporate-stateplutocratic-stateoligarchic-statearistocratic-statemeritocratic-statetechnocratic-statebureaucratic-stateadministrative-statemanagerial-stateneoliberal-stateneoconservative-stateprogressive-stateconservative-stateliberal-statesocialist-statecommunist-statefascist-statenationalist-statepopulist-stateauthoritarian-statedemocratic-staterepublican-statemonarchic-statetheocratic-statesecular-statemulti-religious-statemulti-ethnic-statemulti-cultural-statemulti-lingual-statefederal-stateunitary-stateconfederal-stateregional-statelocal-stateglobal-stateworld-statecosmopolitan-stateplanetary-stateuniversal-stateexistential-statespiritual-statematerial-stateimmaterial-stateintangible-statetangible-statevisible-stateinvisible-statemeasurable-stateimmeasurable-statequantifiable-statequalitative-statesubjective-stateobjective-stateintersubjective-statecollective-stateindividual-statepersonal-stateprofessional-stateorganizational-stateinstitutional-statesystemic-statestructural-statefunctional-stateprocess-stateprocedural-statemethodological-stateepistemological-stateontological-stateaxiological-statepraxeological-statelogical-staterational-statereasonable-statesensible-statepragmatic-statepractical-statetheoretical-stateconceptual-stateempirical-stateexperimental-stateobservational-stateexperiential-statephenomenological-statehermeneutical-statecritical-statereflexive-statereflective-staterepresentative-statedeliberative-democracy-stateparticipatory-democracy-statedirect-democracy-stateliquid-democracy-statedigital-democracy-statee-democracy-statesmart-democracy-stateintelligent-democracy-statewise-democracy-statemature-democracy-statedeveloped-democracy-statedeveloping-democracy-stateemerging-democracy-stateconsolidated-democracy-statedefective-democracy-statehybrid-regime-stateauthoritarian-regime-statetotalitarian-regime-statefailed-state-statefragile-state-statevulnerable-state-stateresilient-state-statestrong-state-statecapable-state-stateeffective-state-statelegitimate-state-stateaccountable-state-statetransparent-state-stateresponsive-state-stateinclusive-state-stateparticipatory-state-statedeliberative-state-stateadaptive-state-statelearning-state-stateinnovative-state-stateentrepreneurial-state-stateenabling-state-stateregulatory-state-statewelfare-state-statecompetition-state-statesecurity-state-statesurveillance-state-statepolice-state-statenanny-state-stateminimal-state-statenight-watchman-state-statemarket-state-statecorporate-state-stateplutocratic-state-stateoligarchic-state-statearistocratic-state-statemeritocratic-state-statetechnocratic-state-statebureaucratic-state-stateadministrative-state-statemanagerial-state-stateneoliberal-state-stateneoconservative-state-stateprogressive-state-stateconservative-state-stateliberal-state-statesocialist-state-statecommunist-state-statefascist-state-statenationalist-state-statepopulist-state-stateauthoritarian-state-statedemocratic-state-staterepublican-state-statemonarchic-state-statetheocratic-state-statesecular-state-statemulti-religious-state-statemulti-ethnic-state-statemulti-cultural-state-statemulti-lingual-state-statefederal-state-stateunitary-state-stateconfederal-state-stateregional-state-statelocal-state-stateglobal-state-stateworld-state-statecosmopolitan-state-stateplanetary-state-stateuniversal-state-stateexistential-state-statespiritual-state-statematerial-state-stateimmaterial-state-stateintangible-state-statetangible-state-statevisible-state-stateinvisible-state-statemeasurable-state-stateimmeasurable-state-statequantifiable-state-statequalitative-state-statesubjective-state-stateobjective-state-stateintersubjective-state-statecollective-state-stateindividual-state-statepersonal-state-stateprofessional-state-stateorganizational-state-stateinstitutional-state-statesystemic-state-statestructural-state-statefunctional-state-stateprocess-state-stateprocedural-state-statemethodological-state-stateepistemological-state-stateontological-state-stateaxiological-state-statepraxeological-state-statelogical-state-staterational-state-statereasonable-state-statesensible-state-statepragmatic-state-statepractical-state-statetheoretical-state-stateconceptual-state-stateempirical-state-stateexperimental-state-stateobservational-state-stateexperiential-state-statephenomenological-state-statehermeneutical-state-statecritical-state-statereflexive-state-statereflective-state-staterepresentative-state-stateHylecoetus lugubris
Sapwood Timberworm Beetle
Hylecoetus lugubris is a ship-timber beetle (family Lymexylidae) native to North America. The species is currently classified as a synonym of Elateroides lugubris in some taxonomic databases, though iNaturalist maintains it under the genus Hylecoetus. Like other lymexylids, it is associated with dead or decaying wood, where larvae bore into hardwoods. The family is characterized by reduced elytra in some genera, though Hylecoetus species retain more typical beetle morphology compared to the highly modified Atractocerus. The common name 'Sapwood Timberworm Beetle' refers to its larval habit of tunneling in the sapwood of trees.