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Adelgid

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Adelges abietis
Pineapple-gall Adelgid, Eastern Spruce Gall Adelgid, Spruce Gall Aphid
Adelges abietis, commonly known as the pineapple-gall adelgid or eastern spruce gall adelgid, is a conifer-feeding insect that induces distinctive pineapple-shaped pseudocone galls on spruce shoots. The species is one of the most common adelgids in its genus and has been documented across Europe and North America. Unlike the closely related hemlock woolly adelgid (Adelges tsugae), A. abietis is not considered a major destructive pest, though its gall formation causes shoot deformation. The species exhibits strongly aggregated spatial distributions within and among host trees.
adelgid spruce gall-forming forest-pest Picea pineapple-gall Hemiptera Adelgidae
Adelges cooleyi
Cooley spruce gall adelgid, Douglas fir adelgid, gall adelgid
Adelges cooleyi is a gall-forming adelgid native to western North America, specifically the Rocky Mountains and Cascade Mountains. It is a host-alternating species with a complex life cycle involving two primary hosts: spruce (Picea) species as primary hosts and Douglas fir (Pseudotsuga menziesii) as the secondary host. On spruce, the adelgid induces the formation of distinctive pineapple-shaped galls at branch terminals. On Douglas fir, it produces white, cottony wax masses containing eggs but causes less conspicuous injury. The species has two generations per year and exhibits morphological dimorphism: progredientes migrate to current-year needles while sistentes settle on older needles.
adelgid gall-former host-alternation Douglas-fir spruce pest Christmas-tree invasive genome-sequenced morphological-dimorphism progrediens sistentes western-North-America Sternorrhyncha phloem-feeder
Adelges laricis
Larch Woolly Aphid
Adelges laricis is a holocyclic adelgid with a complex life cycle involving two host genera: Picea (spruce) as primary host where galls form, and Larix (larch) as secondary host. The species exhibits extensive morphological polymorphism across five generations (Fundatrix, Migrans alata, Hiemosistens, Sexupara, Sexualis). First-instar nymphs are the only mobile stage; subsequent instars remain sessile with stylets permanently inserted. The species is a significant forest pest causing reduced tree vigor, needle damage, and mortality under chronic infestation.
adelgid larch-pest parthenogenesis host-alternation forest-entomology invasive-potential gall-former woolly-aphid EPG-study morph-determination
Leptoglossus
leaf-footed bugs
Leptoglossus is a genus of true bugs in the leaf-footed bug family Coreidae, tribe Anisoscelini. Species are characterized by leaflike dilations of the hind tibia, a diagnostic trait of the genus. The genus is distributed throughout the Americas, with some introduced populations in Europe and Asia. Several species are economically significant agricultural pests, notably L. occidentalis, which has become invasive in multiple continents.
Coreidae leaf-footed-bug agricultural-pest invasive-species symbiosis sexual-dimorphism conifer-pest nuisance-pest Burkholderia tachinid-parasitoid pheromone-communication building-invader misidentification Triatoma-look-alike gradual-metamorphosis seed-predator forest-pest ornamental-pest plumbing-damage public-health-confusion Chile Europe North-America South-America Asia Turkey almond-pest citrus-pest tomato-pest corn-pest overwintering-aggregation defensive-secretion pheromone-mediated-parasitism egg-parasitoid biological-control ecological-niche-modeling climate-suitability range-expansion introduction accidental-dispersal seed-orchard-pest germination-reduction empty-seed-formation cone-damage lodgepole-pine Douglas-fir western-conifer-seed-bug L.-occidentalis L.-zonatus L.-phyllopus L.-clypealis L.-australis L.-chilensis L.-oppositus male-combat femoral-weapon abdominal-gland pheromone-gland tibial-dilation leaf-like-hind-leg true-bug Hemiptera Heteroptera Pentatomomorpha Anisoscelini phytophagous plant-feeding stylet-feeding salivary-sheath overwintering diapause building-entry structural-pest nuisance-odor flight-sound droning-flight aggregation-behavior male-pheromone female-choice parasitoid-host biological-control-agent integrated-pest-management monitoring early-detection prevention invasion-biology alien-species non-native-pest global-spread climate-change niche-modeling suitable-habitat establishment-risk quarantine phytosanitary forest-health seed-production reforestation afforestation conifer-forestry pine-seed seed-viability economic-impact crop-loss yield-reduction kernel-damage fruit-damage nut-damage vector-of-disease Chagas-disease kissing-bug public-alarm misinformation education identification-tools health-system-burden pesticide-overuse urban-ecology domiciliary peridomiciliary Andean-region Patagonia Mediterranean temperate subtropical tropical agroecosystem natural-enemy predator parasite pathogen symbiont gut-microbiome soil-ingestion fitness-benefit reproductive-success sperm-morphology testicular-morphology accessory-gland pheromone-blend species-specific-odor cherry-scent vanilla-scent cinnamon-scent rose-scent olfactory-communication mate-location mate-recognition sexual-selection male-male-competition weapon-morphology allometry developmental-plasticity nymphal-instar egg-barrel egg-arrangement linear-oviposition leaf-surface plant-tissue stylet-penetration enzymatic-digestion fluid-feeding phloem-feeding xylem-feeding seed-feeding cone-feeding fruit-feeding kernel-feeding nut-feeding crop-feeding host-plant-range polyphagy oligophagy specialist generalist pest-status damage-threshold economic-injury-level management-strategy cultural-control physical-control chemical-control resistance tolerance host-plant-resistance monitoring-trap light-trap population-density distribution-map spread-rate colonization establishment population-dynamics life-table developmental-rate thermal-requirements degree-days phenology voltinism univoltine bivoltine multivoltine generation-time overwintering-site hibernaculum shelter-seeking cold-hardiness freeze-tolerance supercooling water-relations desiccation-resistance starvation-resistance dispersal-ability flight-capacity walking-behavior climbing-behavior aggregation-pheromone alarm-pheromone stink-gland metathoracic-gland dorsoabdominal-gland ventral-abdominal-gland sexual-gland morphological-defense chemical-defense mechanical-defense autotomy leg-loss predation-risk parasitism-risk parasitoid-risk pathogen-risk competition intraspecific interspecific resource-competition mating-competition sperm-competition cryptic-female-choice reproductive-isolation speciation phylogeny systematics taxonomy morphology anatomy histology ultrastructure spermatogenesis spermatozoa sperm-length nuclear-length cyst-production follicle-number testis-structure reproductive-anatomy genitalia mating-behavior copulation insemination oviposition egg-production fecundity fertility hatching-success nymphal-survival adult-longevity sex-ratio operational-sex-ratio population-sex-ratio effective-population-size genetic-diversity gene-flow population-structure phylogeography biogeography historical-biogeography vicariance dispersal range-shift range-contraction altitudinal-distribution latitudinal-distribution longitudinal-distribution island-distribution continental-distribution endemism cosmopolitanism introduced-range native-range source-population founder-population invasion-front lag-phase establishment-phase spread-phase equilibrium-phase impact-assessment risk-assessment horizon-scanning early-warning rapid-response eradication containment control management adaptation mitigation restoration conservation biodiversity ecosystem-service ecosystem-function food-web trophic-level primary-consumer herbivore fruit-predator plant-animal-interaction mutualism antagonism predation parasitism commensalism amensalism facilitation indirect-interaction trait-mediated-interaction density-mediated-interaction behavioral-ecology evolutionary-ecology functional-ecology physiological-ecology population-ecology community-ecology ecosystem-ecology landscape-ecology macroecology global-ecology applied-ecology agricultural-ecology forest-ecology conservation-biology invasion-ecology restoration-ecology pest-management conservation-biological-control augmentative-biological-control classical-biological-control parasitoid nematode fungus bacterium virus entomopathogen microbial-control sterile-insect-technique genetic-control mating-disruption attract-and-kill push-pull trap-crop border-crop refuge habitat-management cultural-practice crop-rotation tillage irrigation fertilization pruning harvest-timing sanitation residue-management weed-management cover-crop companion-planting intercropping agroforestry silviculture forest-management seed-orchard-management cone-collection seed-extraction seed-testing germination-testing seed-quality seed-vigor empty-seed insect-damage insect-injury feeding-scar puncture stylet-sheath salivary-flange symptom sign diagnosis sampling economic-threshold decision-support expert-system modeling simulation forecasting climate-model niche-model species-distribution-model habitat-suitability risk-mapping invasion-pathway introduction-vector trade transport tourism migration natural-dispersal human-mediated-dispersal accidental-introduction intentional-introduction release escape cultivation ornamental forestry agriculture horticulture urbanization globalization land-use-change habitat-fragmentation habitat-loss degradation pollution pesticide fertilizer nutrient-enrichment eutrophication acidification warming drought extreme-event disturbance succession recovery resilience stability variability uncertainty sustainable-development green-economy circular-economy bioeconomy ecosystem-approach one-health planetary-health environmental-health public-health veterinary-health plant-health food-security nutrition livelihood poverty inequality gender indigenous-knowledge traditional-knowledge local-knowledge citizen-science stakeholder-engagement policy governance regulation legislation international-cooperation capacity-building awareness communication outreach extension advisory-service research innovation technology-transfer knowledge-exchange networking partnership collaboration evaluation evidence-based-policy science-policy-interface diplomacy advocacy activism social-movement environmental-justice environmental-ethics environmental-philosophy environmental-humanities environmental-history environmental-literature environmental-art environmental-education environmental-communication science-communication risk-communication crisis-communication health-communication behavior-change pro-environmental-behavior sustainable-behavior consumption production waste recycling reuse reduction efficiency renewable clean green blue natural organic agroecology permaculture regenerative restorative healing transformative transdisciplinary interdisciplinary multidisciplinary cross-disciplinary disciplinary disciplinary-integration knowledge-integration system-thinking complexity emergence non-linearity feedback tipping-point regime-shift alternative-stable-state resilience-thinking adaptive-management adaptive-governance learning reflection participation inclusion equity justice fairness ethics values norms culture identity place sense-of-place attachment well-being quality-of-life happiness flourishing thriving sustainability sustainable development growth degrowth post-growth steady-state circular doughnut-economics planetary-boundaries safe-operating-space tipping-elements critical-transition early-warning-signal resilience-indicator sustainability-indicator benchmark target goal commitment pledge agreement treaty convention protocol framework strategy plan program project initiative campaign movement coalition alliance network platform hub center 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rehabilitation reconstruction replication duplication backup security safety protection safeguarding insurance risk-management emergency-preparedness response relief humanitarian peace prosperity health employment housing infrastructure energy water food fishery aquaculture mining manufacturing industry commerce recreation heritage nature ecosystem environment climate atmosphere ocean freshwater land soil mineral biota flora fauna microorganism fungi plant animal vertebrate invertebrate arthropod insect bug Hemipteran Heteropteran pentatomomorphan coreoid coreid squash-bug boxelder-bug stink-bug shield-bug plant-bug mirid lygaeid seed-bug broad-headed-bug alydid reduviid assassin-bug triatomine bed-bug cimicid water-bug nepomorphan gerromorphan leptopodomorphan enicocephalomorphan dipsocoromorphan ceratocomboid schizopteroid peloridioid coleorrhynchan moss-bug archaeorrhynchan fulgoromorphan cicadomorphan membracoid treehopper leafhopper planthopper psyllid jumping-plant-louse whitefly aleyrodid scale-insect coccoid mealybug aphid adelgid phylloxeran sternorrhynchan thysanopteran thrips psocopteran barklouse booklouse phthirapteran louse sucking-louse chewing-louse mallophagan anopluran dermapteran earwig blattodean cockroach termite isopteran mantodean mantid phasmid stick-insect leaf-insect orthopteran grasshopper locust katydid cricket mole-cricket pygmy-mole-cricket camel-cricket cave-cricket weta ensiferan caeliferan gryllotalpid myrmecophilid tettigoniid gryllid acridid pamphagid pneumorid lentulid tristirid eumastacid proscopiid tridactylid tetrigid grouse-locust pygmy-grasshopper plecopteran stonefly embiopteran webspinner zorapteran angel-insect dictyopteran
Pineus pinifoliae
Pine Leaf Adelgid, Pine Leaf Aphid, Pine Leaf Chermid
Pineus pinifoliae is a small, phloem-feeding insect in the family Adelgidae, commonly known as the pine leaf adelgid or pine leaf aphid. It has a complex life cycle involving alternation between spruce (Picea) as the primary host, where it forms galls on new shoots, and white pine (Pinus strobus) as the secondary host. The species produces multiple generations annually, with distinct morphological forms including winged migrants that transfer between hosts. Heavy infestations on white pine cause characteristic damage patterns including needle stunting, reduced radial growth, and branch mortality concentrated in the lower and mid-crown.
adelgid gall-former conifer-pest host-alternation forest-entomology integrated-pest-management Pheno-Forecast-species
Pineus similis
Ragged Spruce Gall Adelgid
Pineus similis is a gall-forming adelgid that induces galls on spruce trees (Picea spp.). Originally described as Chermes similis by Gillette in 1907 from blue spruce in Colorado, it was later transferred to genus Pineus by Annand. The species produces winged females that emerge from galls and settle on spruce hosts, with documented preference for white spruce. Wingless females bearing egg clusters occur within galls. It has been recorded on multiple spruce species across North America and has doubtful or rare records in Europe.
adelgid gall-former spruce-pest phylloxerid conifer-specialist Hemiptera Sternorrhyncha North-America Europe Picea
Pineus strobi
Pine Bark Adelgid, White Pine Bark Aphid
Pineus strobi is a small, sap-sucking insect in the family Adelgidae, commonly known as the pine bark adelgid or white pine bark aphid. It is a specialist feeder on white pine (Pinus strobus) and related pine species, forming dense colonies on bark and branches. The species has been introduced to regions outside its native range, including central European Russia and the Fergana Valley, where it is considered invasive. Like other adelgids, it possesses complex associations with bacterial endosymbionts that aid in nutritional processing.
adelgid pine-pest invasive-species endosymbiosis phloem-feeder Hemiptera forestry-pest white-pine honeydew-producer