Timberlake
Guides
Microterys
Microterys is a large genus of parasitoid wasps in the family Encyrtidae (Chalcidoidea), with its center of distribution in the northern parts of the Northern Hemisphere. Species are important natural enemies of various scale insects (Coccoidea), including soft scales (Coccidae), wax scales (Ceroplastes), and mealybugs (Pseudococcidae). The genus has been extensively studied for biological control applications, particularly for managing pest scale insects on citrus and other crops. Several species have been introduced to new regions as biocontrol agents, including Microterys flavus in California.
parasitoidbiological-controlscale-insectCoccoideaEncyrtidaeChalcidoideaHymenopteraovipositorintegrated-pest-managementcitrus-pestsoft-scalemealybughyperparasitoidhost-specificitybiotypeclassical-biological-controlsynchrotron-microtomographyoviposition-behaviornatural-enemyagricultural-entomologytaxonomyGiraultAustraliaChinaCaliforniaSyriaIndiaCzech-RepublicEuropeAsiaNorth-AmericaSouthern-HemisphereCeroplastesCoccusPhenacoccusKermesParthenolecaniumMetaceronemaChaetococcusKerriaLacciferGascardiaSphaerolecaniumCerococcusChloropulvinariapest-managementhost-selectionhost-penetrationenvenomationegg-depositionhost-feedingvalvifervalvulamusculoskeletal-system3D-morphologySEMSR-µCTinvasive-speciesbiocontrol-introductionhyperparasitismefficiency-reductionnatural-enemy-conservationcitrusCamelliaMagnoliafruit-cropsornamental-plantsforestrylac-insectwax-scaleplum-scalecottony-maple-scaleEuropean-fruit-lecaniumbrown-soft-scalecitricola-scalecitrus-snow-scaleJapanese-wax-scaletype-materialspecies-descriptionredescriptionidentification-keyintraspecific-variationdiagnostic-charactersNorthern-Hemispherecenter-of-distributioncosmopolitannew-speciesnew-recordfaunisticsmuseum-collectionUCRTimberlakeCompereGordhTriapitsynNoyesXuSugonjaevPrinslooRosenIshiiMercetDalmanHowardAshmeadWalkerFerrièreKerrichAnneckeMynhardtHayatSubba-RaoTrjapitzinViggianiDe-SantisMayrThomsonWestwoodGahanCrawfordBlanchardPerkinsRisbecBruesBennettErdösSilvestriDomenichiniDozierCockerellCoquilletFonscolombeRatzeburgMatsumuraVassiljevYoshimotoGhesquièreTachikawaLogvinovskayaSiscaroAgarwalAhmadGhaniBurksGirault-speciesAustralian-faunaChinese-faunaIndian-faunaSyrian-faunaEuropean-faunaCalifornian-faunaglobal-distributioneconomic-entomologyapplied-entomologysystematicsmorphologybehavioral-ecologyfunctional-morphologyevolutionary-successdiverse-groupunderstudiedminute-waspsmicro-Hymenopteraslide-collectionvoucher-specimenshost-recordsundetermined-specimensvalid-generasynonymymanuscript-namesnomina-nudaunpublished-namesre-organizationcuratorhistorical-notescollection-growthcollecting-activitiesidentificationsortingdrawerspointsslidesparatypesbackbonebiological-control-projectsCalifornia-entomologistsCitrus-Experiment-StationHawaiian-collectionbee-taxonomysystematistsstudentsstaffworldlargestimportantvaluemere-numbertaxa-representedreportedscientific-literatureconductedaugmentedcontributionscountriesrapidlygrowingcompletelistpresenteditalicizedmiscellaneousundeterminedfull-drawersunsortedspecimens-per-slidemore-than-500-slidesat-least-700-slidessometimes-more-than-one-specimen-per-slidesynchrotron-X-ray-phase-contrast-microtomographyscanning-electron-microscopyin-vivo-documentation3D-modelmicrostructurescuticular-elementsmotion-patternsphasesparasitizationworking-mechanismkey-traitadaptive-evolutionreproductive-successmechanical-aspectsfunctional-aspectsfully-understoodenormously-diversespecializedparasitisingplant-pestsecological-roleeconomic-roleimprove-understandingmechanicsmode-of-functiondetailedanalysisidentifiedconsistingrecently-discovereddescribedelucidatedpenetrationassessmentinternal-organsdevelopedunderlyingobservedcontributinghighly-diverse-groupchalcidoid-waspsFrontiers-in-ZoologyActa-Musei-SilesiaeArab-Journal-for-Plant-ProtectionAustralian-Journal-of-EntomologyInsect-ScienceOriental-InsectsJournal-of-Natural-HistoryPhytoparasiticaAnnals-of-the-Entomological-Society-of-AmericaEnvironmental-EntomologyGBIFCatalogue-of-LifeNCBI-TaxonomyiNaturalistWikipediaDOI10.1186/s12983-025-00575-110.2478/cszma-2021-000710.22268/AJPP-41.3.29230510.1111/j.1440-6055.1975.tb02058.x10.1111/j.1744-7917.2000.tb00345.x10.1080/00305316.2011.64682710.1111/j.1440-6055.1973.tb01670.x10.1080/00222933.2020.178660810.1007/s12600-019-00727-010.1007/s12600-018-0691-510.1093/aesa/54.2.22210.1093/ee/7.6.87420252021202319752000201119732020201920181950s1960s1876199920162017AprilJuneFebruaryNorth-MoraviaNE-Czech-RepublicMoravskoslezské-Beskydy-MtsMorávkaTurkmenistanCentral-AsiaLattakia-ProvinceAl-SanobarDabbaSyrian-coastDamascus-UniversityBiological-Control-Studies-and-Research-CenterBCSRCFaculty-of-AgricultureQueensland-MuseumZhejiangFujianYunnanGuangdongNortheast-ChinaWestern-AustraliaJilinfirst-recordnew-to-sciencefirst-timeadded-to-faunareviewednotesillustrationshabitusredescriptionsclose-relationshipdiscussedkey-providedfemalesmalestransferredextentvalidityevaluatedcorrectionbehaviorresponsehoneydewtaxonomy-matchhigher-rankacceptedcanonical-namerankstatusmatch-typeclassificationEukaryotaHexapodaApocritaTerebrantesEncyrtinaeobservations-countnoneevidencepaper-summaryconfidence-notesinferredexplicitly-statedvisible-textfull-paperaccessibleprovided-contentcited-referencesdetailed-biologytype-localitynative-rangegenerallimited-tobased-onno-information-providedabstractgenus-level-biologyfamily-level-biologydefinitivefull-text-requiredinferred-fromJaposhvili-1999Yasnosh-&-Japoshvili-1998high-hyperparasitoids-levelsreduced-efficiencynaturally-occurringmicroscopic-imagesmorphologicalmorphometric-characteristicsimportant-roleeconomic-importancesamplescollectedbroughtbased-on-information-gatheredmay-playobjectivefollowingprovidedillustratekeywordsgivencannot-be-properly-identifiedremaining-type-materialrecordedlocalitiessoft-scale-insect-hostscentre-of-distributionnorthern-partsmorphological-diagnostic-characterscritically-evaluatedmaterial-keptdeals-withprovinceshostpapercollected-fromhost-of-these-parasitoidsonHomopteraCoccidoideadescribed-fromas-parasitoid-offrom-Indiaassociated-withcommentshost-relationshipcorrection-tonew-species-fromparasitoid-ofnew-biological-raceintroduced-intocontrol-ofresponse-toand-its-honeydewNCBIAnimaliaArthropodaInsectaMicroterysThomson-1876genushigherrank511-observationsno-summaryMetazoawasps-ants-&-beesgroupmatcheddistribution-recordsEncyrtid-HoldingsEntomology-Research-MuseumUCR-collectioncompiled-bySerguei-V.-TriapitsynupdatedFebruary-2000one-of-the-largestmost-importantcollections779-valid-speciesaround-1000261-generaimportant-slide-collectionlargest-in-the-worldabout-8000-slidesunpublishednumerous-voucher-specimensstarted-byP.H.-Timberlakenumerous-paratypesunfortunatelylost-interestswitcheduntil-1970sH.-Comperenotable-contributionsP.-TimberlakeJ.-MapleP.-DeBachC.-Clausen1970s-1980sG.-GordhJ.-LaSalleJ.-WoolleyJ.-PintoJ.-BeardsleyMexicoUSother-countriespresentlygrowing-rapidlyJ.-HeratymajorityV.-TrjapitzinJ.-Noyescomplete-re-organizationS.-Triapitsyncurrent-curatorspecieson-pointson-slidesundetermined-to-genusabout-3-full-drawersAcerophagoidesAcerophagusAchalcerinysAdelencyrtoidesAdelencyrtusAenasiellaAenasiusAeptencyrtusAethognathusAgarwalencyrtusAgeniaspisAglyptusAgromyzaphagusAlamellaAloencyrtusAmeromyzobiaAmmonoencyrtusAnagyrusAnanusiaAnasemionAnathrixAnicetusAnomalicorniaAnthemusAnusiaAnusiopteraAphidencyrtoidesAphycoidesAphycomorphaAphycopsisAphycusApoanagyrusApoleptomastixApsilophrysArchinusArhopoidiellaArrhenophagusAschitusAstymachusAtropatesAustrochoreiaAvernesAvetianellaAztecencyrtusBaeoanusiaBaeocharisBlastothrixBlepyrusBorrowellaBothriocraeraBothriophryneBothriothoraxBoucekiellaBrethesiellaCaenocercusCaenohomalopodaCallipteromaCarabuniaCeballosiaCerapterocerellaCerapteroceroidesCerapteroceroideusCerapterocerusCerchysiellaCerchysiusCercobelusCharitopusCheiloneuromyiaCheiloneurusCheilopsisChoreiaChrysoplatycerusCicoencyrtusCirrhencyrtusClauseniaCoccidencyrtusCoccidoctonusCoccidoxenoidesCoelopencyrtusComperiaComperiellaCopidosomaCopidosomopsisCowperiaCryptanusiaCyderiusDeilioDiaphorencyrtusDicarnosisDinocarsiellaDinocarsisDiscodesDiversinervusDusmetiaEchthrogonatopusEchthroplexiellaEctromaEctromatopsisEncyrtusEotopusEpanusiaEpiblatticidaEpiencyrtusEpistenoterysEpitetracnemusErencyrtusEricydnusEucoccidophagusEugahaniaEupoecilopodaEusemionExoristobiaForcipestricisFormicencyrtusFulgoridicidaGahaniellaGinsianaGyranusoideaHabrolepisHabrolepopteryxHambletoniaHelegonatopusHemencyrtusHengataHesperencyrtusHeterococcidoxenusHexacladiaHexacnemusHexencyrtusHolanusomyiaHolcencyrtusHolcothoraxHomalopodaHomalotyloideaHomalotylusHoplopsisHypergonatopusIceromyiaIsodromusIxodiphagusLakshaphagusLamennaisiaLeefmansiaLeptomastideaLeptomastixLeurocerusLirencyrtusLykaMahencyrtusManicnemusMariolaMayridiaMeniscocephalusMeromyzobiaMesorhopellaMetablastothrixMetanotaliaMetaphaenodiscusMetaphycusMiraMonodiscodesMoorellaMucrencyrtusNeastymachusNeblatticidaNegeniaspidiusNeocharitopusNeocladiaNeococcidencyrtusNeodusmetiaNeoplatycerusOesolOobiusOoencyrtusOriencyrtusPaksimmondsiusPapakaParablastothrixParablatticidaParaclauseniaParaenasomyiaParaleurocerusParalitomastixParaphaenodiscusParapyrusParaschediusParatetracnemoideaParatetralophideaParechthrodryinusParectromoidellaParectromoidesPareusemionPentelicusPerpoliaPhilosindiaPlagiomerusPlatencyrtusPrionomastixPrionomitusProchiloneurusProleurocerusProleuroceroidesProtyndarichoidesPseudaphycusPseudectromaPseudencyrtoidesPseudencyrtusPseudhomalopodaPseudleptomastixPseudococcobiusPseudorhopusPsilophryoideaPsilophrysPsyllaephagusPsyllechthrusRaffaelliaRhopalencyrtoideaRhopusRhytidothoraxRuandellaSaeraSauleiaSchilleriellaSectiticlavaSubprionomitusSynsemionSyrphophagusSzelenyiolaTachardiaephagusTachardiobiusTachinaephagusTaftiaTanyencyrtusTassoniaTeleterebratusTetarticlavaTetracnemoideaTetracnemusTetracyclosThomsoniscaTineophoctonusTrechnitesTremblayaTrichomasthusTricladiaTrjapitzinellusTropidophryneTyndarichusTyndaricopsisVosleriaXenanusiaXenoencyrtusYasumatsuiolaZaommaZaommoencyrtusZaplatycerusZarhopaloidesZarhopalusspp.sp.sp(p).?nr.ms-namemanuscript-namenomen-nudumincludingprobably-mixed-withunsorted-miscellaneousUniversity-of-California-Riversidecollectionmuseumspecimensvouchershostsparasitoidsscale-insectsmealybugssoft-scaleswax-scalescitrus-pestsagricultural-pestsnatural-enemieshyperparasitoidsefficiencyovipositionanatomy3D-reconstructionmicrotomographyin-vivo-observationevolutionadaptive-radiationdiversityAfricaSouth-AmericaMiddle-EastHawaiihorticulturecrop-protectionaugmentative-biological-controlconservation-biological-controlegg-la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timberlakei
Timberlake's Desert Long-horned Grasshopper
Mohavacris timberlakei is a species of desert grasshopper in the family Tanaoceridae, endemic to arid regions of California. It belongs to a small family of specialized orthopterans adapted to extreme desert environments. The species was described by Rehn in 1948 and named in honor of the entomologist P.H. Timberlake, who extensively collected insects in the southwestern United States.
Perdita nasuta
Perdita nasuta is a species of mining bee in the genus Perdita, family Andrenidae. It was first described by entomologist P.H. Timberlake in 1962. As a member of the large Perdita genus—one of the most diverse bee genera in North America with over 800 species—it is a solitary, ground-nesting bee. Like other Perdita species, it is likely a specialist pollinator, though specific host plant associations for this species have not been documented.
Stemmatosteres
Stemmatosteres is a genus of parasitoid wasps in the family Encyrtidae, established by Timberlake in 1918. The genus is characterized by reduced or absent wings in some species, as indicated by the species epithet *apterus* (wingless) in the type species *Stemmatosteres apterus*. The genus belongs to the subfamily Encyrtinae, a diverse group of chalcid wasps primarily known as parasitoids of scale insects and other Hemiptera. Records from the UCR Encyrtidae collection indicate at least two species are recognized, with specimens preserved both on points and on slides.
Stictiella emarginata
sand wasp
Stictiella emarginata is a solitary sand wasp in the family Crabronidae, subfamily Bembicinae. It constructs nests in sandy soils and provisions cells with lepidopteran larvae, primarily Noctuidae and Hesperiidae. The species occurs across eastern North America from northern Michigan to the Atlantic Coast, with a flight season concentrated in late June through early August. Nesting behavior includes temporary nest closure, mound leveling, and orientation flights.
solitary-waspsand-waspnest-provisioninglepidopteran-predatoreastern-North-AmericaBembicinaeCrabronidaeHymenopterasandy-soil-habitattemporary-nest-closureorientation-flightmound-levelingNoctuidaeHesperiidaelate-summer-flight-seasonOntarioMichiganNew-Yorksand-prairieinsect-predatorground-nesting-waspsand-blowout-habitatCanadian-Forces-Base-BordenSimcoe-CountyCresson-1865emarginate-clypeusdisjunct-distribution-misconceptiongeographic-bridgemuseum-recordsfield-studiesnest-architectureprey-transportprovisioning-behaviorcell-constructionprey-storagelarval-developmentsolitary-HymenopteraApoideaSpheciformesAculeataApocritaHexapodaArthropodaInsectaAnimaliaEukaryotaCatalogue-of-LifeGBIFiNaturalistCanadian-Field-NaturalistKurczewskiBoyle2005KrombeinWillinkGillaspyBohartEvansMatthewsPateTimberlakeCooperHallIrwinSchlingerBallmerYanegaUCR-Entomology-Research-MuseumBembicid-HoldingsNyssonidaeBembicidaeBembiciniStictiellinaStictiellaStictiella-emarginataStictiella-bohartiStictiella-corniculataStictiella-evansiStictiella-fergusoniStictiella-flavescensStictiella-pulchellaStictiella-pulchella-serrataStictiella-p.-pulchellaXerostictiaMicrostictiaGlenostictiaStenioliaBembixBicyrtesEdithaMicrobembexRubricaStenogorytesStizusBembecinusStizoidesGorytesArgogorytesArigorytesClitemnestraHarpactusHoplisoidesLestiphorusMegistommumOryttusPsammaletesPseudoplisusSagenistaSpheciusTanyoprymnusTrichogorytesXerogorytesAlyssonDidineisAmmatomusAfrogorytesHapalomellinusNyssonEpinyssonFoxiaHyponyssonLosadaMetanyssonZanyssonAcanthostethusTetracnemoidea sydneyensis
Tetracnemoidea sydneyensis is a species of parasitoid wasp in the family Encyrtidae, described by Timberlake in 1929. The species is native to Australia, with the specific epithet referencing Sydney, and has been introduced to Hawaii as part of biological control programs. Like other encyrtids, it is presumed to parasitize scale insects or other homopteran hosts, though specific host records for this species remain limited in published literature.