Bembicinae
Guides
Alysson
Alysson is a genus of solitary wasps in the family Crabronidae (subfamily Bembicinae, tribe Alyssontini). The genus comprises at least 40 described species distributed across multiple continents. These wasps are part of the diverse assemblage of digger wasps, though specific biological details for the genus as a whole remain incompletely documented. The genus was established by Panzer in 1806 and has undergone taxonomic revisions, with some authorities historically placing it in the family Bembicidae.
Bembecinus nanus
Bembecinus nanus is a small sand wasp species in the tribe Stizina, family Crabronidae. Like other members of its genus, it excavates burrows in sandy substrates and provisions nests with leafhoppers (Cicadellidae) for its larvae. The species practices progressive provisioning, bringing prey to the developing larva on an as-needed basis rather than mass provisioning. It is one of approximately 190 Bembecinus species worldwide and among ten species recorded from North America north of Mexico.
Bembecinus neglectus
Bembecinus neglectus is a species of sand wasp in the family Crabronidae (formerly treated as Bembicidae). It was first described by Cresson in 1873 and is known to occur in North America. As a member of the tribe Bembicini, it belongs to a group of solitary wasps commonly associated with sandy habitats. The species has been documented in museum collections and through citizen science observations, though detailed biological studies remain limited.
Bembix americana spinolae
Eastern Sand Wasp
A sand wasp subspecies common on barren sand exposures where females dig burrows into loose sand. Adults provision nests with paralyzed flies, with females providing progressively more prey as larvae grow. Formerly placed in family Sphecidae, now classified in Crabronidae. The subspecies is part of a variable species complex with intermediate characteristics between B. americana comata and B. americana spinolae in some populations.
Bembix belfragei
A solitary sand wasp in the genus Bembix, characterized by ground-nesting behavior in loose, sandy substrates. Females excavate burrows to provision with paralyzed flies, exhibiting progressive parental care by delivering prey as larvae develop. Adults of both sexes visit flowers for nectar, particularly composites. The species is part of a diverse North American radiation of sand wasps with complex nesting behaviors.
Bembix occidentalis
sand wasp
Bembix occidentalis is a species of solitary sand wasp in the family Crabronidae. The species was described by W. Fox in 1893 and occurs in Central America and North America. Like other members of the genus Bembix, it is associated with sandy habitats where females excavate burrows to provision with prey for their larvae.
Bembix rugosa
Bembix rugosa is a species of sand wasp in the family Crabronidae, described by J. Parker in 1917. As a member of the genus Bembix, it shares the characteristic behaviors of solitary digger wasps that excavate burrows in sandy substrates and provision nests with paralyzed flies for their larvae. The species occurs in North America, though specific details about its biology and distribution remain limited compared to better-studied congeners such as Bembix americana.
Bicyrtes
Bicyrtes is a genus of large, often brightly colored predatory sand wasps in the family Crabronidae. The genus name derives from the two noticeable projections arising from the rear of the mesosoma. Most species nest in sandy or coarse soils and provision their burrows with paralyzed true bugs, primarily Pentatomidae (stink bugs), though some species utilize Reduviidae (assassin bugs) and Coreidae (leaf-footed bugs) as prey. The genus contains approximately 11 described species distributed across North America, with some species extending into Central and South America and the Caribbean, including the Galápagos Islands.
Bicyrtes capnopterus
Bicyrtes capnopterus is a species of sand wasp in the family Crabronidae. It occurs in Central America and North America. Like other members of the genus Bicyrtes, females are fossorial, excavating burrows in soil to provision with prey for their larval offspring. The species is part of a group of wasps known for preying on true bugs, contributing to natural control of pest populations.
Bicyrtes fodiens
Bicyrtes fodiens is a solitary sand wasp in the family Crabronidae, native to North and Middle America. Like other members of its genus, it is a fossorial species that excavates burrows in sandy or coarse soil to provision with paralyzed true bugs for its larval offspring. The species is part of a guild of beneficial predatory wasps that help control pest insect populations, including stink bugs. Adults are nectar-feeders and can often be observed visiting flowers.
Bicyrtes insidiatrix
Bicyrtes insidiatrix is a species of sand wasp in the family Crabronidae. It is found in North America. Like other members of the genus Bicyrtes, it is a solitary, fossorial wasp that nests in sandy or coarse soils. The species is part of a group of wasps known for preying on true bugs (Hemiptera), though specific prey records for this species are not well documented in the available literature.
Clitemnestra
Clitemnestra is a genus of small sand wasps in the family Crabronidae, containing at least 60 described species. Members of this genus are predatory wasps that hunt various sap-feeding insects, particularly planthoppers, leafhoppers, and related Hemiptera. The genus was revised by Bohart in 2000, which included the merger of the former genus Ochleroptera into Clitemnestra. These wasps are generally inconspicuous due to their small size, with some species measuring only 5–6 millimeters in body length.
Didineis latimana
Didineis latimana is a species of solitary wasp in the family Crabronidae (formerly treated as Bembicidae). It belongs to the subfamily Bembicinae and tribe Alyssontini. The species was described by Malloch and Rohwer in 1930 and is known from North America, with records from Canada including Ontario.
Didineis texana
Didineis texana is a species of sand wasp in the family Crabronidae, subfamily Bembicinae, tribe Alyssontini. It is one of several species in the genus Didineis, which are small to medium-sized wasps adapted to sandy habitats. The species was described by Cresson in 1873 and is known from the southern United States and Mexico. Like other members of its tribe, it is likely a predator of small insects, though specific biological details remain poorly documented.
Epinysson bellus
Epinysson bellus is a species of digger wasp in the family Crabronidae, subfamily Bembicinae, tribe Nyssonini. The species was first described by Cresson in 1882. It belongs to a group of solitary wasps that are part of the diverse nyssonine lineage within the Bembicinae. Like other members of its genus, it is likely a kleptoparasite or predator associated with other ground-nesting wasps, though specific biological details remain poorly documented.
Foxia
Foxia is a genus of cleptoparasitic wasps in the family Crabronidae, subfamily Bembicinae. The genus comprises 10 described species distributed across arid regions of the New World, from the United States south through Chile and Argentina. These wasps are obligate cleptoparasites, meaning they exploit the nests and provisions of other wasp species rather than constructing their own nests or hunting prey directly.
Foxia navajo
Foxia navajo is a species of solitary wasp in the family Crabronidae (formerly Bembicidae), described by Pate in 1938. The genus Foxia belongs to the tribe Nyssonini within the subfamily Bembicinae. Like other members of its tribe, this species is presumed to be a kleptoparasite or predator of other ground-nesting wasps, though specific biological details remain poorly documented. The species is known from North America, with the specific epithet 'navajo' suggesting a connection to the Navajo Nation or southwestern United States region.
Foxia pacifica
Foxia pacifica is a species of sand wasp in the family Crabronidae, subfamily Bembicinae. It was described by Ashmead in 1898. The species is part of the genus Foxia, which belongs to the tribe Nyssonini. Based on the family-level classification and the genus name, members of Foxia are likely cleptoparasitic wasps that exploit the nests of other ground-nesting hymenopterans.
Gorytes smithii
Gorytes smithii is a species of sand wasp in the family Crabronidae, subfamily Bembicinae. It belongs to the tribe Bembicini, which includes solitary wasps that typically nest in sandy soils and provision their burrows with paralyzed prey. The species was described by Cresson in 1880 and occurs in North America. Like other members of the genus Gorytes, it is likely associated with open sandy habitats where it excavates burrows for nesting.
Gorytes venustus
Gorytes venustus is a species of sand wasp in the family Crabronidae, described by Cresson in 1865. It belongs to the tribe Bembicini, a group of solitary wasps known for nesting in sandy soils and provisioning their young with paralyzed prey. The species is documented from North America. As a member of the genus Gorytes, it shares the general biology of related sand wasps, though species-specific details remain limited in available sources.
Gorytes willcoxi
Gorytes willcoxi is a species of sand wasp in the family Crabronidae (formerly placed in Bembicidae), described by Ohl in 2009. The species is known from North America. Like other members of the genus Gorytes, it is a solitary wasp that excavates burrows in sandy soils.
Hapalomellinus
Hapalomellinus is a genus of digger wasps in the family Crabronidae, subfamily Bembicinae. Established by Ashmead in 1899, it belongs to the tribe Bembicini within the subtribe Gorytina. The genus contains at least three described species: Hapalomellinus albitomentosus, H. pulvis, and H. teren. Like other Bembicinae, these wasps are solitary and nest in soil.
Hapalomellinus albitomentosus
Hapalomellinus albitomentosus is a species of sand wasp in the family Crabronidae, originally described by Bradley in 1920 as Gorytes albitomentosus. It belongs to the subfamily Bembicinae, a group of solitary wasps known for nesting in soil and provisioning their young with paralyzed prey. The species is part of the genus Hapalomellinus, which contains only three described species. Records indicate presence in North America.
Harpactus pictifrons
Harpactus pictifrons is a species of sand wasp in the family Crabronidae, first described by W. Fox in 1894. It belongs to the subfamily Bembicinae, a group of solitary wasps known for nesting in sandy soils and provisioning their larvae with paralyzed prey. The species is part of the genus Harpactus, which comprises small to medium-sized wasps with distinctive facial markings. Records indicate presence in North America, specifically in Canada including Alberta. Like other members of its genus, it is presumed to be a predator of true bugs (Hemiptera), though direct observations of its biology are limited.
Hoplisoides
sand wasps
Hoplisoides is a genus of solitary sand wasps in the family Crabronidae, containing at least 70 described species worldwide. These small wasps are characterized by their spotted wings and specialized predatory behavior targeting treehoppers (Membracidae). Females construct short burrows in sandy soil to provision with paralyzed prey for their larvae. The genus is distributed globally except Australia, with approximately 18 species in North America.
Hoplisoides hamatus
Hoplisoides hamatus is a sand wasp in the family Crabronidae. It is found in North America, with records from Canada including British Columbia. Like other members of the genus Hoplisoides, it is a solitary wasp that hunts treehoppers in the family Membracidae as prey for its larvae.
Hoplisoides tricolor
sand wasp
Hoplisoides tricolor is a species of solitary sand wasp in the family Crabronidae, originally described by Cresson in 1868. It is one of approximately eighteen Hoplisoides species found in North America north of Mexico. Like congeners, it is a ground-nesting wasp that hunts treehoppers (Membracidae) as prey for its larvae. The species occurs across Central America and North America, with most Hoplisoides species concentrated in western North America.
Microbembex californica
Microbembex californica is a species of sand wasp in the family Crabronidae, first described by R. Bohart in 1970. It belongs to a genus of small to medium-sized predatory wasps that inhabit sandy environments. Like other members of the Bembicinae, this species likely excavates burrows in soil and provisions them with prey for their larvae. The specific epithet "californica" indicates a geographic association with California.
Nysson
Nysson is a Holarctic genus of kleptoparasitic wasps in the family Crabronidae. Over 100 species are known. These wasps are obligate kleptoparasites, meaning females exploit the food caches of other solitary wasps by locating host burrows, digging them open, destroying the host egg, and replacing it with their own. The larva then consumes the prey provisioned by the host mother.
Nysson intermedius
Nysson intermedius is a species of cuckoo wasp in the family Crabronidae, subfamily Bembicinae. It was described by Viereck in 1908. Members of the genus Nysson are kleptoparasitic, laying their eggs in the nests of other digger wasps, particularly those in the genus Bembix and related taxa. The species occurs in North America.
Nysson lateralis
Nysson lateralis is a species of solitary wasp in the family Crabronidae, distributed across North America. It belongs to the subfamily Bembicinae and tribe Nyssonini, a group that includes kleptoparasitic wasps that exploit the nests of other crabronid wasps. The species was described by Packard in 1867. Like other members of the genus Nysson, it is presumed to be a nest parasite, though specific behavioral details for this species remain poorly documented.
Nysson subtilis
Nysson subtilis is a species of solitary wasp in the family Crabronidae, subfamily Bembicinae. It occurs in North America, with records from Canada including Nova Scotia. The species is a cleptoparasite (cuckoo wasp) that targets nests of other ground-nesting crabronid wasps, particularly those in the genus Hoplisoides. It can be distinguished from congeners by its entirely black front.
Saygorytes
Saygorytes is a genus of sand wasps in the family Crabronidae, established by Nemkov in 2007. The genus comprises approximately seven described species distributed in North America. Members are solitary wasps that visit sunflowers and other plants for nectar. They are part of the diverse community of wasps attracted to extrafloral nectaries on sunflowers in arid and disturbed habitats.
Saygorytes phaleratus
Saygorytes phaleratus is a species of sand wasp in the family Crabronidae. It is found in Central America and North America. The species was originally described by Thomas Say in 1837. As a member of the sand wasp group, it is likely associated with sandy habitats where it hunts for prey to provision its nest.
Steniolia
sand wasps
Steniolia is a genus of solitary sand wasps in the family Crabronidae, containing approximately 15 described species. All North American species are western in distribution, with four species extending only as far north as Mexico. Females are fossorial, constructing single-celled burrows in dry, powdery soil to provision with paralyzed flies. The genus is notable for ornate coloration, elongated mouthparts adapted for nectar feeding, and distinctive nightly sleeping clusters formed by both sexes.
Steniolia obliqua
Steniolia obliqua is a species of sand wasp in the family Crabronidae, originally described as Monedula obliqua by Cresson in 1865. It belongs to a genus of fourteen North American species, all with western distributions. Like other Steniolia species, it is a solitary wasp that hunts flies to provision underground nests for its larvae. The species has been recorded in British Columbia, Canada.
Stictiella emarginata
sand wasp
Stictiella emarginata is a solitary sand wasp in the family Crabronidae, subfamily Bembicinae. It constructs nests in sandy soils and provisions cells with lepidopteran larvae, primarily Noctuidae and Hesperiidae. The species occurs across eastern North America from northern Michigan to the Atlantic Coast, with a flight season concentrated in late June through early August. Nesting behavior includes temporary nest closure, mound leveling, and orientation flights.
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sand wasp
Stictiella formosa is a species of sand wasp in the family Crabronidae, subfamily Bembicinae. It is distributed across Central America and North America. As a member of the Bembicini tribe, it is likely associated with sandy habitats, though specific ecological details are poorly documented in available sources.
Stizus texanus
Stizus texanus is a species of sand wasp in the family Crabronidae, first described by Cresson in 1873. It belongs to the tribe Stizini within the subfamily Bembicinae. The species is distributed in North America and Middle America, with museum holdings documented from the UCR Entomology Research Museum. As a member of the Crabronidae, it is likely a predatory wasp, though specific biological details for this species remain poorly documented in available sources.
Tanyoprymnus
Tanyoprymnus is a genus of digger wasps in the family Crabronidae, subfamily Bembicinae. The genus was established by Cameron in 1905. The best-known species is Tanyoprymnus moneduloides (Packard), which has been studied for its nesting biology. Members of this genus are solitary wasps that construct nests in soil.
Tanyoprymnus moneduloides
Tanyoprymnus moneduloides is a species of sand wasp in the family Crabronidae, subfamily Bembicinae. The species has been the subject of dedicated nesting biology research, with a 1981 study documenting its nest architecture and provisioning behavior. It is present in North America and Middle America according to distribution records. The species was originally described by Packard in 1867.
Zanysson
Zanysson is a strictly New World genus of digger wasps in the family Crabronidae, subfamily Bembicinae. The genus was established by Rohwer in 1921 and remained taxonomically stagnant for over 80 years until the description of Z. gemmatus from Colombia in 2007. Members are assumed to be cleptoparasites of other crabronid wasps, though direct biological observations are lacking for most species. The genus includes at least three described species distributed from the southwestern United States through Mexico to northwestern South America.
Zanysson plesius
Zanysson plesius is a species of wasp in the family Crabronidae, subfamily Bembicinae, first described by Rohwer in 1921. It belongs to the tribe Nyssonini, a group of sand wasps. The species is part of a genus containing at least three recognized species in North America, including Z. texanus and its subspecies. Specimens have been documented in museum collections, including holdings at the UCR Entomology Research Museum.
Zanysson texanus
Zanysson texanus is a species of wasp in the family Crabronidae, subfamily Bembicinae. It was originally described as Nysson texanus by Cresson in 1873. The genus Zanysson belongs to the tribe Nyssonini, a group of solitary wasps commonly known as cuckoo wasps or kleptoparasitic wasps that typically exploit the nests of other wasps. The species epithet 'texanus' indicates a Texas association, likely referring to the type locality or primary distribution.