Running-water
Guides
Ambrysus arizonus
Ambrysus arizonus is a creeping water bug (family Naucoridae) in the Ambrysus guttatipennis species complex. It is distributed in both the United States and Mexico, with recent records extending its known range into Mexico. As a member of the Cryphocricinae subfamily, it shares the characteristic saucer-like, flattened body form and aquatic lifestyle typical of this group.
Ambrysus californicus
California creeping water bug
Ambrysus californicus is a species of creeping water bug in the family Naucoridae. It was described by Montandon in 1897. The species is native to North America and belongs to the subfamily Cryphocricinae, commonly known as saucer bugs due to their round, flat body shape. Like other members of the genus Ambrysus, it is an aquatic insect adapted to life in running streams.
Ameletus validus
combmouthed minnow mayfly
Ameletus validus is a species of mayfly in the family Ameletidae, first described by McDunnough in 1923. It is one of approximately 30 bisexual species recognized in the genus Ameletus across North America. The species occurs in running waters from small headwater streams to large rivers, with documented populations in western North America including Alberta, British Columbia, and the western United States. Taxonomic work has clarified that adult wings are lightly and evenly suffused with brownish-amber, not speckled as previously miskeyed.
Ameletus vernalis
combmouthed minnow mayfly
Ameletus vernalis is a species of combmouthed minnow mayfly in the family Ameletidae. It is found in southwestern Canada and the western United States, particularly in western Alberta where it occurs across diverse watersheds from small headwater streams to large rivers. The species was described by McDunnough in 1924, and its nymphs were associated with adults through field work in southern British Columbia in 1935.
Calopteryx
Jewelwings
Calopteryx is a genus of large damselflies in the family Calopterygidae, commonly known as jewelwings. Males typically display brightly colored wings while females usually have clear wings, though some females develop male-like (androchrome) wing characteristics. The genus lacks a pterostigma, a wing cell found in most other odonates. Species occupy distinct zones within running water systems, with differential thermal adaptations driving habitat segregation.
Carapacea
Carapacea is a suborder of mayflies (Ephemeroptera) characterized by nymphs possessing a distinctive carapace-like structure formed from expanded and fused thoracic tergites. This morphological adaptation distinguishes them from other mayfly suborders. The group includes families such as Baetiscidae and Prosopistomatidae. Carapacean mayflies are primarily found in running water habitats.
Cinygmula reticulata
Cinygmula reticulata is a species of mayfly in the family Heptageniidae, first described by McDunnough in 1934. It is a member of the Nearctic fauna, occurring in North America. Like other members of the genus Cinygmula, it is associated with running water habitats where the aquatic nymphs develop. The species is represented by limited observation records, with only four observations documented on iNaturalist.
Dubiraphia minima
riffle beetle
Dubiraphia minima is a species of riffle beetle in the family Elmidae, first described by Hilsenhoff in 1973. As a member of this family, it is adapted to aquatic or semi-aquatic habitats in flowing water systems. The species is known from North America, with confirmed records from Manitoba and Ontario in Canada.
Elminae
riffle beetles
Elminae is a subfamily of riffle beetles (family Elmidae) comprising at least 120 genera and over 1,300 described species. Members are primarily aquatic, inhabiting running water habitats across all major biogeographic regions. The subfamily exhibits considerable morphological diversity, with some genera showing pronounced secondary sexual dimorphism and miniaturization traits.
Enicocerus
Enicocerus is a subgenus of minute moss beetles within the genus Ochthebius (family Hydraenidae). Species in this subgenus are small aquatic beetles associated with moist microhabitats in running water systems. The subgenus was established by Stephens in 1829 and is currently treated as valid. One documented species, Ochthebius (Enicocerus) colveranus, inhabits algae and moss growth on spray zones of rocks in streams.
Hemerodromiini
Hemerodromiini is a tribe of dance flies (family Empididae) within the subfamily Empidinae. Members of this tribe are small to medium-sized predatory flies characterized by their distinctive leg structure and behavior. The tribe is distinguished from related groups by morphological adaptations associated with their characteristic hunting strategy. Hemerodromiini species are found across various habitats, particularly in association with running water and moist environments.
Himalopsyche
Himalopsyche is a genus of free-living caddisflies in the family Rhyacophilidae, containing more than 40 described species. The genus is primarily distributed in alpine to montane environments across Central and East Asia and North America, with its greatest diversity concentrated in the Himalayas and surrounding mountain ranges. Species within the genus have been divided into five morphological groups: kuldschensis, lepcha, navasi, phryganea, and tibetana. The genus has served as a model system for studying mountain biodiversity, gene flow, and the mountain-geobiodiversity hypothesis due to its endemic distribution in geologically dynamic regions.
caddisflyRhyacophilidaemountain-biodiversityTibeto-Himalayanaquatic-insectphylogeographygenome-assemblyindicator-speciesgene-flowHengduan-MountainsHimalayasalpinemontanerunning-waterfree-living-caddisflycocoon-makerpredatory-larvaewater-quality-monitoringadaptive-evolutionmountain-geobiodiversity-hypothesisHydropsyche
Spotted Sedges
Hydropsyche is a large genus of netspinning caddisflies comprising at least 260 described species worldwide. Larvae construct fixed capture nets across flowing water to filter food particles, a behavior that defines the family Hydropsychidae. The genus exhibits substantial diversity in the Aegean region and Mediterranean, with numerous endemic species on Greek islands. Both larvae and adults have been extensively studied for their morphological characteristics, respiratory physiology, and responses to aquatic hypoxia.
Leucotrichiinae
Leucotrichiinae is a subfamily of microcaddisflies (Hydroptilidae) in the order Trichoptera. The subfamily includes genera such as Betrichia and Mejicanotrichia, distributed primarily in the Neotropical region. Diagnostic characteristics involve features of the antennae, wing venation, and male genitalia. Larvae of at least some genera are dorsoventrally flattened with numerous broad setae and inhabit high-flow aquatic environments.
Leucrocuta minerva
Leucrocuta minerva is a species of mayfly in the family Heptageniidae, described by McDunnough in 1924. It belongs to a genus of small to medium-sized flat-headed mayflies commonly known as "little yellow quills" or similar vernacular names. Species in this genus are typically associated with running water habitats. The specific epithet "minerva" refers to the Roman goddess of wisdom, though the etymological reasoning behind this choice is not documented in available sources.
Lutrochus
travertine beetles
Lutrochus is the sole genus in the family Lutrochidae, a group of small aquatic beetles commonly called travertine beetles. The genus contains approximately 21 species distributed from the southern United States through Central America to Brazil. Both adults and larvae inhabit shallow, fast-flowing streams where they are specifically associated with submerged decaying wood and travertine deposits.
Macronychini
riffle beetles
Macronychini is a tribe of riffle beetles within the family Elmidae, first described by Gistel in 1848. The tribe comprises more than 20 genera and over 80 described species. Members of this tribe are aquatic beetles associated with running water habitats. The tribe is taxonomically well-established but individual species continue to be described, particularly from underexplored regions such as China.
Optioservus castanipennis
Optioservus castanipennis is a species of riffle beetle in the family Elmidae, first described by Fall in 1925. It belongs to a genus of small aquatic beetles inhabiting running water systems. The species is recognized as part of the Nearctic fauna and has been subject to taxonomic revision based on male genitalia and elytral characteristics.
Optioservus divergens
Optioservus divergens is a species of riffle beetle in the family Elmidae. The species was described by LeConte in 1874 and is currently accepted under this name, though it was formerly classified in the genus Heterlimnius. Riffle beetles in this family are aquatic insects typically associated with running water habitats.
Optioservus phaeus
Scott optioservus riffle beetle
Optioservus phaeus is a species of riffle beetle (family Elmidae) described by White in 1978. It belongs to a genus of small aquatic beetles inhabiting running waters. The species was described as part of a taxonomic revision based on male genitalia, elytral patterns, and geographic distribution. It is known from a highly restricted range in Kansas.
Optioservus quadrimaculatus
four-spotted riffle beetle
Optioservus quadrimaculatus is a species of riffle beetle in the family Elmidae, described by Horn in 1870. The species is found in North America, with records from Canadian provinces including Alberta, British Columbia, and Saskatchewan. Riffle beetles in this family are aquatic insects associated with running water habitats. The specific epithet "quadrimaculatus" refers to four spots, likely describing a distinctive elytral pattern.
Oreodytes humboldtensis
Oreodytes humboldtensis is a species of predaceous diving beetle in the family Dytiscidae, described by Zimmerman in 1985. As a member of the genus Oreodytes, it belongs to a group of small diving beetles primarily associated with running water habitats. The species epithet suggests a connection to the Humboldt region, likely referring to Humboldt County or the broader Humboldt Bay area of coastal northern California. Like other Oreodytes species, it is presumed to be an aquatic predator in freshwater systems.
Petrophila fulicalis
Feather-edged Petrophila
Petrophila fulicalis is a crambid moth described by James Brackenridge Clemens in 1860. It belongs to a species group that has been taxonomically clarified using citizen science data, distinguishing it from similar species P. confusalis and P. bella. Adults are diurnal and often found near running water. The species has a broad North American distribution with two distinct population centers: eastern records from the Great Lakes and Appalachian regions, and western records from the southwestern United States and northern Mexico.
Plectrocnemia
tube maker caddisflies
Plectrocnemia is a genus of tube maker caddisflies in the family Polycentropodidae comprising more than 120 described species. Larvae are aquatic predators that construct silken capture nets to intercept prey. The genus has been extensively studied for its larval silk production, vibration-mediated predatory behavior, and population genetics. Species occur across Europe and into western Asia, with detailed biological information available for several well-studied species including P. conspersa and P. brevis.
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predatory-behaviourvariations-in-the-frequencymore-marked-effectvariations-in-amplitudestage-2orientation-and-displacement-towards-the-lurestage-3capture-of-the-lure-with-mandibleslarvae-live-in-running-waterscatch-with-a-netpreymainly-chironomid-larvae-and-oligochaeteseffect-of-vibrations-generated-by-prey-in-the-netvery-irregularly-woven-netopen-ended-dwelling-tube-at-both-endsvibration-weakly-dampenedfrequency-does-not-changevibration-excitesattentionorientation-and-movementcapture-attemptsorientation-and-movement-to-preythe-fasterthe-more-the-vibration-exceeds-0.28-Hzfrequencies-of-0.15-to-0.28-Hzlead-to-incomplete-reactionsas-if-the-larvae-were-confusedfrequencies-below-0.075-Hzgenerate-building-movementsbuilding-behavior-instead-of-prey-captureclosely-connectedrecruitment-kin-and-spatial-genetic-structureoviposition-and-genetic-relatednessstream-dwelling-caddisbeginning-of-the-aquatic-phasefour-sample-dateswithin-one-generationmean-relatedness-coefficientwithin-reared-egg-massesdiffered-significantlypopulation-as-a-wholemarkers-sufficiently-powerfulidentify-groups-of-siblingssmall-contribution-from-a-second-malemean-relatedness-within-spatially-structured-groupsdid-not-differ-from-backgroundsiblings-disperse-away-from-each-otherkin-structure-does-not-persistchanges-in-spatial-genetic-structure-late-in-larval-lifeneighbouring-larvae-less-closely-relatedapproaching-onset-of-pupationsurvival-through-egg-stage-and-early-larval-lifevery-highgreater-than-50%non-social-insectconsequence-of-colonial-netbriefly-occupied-by-first-instar-larvaelack-of-spatial-genetic-structurehigh-survivalrefute-patchy-recruitment-hypothesislarva-of-Plectrocnemia-renettaincluding-discriminatory-matrixlarvae-of-Plectrocnemia-Stephens-1836-species-of-Greecemorphology-of-final-instar-larvamost-important-diagnostic-features-illustratedpreliminary-discriminatory-matrixstrongly-different-in-lengthseparated-from-each-othermuscle-attachment-spots-on-head-capsulenumber-and-length-of-setae-on-abdominal-sternum-IXreported-from-Cyprus-Turkey-Greek-islandsexploit-wide-range-of-ecological-nichesfive-genomes-publishedvariable-qualitiessingle-Oxford-Nanopore-flow-cellde-novo-assembly-methods-comparedassembly-of-low-coverage-Nanopore-readssubsequent-polishingyielded-highest-genome-qualitycontiguity-and-BUSCO-completenessshortest-genomes-to-dateextend-knowledge-of-genome-sizegenomic-region-encodes-for-light-L-chain-fibroinprotein-component-of-larval-caddisfly-silkidentified-and-comparednew-genomic-resourcesamong-highest-quality-Trichoptera-genomesincrease-knowledgebasis-for-phylogenomic-and-comparative-genomic-studiesrevised-key-to-larvaedistinguishes-previously-unknown-larvaother-two-British-speciesnotes-on-larval-habitat-life-cycle-and-identification-of-adultgenetic-population-structure-and-neighbourhood-population-size-estimatesrole-of-history-and-scale-of-colonisationno-genetic-differentiation-between-sites-up-to-20-kmdespite-population-sizes-suggesting-genetic-driftgenetic-differentiation-between-populations-separated-by-more-than-20-kmneighbourhood-population-size-implausibly-smallevolutionary-processes-do-not-explain-differentiationrelatively-short-flights-by-winged-adultsspread-over-number-of-generationsfound-small-often-temporary-populationsgrow-and-exchange-genes-with-larger-permanent-local-populationsamplify-effects-of-initial-gene-flowsubstantial-gaps-between-regions-containing-suitable-habitat-patchesreduce-number-of-colonisation-eventsgenetic-patterns-may-date-from-time-last-colonisedecological-studies-rarely-examined-dynamics-over-larger-geographical-scalescentral-to-persistence-and-spreadbiology-of-Plectrocnemia-latissimarivers-and-streams-of-Central-Caucasus-northern-slopessprings-streams-and-riversrequires-high-oxygen-concentrationgood-indicator-of-water-qualityaspects-of-biology-reportedWikipedia-summaryrank-GENUSstatus-ACCEPTEDmatch-type-HIGHERRANKdistribution-records-DK-NO-SE-Vermont-US-USscientific-nameauthorship-Stephens-1836classification-Eukaryota-Animalia-Arthropoda-Hexapoda-Insecta-Trichoptera-Hydropsychoidea-Polycentropodidae-Polycentropodinae-Plectrocnemiascientific-name-Plectrocnemiagroup-caddisflieskingdom-Metazoainstructionsfill-all-fieldsif-a-field-cannot-be-supported-return-nulldo-not-repeat-information-across-fieldskeep-each-section-focused-on-its-purposeprovide-useful-detail-where-possiblecritical-rulesfactual-correctness-over-completenessclarity-over-verbosityusefulness-over-speculationif-information-is-not-clearly-supported-return-nulldo-not-infer-species-level-traits-from-higher-taxa-unless-explicitly-justifieddo-not-repeat-the-same-information-across-multiple-fieldseach-field-must-contain-unique-non-overlapping-contentavoid-vague-generalizationslike-most-insectstypically-feeds-on-plantsuse-cautious-language-when-necessaryhas-been-observedis-known-todo-not-fabricatebehaviorsdietlife-cycle-detailshost-relationshipsfield-intentsummary-high-level-overview-3-5-sentencesappearance-physical-description-onlyidentif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Net Tube Caddisflies, tube-making caddisflies, trumpet-net caddisflies
Psychomyiidae is a family of tube-making caddisflies (order Trichoptera) characterized by larvae that construct silken galleries or tubes on submerged surfaces. Adults are typically small with reduced wing venation, including absence of forewing fork I and shortened hindwing forks. The family is distinguished from similar Polycentropodidae by leg spur formula 2-4-4, short thyridial cell, and male genitalia with elongate preanal appendages and reduced tergum IX. Larvae possess separated submental sclerites, broad foretrochantin, and whip-like pupal mandible apices. The family contains approximately 10 extant genera and has a global distribution with notable diversity in the Oriental and Western Palearctic regions.
Rhagovelia
smaller water striders, riffle bugs
Rhagovelia is a large genus of small aquatic bugs in the family Veliidae, with over 390 described species. Members are distinguished by a swimming fan on the distal tarsomere of the middle leg, an adaptation that enables rapid maneuvering on water surfaces. The genus has colonized lotic freshwater systems and coastal marine environments across the Americas, with additional species in Africa, Asia, and Oceania. Evolutionary origins trace to gene duplications ('geisha' and 'mother-of-geisha') that created this propelling fan structure, allowing exploitation of faster-flowing and more turbulent streams than relatives lacking this adaptation.
Rhagovelia torreyana
Rhagovelia torreyana is a species of small water strider in the family Veliidae, described by Bacon in 1956. It belongs to the genus Rhagovelia, a diverse group of semiaquatic bugs commonly found in running water habitats. The species is known from North America, though detailed biological information remains limited in available sources. Like other members of the family Veliidae, it is adapted to life on the water surface, using surface tension to move and forage.
Rhithrogena
March Browns, Olive Uprights
Rhithrogena is a genus of flatheaded mayflies in the family Heptageniidae, commonly known as March Browns or Olive Uprights. The genus contains approximately 153 species distributed across the Holarctic region. Larvae are primarily scrapers that feed on periphyton from stones in running waters. Several species serve as important food sources for fish and have been studied for their bioindicator potential in freshwater monitoring.
Rhyacophila
caseless caddisflies
Rhyacophila is a large genus of caddisflies (Trichoptera) in the family Rhyacophilidae, comprising at least 640 described species. The genus is cosmopolitan in distribution, with significant diversity in Europe, Asia, and North America. Larvae are commonly known as 'caseless caddis' because they do not construct portable cases, instead using silk to anchor themselves to substrates in running water. The genus exhibits high endemism in certain regions, with multiple species restricted to specific mountain ranges or islands.
Shipsa rotunda
Intrepid Forestfly
Shipsa rotunda is a Nearctic stonefly species in the family Nemouridae with an extensive distribution across Canada and the United States. Despite its wide geographic range, the species exhibits remarkably little morphological variation in both males and females. Reproductive terminalia have been examined using scanning electron microscopy from specimens across its range, revealing consistent structural features. The species was originally described as Nemoura rotunda by Claassen in 1923 and later transferred to the genus Shipsa.
Tanypezidae
Stretched-foot Flies
Tanypezidae is a small family of acalyptrate Diptera containing 28 species in two genera. The family is primarily New World in distribution, with Tanypeza (2 species) occurring in North America and the Palearctic, and Neotanypeza (26 species) restricted to the Neotropics. Species are characterized by their relatively large size, semispherical heads, stout bodies, and notably long, thin legs that give the family its common name. The biology of most species remains poorly known.
Torrenticolidae
torrent mites
Torrenticolidae is a family of water mites (Acari: Hydrachnidia) in the order Trombidiformes. Members are tiny, heavily sclerotized, crawling arthropods found in running freshwater habitats. The family contains approximately 5 genera and more than 400 described species, with the genus Torrenticola being the most diverse and widespread. Species diversity is highest in tropical regions, though the family occurs on all continents except Antarctica.