Aquatic
Guides
Acentropinae
China-mark Moths
Acentropinae is a subfamily of aquatic moths within the family Crambidae, comprising approximately 730 species in 78 genera. It represents the largest lineage of moths with at least one life stage adapted to aquatic environments. The subfamily includes two tribes: Argyractini and Acentropini (the latter including the former Nymphulinae). Larvae are aquatic and feed on submerged vegetation, while adults are terrestrial but associated with wetland habitats.
Acentropini
China-mark moths
Acentropini is a tribe of aquatic moths within the family Crambidae, commonly known as china-mark moths. Adults are small to medium-sized moths with wingspans typically ranging from 15–35 mm. The tribe is distinguished by larvae that are fully aquatic, breathing through gills and feeding on submerged aquatic plants. These moths are found across temperate and tropical regions worldwide, with highest diversity in the Old World tropics.
Acilius abbreviatus
Abbreviated Predaceous Diving Beetle
Acilius abbreviatus is a predaceous diving beetle in the family Dytiscidae. It is found in North America, with records from Alberta and British Columbia in Canada. As a member of the genus Acilius, it is an aquatic predator inhabiting freshwater environments.
Acilius athabascae
Athabascan Predaceous Diving Beetle
Acilius athabascae is a species of predaceous diving beetle described by Larson in 1975. It belongs to the genus Acilius, a group of large diving beetles known for their streamlined bodies and predatory habits in aquatic environments. The species is found in North America, with records from Alberta and British Columbia in Canada.
Acilius sylvanus
Woodland Predaceous Diving Beetle
Acilius sylvanus is a predaceous diving beetle described by Hilsenhoff in 1975. It belongs to the family Dytiscidae, a group of aquatic beetles adapted for underwater predation. The species is known from forested wetland habitats in North America.
Acneus
water-penny beetles
Acneus is a genus of water-penny beetles (family Psephenidae) containing four described species. The genus is characterized by flattened, oval larvae that cling to submerged rocks in flowing water. Adults are terrestrial and rarely encountered. The genus was established by Horn in 1880 and is restricted to western North America.
Acneus quadrimaculatus
water penny beetle
Acneus quadrimaculatus is a species of water penny beetle in the family Psephenidae. It is endemic to the United States with confirmed records from California and Oregon. Adults are found in riparian vegetation near aquatic habitats. The species was described by Horn in 1880.
Aeshna palmata
Paddle-tailed Darner
Aeshna palmata, commonly known as the Paddle-tailed Darner, is a species of dragonfly in the family Aeshnidae. It is widely distributed across western Canada and the western United States. The species is named for its distinctive paddle-shaped cerci (appendages at the tip of the abdomen). It inhabits lakes, ponds, and slow-moving streams, typically with dense vegetation along the shorelines. The species was first scientifically described by Hermann Hagen in 1856.
Agabus
Agabus is a large genus of predatory aquatic beetles in the family Dytiscidae, established by William Elford Leach in 1817. Adults range from 5 to 14 mm in length. The genus has a primarily Holarctic distribution, with limited representation in the Afrotropic and Neotropic realms. Some species are of conservation concern, with A. clypealis, A. discicollis, and A. hozgargantae listed as endangered by the IUCN.
Agabus ajax
Agabus ajax is a predaceous diving beetle in the family Dytiscidae. The species was described by Fall in 1922. It is known from North America, with records from Alberta and British Columbia in Canada. As a member of Agabus, it is an aquatic predator.
Agabus antennatus
Agabus antennatus is a species of predaceous diving beetle in the family Dytiscidae, first described by Leech in 1939. It is found in North America, with records from Alberta and British Columbia in Canada. As a member of the genus Agabus, it is an aquatic predator inhabiting freshwater environments.
Agabus canadensis
Agabus canadensis is a predaceous diving beetle in the family Dytiscidae, described by Fall in 1922. It is found in North America, with records from Canada (Alberta, British Columbia) and the United States. As a member of the genus Agabus, it is an aquatic predator inhabiting freshwater environments.
Agabus clavicornis
Agabus clavicornis is a predaceous diving beetle in the family Dytiscidae. It is distributed across the Holarctic region, occurring in both North America and the Palearctic. The species was described by Sharp in 1882. Like other members of the genus Agabus, it is an aquatic predator.
Agabus crassipes
Agabus crassipes is a diving beetle species in the family Dytiscidae, described by Fall in 1922. It is native to North America, with records from western Canada including Alberta and British Columbia. As a member of the genus Agabus, it is an aquatic predator inhabiting freshwater environments. The specific epithet 'crassipes' refers to its stout or thick legs.
Agabus falli
Agabus falli is a species of predaceous diving beetle in the family Dytiscidae. It is known from North America, with records from Alberta and Manitoba in Canada. As a member of the genus Agabus, it is an aquatic predator inhabiting freshwater environments. The species was described by Zimmermann in 1934.
Agabus punctulatus
Agabus punctulatus is a predaceous diving beetle in the family Dytiscidae. It inhabits freshwater environments and is distributed across North America, with records from western Canada including Alberta and British Columbia. As a member of the genus Agabus, it is part of a diverse group of aquatic beetles adapted to life in ponds, lakes, and slow-moving waters.
Agabus taeniolatus
Agabus taeniolatus is a species of predaceous diving beetle in the family Dytiscidae. It belongs to the large genus Agabus, which contains numerous aquatic beetle species found primarily in the Holarctic region. This species is documented from North America and shares the general ecological characteristics of the genus as an aquatic predator in freshwater habitats.
Agabus tristis
Agabus tristis is a predaceous diving beetle in the family Dytiscidae. It belongs to the tristis species group, which has a Holarctic distribution. The species occurs in North America and the Palearctic region. It is one of two North American members of the tristis group, alongside A. leptapsis.
Allocapnia rickeri
Midwest Snowfly
Allocapnia rickeri is a small winter stonefly in the family Capniidae, commonly known as the Midwest Snowfly. It is one of numerous small, dark stoneflies in the genus Allocapnia that emerge during cold months when few other insects are active. The species has been documented across the central and eastern United States. Like other capniids, it is associated with clean, cold streams and is an important indicator of water quality.
winter-stoneflybioindicatorcoldwaterPlecopteraCapniidaeAllocapnialoticemergencebrachypteryapterygenitalia-identificationFrison-1942Midwestsoutheastern-USclean-water-indicatorJanuary-Marchsmall-stoneflywingless-femalestream-insectshreddergathererseasonal-resourcewater-qualityaquatic-insectterrestrial-adultshort-lived-adultovipositionsubmerged-eggshigh-dissolved-oxygenlow-temperaturecentral-USeastern-USAlabamaArkansasDelawareGeorgiaIllinoishexapodhemimetabolousEuholognathaNemouroideaArctoperlariaInsectaArthropodaAnimaliaGBIFCatalogue-of-LifeiNaturalistNCBItaxonomyaccepted-species1942FrisonRickerMidwest-Snowflysnowflysmall-dark-stoneflyclean-streamsriverswell-oxygenatedlotic-habitatcold-monthswinter-activitywing-reductionfemale-apterymale-flightepiproctparaproctterminaliataxonomic-revisioncongenersdistribution-recordsobservations9-observationseukaryotemetazoanarthropodinsectstoneflywinter-emergingJanuaryFebruaryMarchcold-weathernear-freezingbelow-freezingwater-surfacesubmerged-substratesallochthonous-organic-materialstream-ecosystemsseasonal-food-resourceinsectivorous-birdspredatorsscarce-preyunpollutedno-economic-importancestream-monitoringwater-quality-indicatorhigh-quality-coldwatermicroscopic-examinationtaxonomic-keysmale-terminaliareliable-separationgenitalic-examinationoverlapping-distributionsimilar-habitatsmall-sizeunder-10-mmbody-lengthreduced-wingsabsent-wingsfully-developed-wingsspecific-identificationpublished-descriptionsillustrationssubsequent-revisionscharacteristicfamily-Capniidaecommon-nameextended-nymphal-periodone-to-two-yearsshort-liveddoes-not-feedaquatic-nymphclean-cold-streamslow-temperaturesyear-roundwinter-monthsJanuary-through-Marchfamilycentered-Midwestextends-southeasternUnited-Statesdocumentedappearsmost-reliablydistinguishedsubtle-differencesterminal-abdominal-structuresshould-be-comparedagainstpublishedsubsequentgenus-levelcharacterizedreducedabsentfemalesfully-developedmalesrequires-examinationmale-genitaliastructureparaproctsreliableseparationoverlapssimilarmanyexternallydefinitivereliesmicroscopicexaminationcomparisonkeysusedbiologicalindicatorprogramspresenceindicatescoldconditionsno-directeconomicimportanceshreddersgatherersprocessingallochthonousorganicmaterialstreamecosystemsseasonalfoodresourceinsectivorousbirdsotherwhenalternativepreyscarceserveshigh-qualityhabitatsdevelopmentaquaticnymphalstagesterrestrialadultstagenymphsdevelopstreamsextendedperiodlikelyonetwoyearsbasedrelatedspeciesadultsdo-notfeedactiveduringweatherairtemperaturesmaynearbelowfreezingwingedcapableflightwinglessshort-wingedremainwatersurfacematingoccurwinterenteringdepositeggssubmergedsubstratessmallcommonlyknownnumerousdarkemergefewinsectscentraleasternassociatedcleanimportantundermmbodylengthmembersgenuswingspossessfullydevelopedspecificidentificationlevelwithinrequiresmalegenitaliaparticularlymostreliablysubtledifferencestheseterminalabdominalstructuresshouldcompareddescriptionstaxonomicrevisionswinter-emergingmaintainlowhighdissolvedoxygenlevelsthroughoutyearUnitedStatesdistributioncenteredextendssoutheasternmonthstypicallythroughthisactivitygivesrisecommonnamedonotprovidesqualitymonitoringnodirecthabitatmorphologysizegenitalicAmbrysus
creeping water bugs, saucer bugs
Ambrysus is a genus of creeping water bugs (family Naucoridae) containing more than 90 described species. These aquatic insects are found primarily in running streams across the New World, from northern North America to Argentina, with greatest diversity in Mexico. They possess flattened, rounded bodies and powerful claws for clinging to rocks in turbulent water. The genus includes two subgenera: Ambrysus (Syncollus) and Ambrysus (Ambrysus).
Ambrysus californicus
California creeping water bug
Ambrysus californicus is a species of creeping water bug in the family Naucoridae. It was described by Montandon in 1897. The species is native to North America and belongs to the subfamily Cryphocricinae, commonly known as saucer bugs due to their round, flat body shape. Like other members of the genus Ambrysus, it is an aquatic insect adapted to life in running streams.
Ambrysus circumcinctus
creeping water bug
Ambrysus circumcinctus is a species of creeping water bug in the family Naucoridae, distributed across Central America and North America. As a member of the subfamily Cryphocricinae, it inhabits aquatic environments including running streams. The genus Ambrysus is widespread in the New World, with its greatest diversity in Mexico. Like other naucorids, this species has adaptations for clinging to substrates in flowing water.
Ambrysus hungerfordi
A creeping water bug in the family Naucoridae, described by Usinger in 1946. It belongs to the genus Ambrysus, which is widespread in the New World from northern North America to Argentina, with greatest diversity in Mexican running streams. Like other members of Cryphocricinae, it is aquatic in both nymphal and adult stages.
Ambrysus lunatus
creeping water bug
Ambrysus lunatus is a species of creeping water bug in the family Naucoridae. It occurs in Central America and North America. The subspecies A. lunatus lunatus has been described with documented immature stages. Like other members of the genus Ambrysus, it is an aquatic predator inhabiting running streams.
Ambrysus pudicus
creeping water bug, saucer bug
Ambrysus pudicus is a species of creeping water bug (family Naucoridae) described by Carl Stål in 1862. It belongs to the diverse New World genus Ambrysus, which is distributed from northern North America to Argentina with highest diversity in Mexico. Like other members of its family, it is an aquatic predator inhabiting running streams. The specific epithet 'pudicus' (Latin for 'bashful' or 'modest') is shared with the unrelated pitcher plant Nepenthes pudica, which was named for its hidden underground traps.
Amphizoa
troutstream beetles
Amphizoa is a monogeneric genus of aquatic beetles, the sole representative of the family Amphizoidae. These beetles are commonly called troutstream beetles due to their association with cold, flowing mountain waters. The genus contains five known species, with three distributed in western North America and two in the eastern Palearctic region (China and North Korea). Adults and larvae are predatory, feeding primarily on stonefly larvae. When disturbed, adults release a yellowish, cantaloupe-scented fluid from the anus as a chemical defense.
Amphizoa lecontei
Trout-stream beetle
Amphizoa lecontei is an aquatic beetle species in the family Amphizoidae, commonly known as the trout-stream beetle. Adults measure 11.5–16 mm in body length and possess a diagnostic carina on the fifth interval of the elytron. The species is restricted to western North America, with concentrations in the Rocky Mountains. It is one of few beetles in its family and represents a distinctive lineage of stream-dwelling coleopterans.
Anabolia
Anabolia is a genus of northern caddisflies in the family Limnephilidae, containing approximately 18 described species. Species within this genus inhabit both temporary and permanent aquatic environments, with some species adapted to fluctuating water levels. Larvae exhibit rapid growth rates and extended hatching periods. Adults of some species use sex pheromones for mate attraction.
Anacaenini
Anacaenini is a tribe of small aquatic beetles in the family Hydrophilidae, containing 256 species across 6 genera. Members inhabit freshwater environments and are characterized by their compact body form and aquatic adaptations. The tribe was established by Hansen in 1991 and is classified within the subfamily Hydrophilinae.
Ancyronyx
spider water beetles, spider riffle beetles
Ancyronyx is a genus of aquatic riffle beetles in the family Elmidae, commonly known as spider water beetles or spider riffle beetles. The genus comprises 21 described species distributed across North America, South Asia, China, and Southeast Asia, with the Philippines recognized as a center of diversity containing 11 endemic species. Members are obligate inhabitants of lotic freshwater environments, where both adults and larvae cling to submerged substrates using their elongated legs and strong claws. The genus is notable for its potential use as a bioindicator of water quality due to its sensitivity to pollution and oxygen levels.
Anoplius depressipes
Anoplius depressipes is a spider wasp in the family Pompilidae, notable for its specialized hunting of fishing spiders in the genus Dolomedes. The species exhibits remarkable aquatic adaptations, including the ability to walk on water and dive underwater for several minutes to capture prey that attempts to escape beneath the surface. This wasp represents a striking example of predator-prey coevolution with semi-aquatic spiders.
Apsectrotanypus johnsoni
Apsectrotanypus johnsoni is a species of non-biting midge in the family Chironomidae, subfamily Tanypodinae. It was described by Coquillett in 1901. As a member of the tribe Macropelopini, it belongs to a group of predatory midges whose larvae inhabit aquatic environments. The species is documented in the Catalogue of Life and GBIF as an accepted taxon, with 25 observations recorded on iNaturalist.
Apsilops
Apsilops is a genus of ichneumonid wasps in the subfamily Cryptinae, containing approximately nine described species. Members are parasitoids of lepidopteran larvae, with documented associations including aquatic crambid moths and rice pests. The genus includes both aquatic and terrestrial species, reflecting diverse habitat specializations among its members.
Arcola
Arcola is a genus of snout moths (Lepidoptera: Pyralidae) in the subfamily Phycitinae, described by Shaffer in 1995. The genus includes species associated with aquatic vegetation, most notably Arcola malloi (Alligatorweed Stem Borer), which has been employed as a biological control agent for the invasive aquatic weed alligatorweed (Alternanthera philoxeroides) in the United States. The genus belongs to a diverse group of pyralid moths whose larvae typically feed on plant material.
Argyractini
Argyractini is a tribe of aquatic moths within the subfamily Acentropinae (Crambidae). The tribe includes several genera of small to medium-sized moths whose larvae are adapted to life in freshwater habitats. Adults are generally nocturnal and are attracted to light. The group is primarily distributed in the New World, with most species occurring in the Neotropical region.
Atherix lantha
watersnipe fly
Atherix lantha is a species of watersnipe fly in the family Athericidae. The larvae are aquatic, piercing-sucking predators that feed on other aquatic invertebrates. The species has a univoltine life cycle with overwintering fifth-instar larvae. Mass egg-laying behavior results in large aggregations of dead females attached to egg masses.
Atractelmis wawona
Wawona Riffle Beetle
Atractelmis wawona is a species of riffle beetle in the family Elmidae, described by Chandler in 1954. The common name 'Wawona Riffle Beetle' references Wawona, a location in California's Sierra Nevada. Like other elmids, it inhabits aquatic environments with flowing water. Very few observations or detailed studies of this species have been published.
Atrichomelina pubera
Atrichomelina pubera is a sciomyzid fly whose larvae are aquatic or semi-aquatic predators of freshwater snails. The species has been successfully reared in laboratory conditions, enabling documentation of its complete immature stages and feeding behavior. Its biology was formally described in a 1960 study focusing on life history and developmental morphology.
Bagoinae
Bagoinae is a subfamily of weevils (Curculionidae) containing the genus Bagous and related genera. Members are primarily associated with aquatic and semi-aquatic habitats, though some occupy terrestrial environments. The group has been variously classified, with some treatments placing it within Molytinae. The subfamily includes approximately 130 species in the Palaearctic region, with the genus Bagous distributed globally except Central and South America.
Bagous
Bagous is a genus of semi-aquatic weevils in the family Curculionidae containing at least 360 described species. Many species are associated with aquatic or semi-aquatic habitats, with several species developed as biological control agents for invasive aquatic plants, particularly Hydrilla verticillata. The genus includes both native and introduced species with varying degrees of host specificity. Larvae typically develop within plant stems while adults feed on stems and leaves.
Bagous bituberosus
Bagous bituberosus is a species of true weevil in the family Curculionidae, first described by John Lawrence LeConte in 1876. The species is known from North America, with confirmed records from Canada and the United States. Like other members of the genus Bagous, it is associated with aquatic or semi-aquatic habitats. The specific epithet 'bituberosus' refers to a morphological feature of two tubercles or protuberances.
Bellura anoa
Arrowhead Diver Moth
Bellura anoa is a North American moth in the family Noctuidae. Like other members of its genus, it is associated with aquatic habitats during its larval stage. The species is poorly studied compared to its congener Bellura melanopyga, with limited published biological data available.
Bellura brehmei
Narrow-leaved Cattail Diver Moth
Bellura brehmei is a North American moth in the family Noctuidae, commonly known as the Narrow-leaved Cattail Diver Moth. It is one of approximately 800 species of Lepidoptera with aquatic or semi-aquatic larvae. Like other members of its genus, the caterpillar likely inhabits wetland environments and feeds on aquatic vegetation, though specific details for this species remain poorly documented.
Bellura gortynoides
White-tailed Diver Moth, white-tailed diver
Bellura gortynoides, commonly known as the white-tailed diver moth, is a North American noctuid moth whose aquatic caterpillars live submerged in freshwater habitats. The larvae feed on aquatic plants, including water lilies, and breathe by periodically surfacing to capture air through specialized spiracles. Adults are strictly nocturnal. The species belongs to the poorly studied group of aquatic moths, which have received limited scientific attention despite their unusual ecology.
Belostoma bakeri
giant water bug
Belostoma bakeri is a giant water bug in the family Belostomatidae, described by Montandon in 1913. The species is distributed across Central America and North America. Research at Montezuma Well, Arizona has established it as a significant top predator in fishless desert spring ecosystems, where it contributes substantially to secondary consumer production and trophic energy flow.
Belostoma confusum
Belostoma confusum is a species of giant water bug in the family Belostomatidae, described by Lauck in 1959. As a member of this family, it is an aquatic predator found in freshwater habitats. The species occurs in North America and Middle America based on distribution records. Like other Belostoma species, it possesses raptorial forelegs adapted for capturing prey and likely exhibits the reversed sexual size dimorphism characteristic of the genus, where females are larger than males and males carry eggs on their backs.
Belostoma minor
giant water bug
Belostoma minor is a species of giant water bug in the family Belostomatidae. It occurs in the northern Caribbean bioregion, with documented records from Cuba, the Dominican Republic, Jamaica, Puerto Rico, the U.S. Virgin Islands, and southern Florida. Like other members of its genus, it is an aquatic predator. The species was described by Palisot de Beauvois in 1820.
Belostoma testaceum
giant water bug
Belostoma testaceum is a species of giant water bug in the family Belostomatidae. It is found in the eastern United States, ranging from New York south to southern Florida and west to Texas and Michigan. Like other members of its family, it is an aquatic predator inhabiting freshwater environments.
Belostomatidae
giant water bugs, toe-biters, Indian toe-biters, electric-light bugs, alligator ticks, alligator fleas
Belostomatidae is a family of large freshwater hemipteran insects comprising approximately 170 species worldwide. Members are the largest insects in the order Hemiptera, with some species exceeding 12 cm in length. They are aggressive predators found in ponds, marshes, and slow-flowing streams, feeding on aquatic invertebrates, fish, amphibians, and even small vertebrates. The family exhibits remarkable paternal care, with males of many species carrying eggs on their backs until hatching. Several species are consumed as food in Southeast Asia.
Belostomatinae
giant water bugs
Belostomatinae is a subfamily of large aquatic true bugs within the family Belostomatidae. Members are commonly known as giant water bugs or toe-biters. They are among the largest insects in the order Hemiptera and are entirely aquatic as adults. The subfamily includes genera such as Belostoma and Appasus, distinguished from the other subfamily Lethocerinae by their smaller adult size and different respiratory strategies.
Benacus
Benacus is a genus of giant water bugs in the family Belostomatidae, subfamily Lethocerinae. The genus was established by Stål in 1861 and contains species of large aquatic predatory insects. Spermatogenesis has been studied cytologically in Benacus griseus, revealing 28 spermatogonial chromosomes and distinct cytoplasmic inclusions including chondriosomes and Golgi bodies during sperm development.
Bezzia nobilis
Bezzia nobilis is a biting midge in the family Ceratopogonidae, widely considered one of the most common species in the genus Bezzia. It has a notably broad geographic distribution spanning Eurasia, North America, Central America, and South America. Adults are distinguished by black and yellow banded legs, while larvae and pupae have characteristic coloration and morphological features. The species is predatory, with larvae feeding on immobile prey including dead insects, bacteria, and protozoa.
Blephariceridae
Net-winged Midges
Blephariceridae, or net-winged midges, are a family of nematoceran flies distinguished by wings with a distinctive network of fissures and folds resembling cracked glass. Adults superficially resemble crane flies but differ in wing shape, head structure, and leg posture. Larvae are specialized inhabitants of torrential streams, possessing six ventral suckers that enable attachment to rocks in flows exceeding 2 m/s. The family contains over 200 species across dozens of genera worldwide, with highest diversity in mountainous regions. Larvae are ecologically significant as diatom grazers in lotic ecosystems.
Blethisa
Blethisa is a genus of ground beetles in the family Carabidae, subfamily Elaphrinae. It comprises approximately nine extant species distributed across the Holarctic region, plus one fossil species from the Pleistocene. Members of this genus are associated with aquatic or semi-aquatic habitats, with documented diving behavior in at least one species.
Brychius
crawling water beetles
Brychius is a genus of small aquatic beetles in the family Haliplidae, commonly known as crawling water beetles. The genus comprises five recognized species distributed across parts of Europe and North America. One species, Brychius hungerfordi, is federally endangered in the United States. Members of this genus inhabit freshwater environments and are associated with aquatic vegetation.
Brychius hornii
Horn's Crawling Water Beetle
Brychius hornii is a species of crawling water beetle in the family Haliplidae, first described by George Robert Crotch in 1873. It is native to western North America, where adults inhabit small streams and irrigation ditches, primarily on gravel substrates. The species belongs to a small genus of aquatic beetles adapted to life in running water environments.
Buenoa platycnemis
Buenoa platycnemis is a species of backswimmer in the family Notonectidae. It is a predatory aquatic insect found in freshwater habitats across the Americas. The genus Buenoa is distinguished by males producing audible stridulatory sounds, a rare trait among aquatic Heteroptera. This species has been recorded from North, Central, and South America, including multiple states in Brazil.
Carapacea
Carapacea is a suborder of mayflies (Ephemeroptera) characterized by nymphs possessing a distinctive carapace-like structure formed from expanded and fused thoracic tergites. This morphological adaptation distinguishes them from other mayfly suborders. The group includes families such as Baetiscidae and Prosopistomatidae. Carapacean mayflies are primarily found in running water habitats.
Celina
Celina is a genus of small predaceous diving beetles (family Dytiscidae) first described by Aubé in 1837. These aquatic beetles belong to the subfamily Hydroporinae and tribe Methlini. The genus occurs primarily in the Neotropical region, with records from northern South America including Colombia. Members of this genus are aquatic predators inhabiting freshwater environments.
Cenocorixa
water boatmen
Cenocorixa is a genus of water boatmen (family Corixidae) comprising approximately 12 described species. These aquatic insects are notable for their specialized swimming mechanics using synchronized hind leg rowing, and for their complex acoustic communication systems involving species- and sex-specific stridulatory signals. Both males and females produce sound by rubbing specialized pegs on the fore femora against the head. The genus exhibits significant ecological research interest regarding feeding niche partitioning in sympatric and allopatric populations, with some species inhabiting saline lakes.
Centrocorisa nigripennis
Centrocorisa nigripennis is a species of water boatman in the family Corixidae, first described by Fabricius in 1803. It belongs to the suborder Heteroptera (true bugs) within the order Hemiptera. The genus Centrocorisa is part of the tribe Corixini, a group of aquatic insects commonly known as water boatmen. As a member of the Nepomorpha, it is adapted to an aquatic lifestyle. The species has been recorded across the Caribbean, Middle America, and North America.
Chaoboridae
phantom midges, glassworms
Chaoboridae, known as phantom midges or glassworms, is a family of aquatic midges with cosmopolitan distribution. The family comprises approximately 55–90 species across six genera, including the widespread Chaoborus and the endemic Australian genera Promochlonyx and Australomochlonyx. Adults are delicate, chironomid-like flies distinguished by wing venation and plumose male antennae. Larvae are nearly transparent aquatic predators with modified raptorial antennae used to capture prey.
Chaoborus albatus
phantom midge
Chaoborus albatus is a species of phantom midge described by Johnson in 1921. As a member of the family Chaoboridae, it is an aquatic dipteran whose larvae are known for their transparent, nearly invisible bodies. The genus Chaoborus is characterized by larvae that lack respiratory siphons and possess unique gas-filled tracheal sacs that function as hydrostatic organs for vertical migration in water columns.
Chaoborus festivus
phantom midge
Chaoborus festivus is a species of phantom midge in the family Chaoboridae, first described by Dyar and Shannon in 1924. The genus Chaoborus comprises aquatic midges known for their transparent, nearly invisible larvae that inhabit freshwater systems. Larvae are predatory and possess unique gas-filled sacs that aid in buoyancy control. Adults are short-lived and do not feed.
Cirolanidae
Cirolanidae is a family of aquatic isopod crustaceans established by Dana in 1852. The family encompasses approximately 70 genera and over 500 species, ranging from small interstitial forms to the supergiant deep-sea isopods of the genus Bathynomus that can exceed 30 cm in length. Members occupy diverse aquatic habitats including marine coastal waters, deep-sea environments, freshwater systems, and subterranean groundwater. The family is characterized by dorsoventrally flattened bodies and includes both free-living and troglobitic species.
Cleptelmis addenda
Cleptelmis addenda is a species of riffle beetle in the family Elmidae, first described by Fall in 1907. Riffle beetles are aquatic insects adapted to life in fast-flowing streams and rivers. The species is known from North America, though specific details about its biology remain poorly documented.
Clinocerinae
aquatic dance flies
Clinocerinae is a subfamily of dance flies (Empididae) comprising 16 recognized genera including Clinocera, Wiedemannia, Dolichocephala, Trichoclinocera, and Kowarzia. Members are commonly known as aquatic dance flies due to their strong association with freshwater habitats. The subfamily has a global distribution with particularly diverse faunas in the Palaearctic and Oriental regions. Multiple generic revisions and species-level revisions have been published for Nearctic, Palaearctic, and Oriental faunas.
Colymbetes dolabratus
Colymbetes dolabratus is a predaceous diving beetle in the family Dytiscidae. It has a Holarctic distribution, occurring in both North America and the Palearctic region. The species belongs to the genus Colymbetes, a group of medium-sized diving beetles adapted to aquatic habitats.
Colymbetes longulus
Colymbetes longulus is a species of diving beetle in the family Dytiscidae, first described by LeConte in 1862. It belongs to the subfamily Colymbetinae, a group of predatory aquatic beetles. The genus Colymbetes is among the more diverse genera of diving beetles, with species distributed across the Northern Hemisphere. Like other members of its family, this species is adapted to life in freshwater habitats.
Colymbetes sculptilis
Sculptured Predaceous Diving Beetle
Colymbetes sculptilis is a species of predaceous diving beetle in the family Dytiscidae. It is known from North America, with records from Canada (Alberta, British Columbia) and the United States. As a member of the genus Colymbetes, it is adapted to aquatic habitats and exhibits the typical diving beetle morphology of streamlined body form and hind legs modified for swimming. The specific epithet 'sculptilis' refers to the sculptured or carved appearance of the body surface.
Copepoda
copepods
Copepods are small aquatic crustaceans and one of the most abundant and diverse multicellular organisms on Earth. They occupy nearly every aquatic habitat, from marine plankton to deep ocean floors, freshwater lakes, groundwater systems, and even moist terrestrial environments such as leaf litter and bromeliad phytotelmata. The group includes free-living forms as well as highly modified parasites. Copepods are fundamental components of aquatic food webs, serving as critical prey for fish, whales, and other marine life, while also contributing to nutrient cycling and carbon sequestration through the biological pump.
Coptotomus loticus
Coptotomus loticus is a predaceous diving beetle described by Hilsenhoff in 1980. It belongs to the subfamily Coptotominae within Dytiscidae, a family of aquatic beetles. The species is known from eastern Canada, with records from Ontario and Québec. As a member of Coptotomus, it represents one of the less commonly encountered genera in North American dytiscid fauna.
Corixidae
Water Boatmen, Lesser Water Boatmen
Corixidae is a family of aquatic true bugs in the order Hemiptera, commonly known as water boatmen or lesser water boatmen. The family comprises approximately 500 species across 33-55 genera distributed worldwide in freshwater habitats, with a few species inhabiting saline waters. Members are distinguished from similar aquatic bugs by their right-side-up swimming posture, flattened body shape, and modified front legs adapted for feeding. The family exhibits diverse feeding strategies including herbivory, carnivory, detritivory, and omnivory.
Corixinae
water boatman
Corixinae is a subfamily of aquatic true bugs comprising at least 130 described species. Members are commonly known as water boatmen and are found in freshwater habitats worldwide. The subfamily includes the genera Corixa and Sigara, among others. Larvae develop through five instars, with distinct morphometric and chaetotaxy characteristics separating developmental stages.
Crenitis paradigma
Crenitis paradigma is a species of water scavenger beetle in the family Hydrophilidae. It is a small aquatic beetle known from western North America. The species was described by Orchymont in 1942. Like other members of the genus Crenitis, it is associated with aquatic or semi-aquatic habitats.
Crinodessus
Crinodessus is a monotypic genus of predaceous diving beetles in the family Dytiscidae. The genus contains a single species, Crinodessus amyae, described by K.B. Miller in 1997. This genus belongs to the tribe Bidessini within the subfamily Hydroporinae.
Curicta
water scorpion
Curicta is a genus of aquatic insects in the family Nepidae, commonly known as water scorpions. Members of this genus are predatory true bugs found in freshwater habitats. The genus was established by Stål in 1861 and is placed in the tribe Curictini within the subfamily Nepinae. Curicta scorpio is one documented species, with records extending from Central to South America.
Cybister
Giant Diving Beetles
Cybister is a genus of large predaceous diving beetles (Dytiscidae) distributed across all continents except Antarctica. Adults range up to 43 mm in length, with some species among the largest diving beetles. The genus comprises approximately 98 species organized into four subgenera. Both adults and larvae are aquatic predators, with larvae of some species known to prey on vertebrates including tadpoles and small fish.
Cyrtobagous salviniae
salvinia weevil, Kariba Weed Biocontrol Weevil
Cyrtobagous salviniae is a subaquatic weevil native to Brazil, widely distributed as a classical biological control agent for invasive aquatic ferns of the genus Salvinia. It has proven effective in reducing infestations of Salvinia molesta and Salvinia minima in tropical and subtropical regions, with documented population reductions exceeding 90% in under one year in some cases. The species is subject to natural enemy pressure from pathogens such as Helicosporidium sp., which can reduce its reproductive output and efficacy.
Dicranopselaphus
water-penny beetles
Dicranopselaphus is a genus of water-penny beetles in the family Psephenidae, comprising over 40 described species. The genus exhibits a disjunct distribution spanning Asia and North America. Members are aquatic beetles with distinctive flattened, oval larval forms adapted to clinging to submerged surfaces in flowing water.
Dineutus angustus
whirligig beetle
Dineutus angustus is a species of whirligig beetle in the family Gyrinidae, first described by LeConte in 1878. The species is found in North America. As a member of the genus Dineutus, it is among the larger whirligig beetles, characterized by a hidden scutellum and rounded elytral apices. Like other gyrinids, it inhabits the surface of freshwater bodies where it forms aggregations called rafts.
Dineutus ciliatus
whirligig beetle
Dineutus ciliatus is a species of whirligig beetle in the family Gyrinidae. It is one of two genera of whirligig beetles found in Missouri, distinguished from the smaller Gyrinus by its larger size (~12 mm) and hidden scutellum. The species is found in North America and is distinguished from similar congeners primarily by its dark ventral coloration and dark legs, in contrast to the orange-legged D. emarginatus. Whirligig beetles are aquatic insects that live almost exclusively on the water surface, where they form aggregations called 'rafts' that provide anti-predator benefits through increased vigilance and chemical defense.
Dixidae
Meniscus Midges
Dixidae (meniscus midges) are a small family of nematocerous Diptera comprising fewer than 200 species. Adults are small, frail gnats that remain near aquatic habitats and do not feed. Larvae are aquatic filter-feeders that inhabit unpolluted freshwater margins, where they rest in a distinctive U-shaped posture at the water surface film. The family occurs on all continents except Antarctica and has been documented since the Jurassic period.
Dolichopus albiciliatus
Dolichopus albiciliatus is a species of long-legged fly in the family Dolichopodidae, first described by Loew in 1862. Members of the genus Dolichopus are characterized by elongated legs and often metallic coloration. As with other dolichopodids, adults are predatory on small insects.
Dolichopus setifer
Dolichopus setifer is a species of long-legged fly in the family Dolichopodidae, described by Loew in 1861. As a member of the genus Dolichopus, it shares traits with other species in this group, including elongated legs and predatory habits. Adults are typically found in moist habitats near water. The species has been documented in 46 iNaturalist observations, indicating it is encountered with moderate frequency by naturalists.
Dolomedes okefinokensis
Okefenokee Fishing Spider
Dolomedes okefinokensis is a large nursery web spider endemic to the southeastern United States, named for the Okefenokee Swamp. Females reach approximately 30 mm in body length, while males are substantially smaller at about 8 mm. The species is strongly associated with swampy aquatic habitats and is difficult to distinguish from the similar Dolomedes tenebrosus. As a fishing spider, it exhibits surface-dwelling predatory behaviors characteristic of the genus.
Dolomedes vittatus
Banded Fishing Spider
Dolomedes vittatus is a nursery web spider commonly known as the Banded Fishing Spider. It occurs throughout the United States. As a member of the genus Dolomedes, it is associated with aquatic and semi-aquatic habitats where it hunts prey at the water's surface. The species is among the more frequently observed fishing spiders in North America, with over 5,000 documented observations.
Donacia
Donacia is a large genus of aquatic leaf beetles in the subfamily Donaciinae. Adults are active, capable of sustained flight, and possess adaptations for both aerial and submerged existence. The genus is distinguished by a truncate elytral apex and a pubescent area above the procoxa. Larvae develop on submerged portions of aquatic plants, while adults inhabit surface parts of the same host vegetation. The genus contains approximately 113 described species arranged into 15 proposed subgenera, with major radiations in the Nearctic and Palaearctic regions.
Donacia distincta
Donacia distincta is an aquatic leaf beetle in the family Chrysomelidae, described by LeConte in 1851. The species is known from North America, with records from Canada including Alberta, British Columbia, and Labrador. As a member of the subfamily Donaciinae, it shares the characteristic aquatic or semi-aquatic lifestyle of its genus.
Donacia fulgens
Glittering reed beetle
Donacia fulgens, commonly known as the glittering reed beetle, is a species of leaf beetle in the subfamily Donaciinae. Members of this genus are aquatic or semi-aquatic beetles closely associated with emergent vegetation in freshwater habitats. The species was described by LeConte in 1851 and is known to occur in North America, with records from Canada including Manitoba, New Brunswick, and Nova Scotia.
Donacia hypoleuca
Pale pond lily leaf beetle
Donacia hypoleuca is an aquatic leaf beetle in the family Chrysomelidae, commonly known as the pale pond lily leaf beetle. It is found in North America, with records from regions including Québec, Canada. As a member of the genus Donacia, it is associated with aquatic or semi-aquatic habitats and is likely linked to pond lily (Nymphaea) or other aquatic vegetation as host plants.
Donacia liebecki
Donacia liebecki is an aquatic leaf beetle in the family Chrysomelidae, first described by Schaeffer in 1919. The species is known from North America, with records from Saskatchewan, Canada. As a member of the subfamily Donaciinae, it is adapted to aquatic or semi-aquatic habitats. Specific ecological details for this species remain poorly documented in the available literature.
Donacia magnifica
magnificent aquatic leaf beetle, Magnificent pondweed beetle
Donacia magnifica is an aquatic leaf beetle in the family Chrysomelidae, first described by LeConte in 1851. It is native to North America, with records from Canada and the northern United States. The species is closely associated with pondweeds (Potamogeton spp.), on whose leaves adults feed. Like other members of the genus Donacia, it is adapted to an aquatic lifestyle.
Donacia militaris
Well-armed pond lily beetle
Donacia militaris is an aquatic leaf beetle in the family Chrysomelidae. It is native to the eastern United States. Like other members of the genus Donacia, it is associated with aquatic vegetation. The common name "Well-armed pond lily beetle" reflects both its aquatic habitat and the species epithet.
Donacia parvidens
Donacia parvidens is an aquatic leaf beetle in the family Chrysomelidae. The species was described by C. Schaeffer in 1919. It is found in North America, with records from Ontario, Canada. As a member of the subfamily Donaciinae, it is adapted to life in association with aquatic vegetation.
Donacia pubescens
Pubescent sedge beetle
Donacia pubescens is an aquatic leaf beetle in the family Chrysomelidae, first described by LeConte in 1868. It is found across North America, with records from Canada (Alberta, British Columbia, Manitoba) and the United States. Like other members of the genus Donacia, it is associated with aquatic habitats and is likely a specialist on aquatic plants.
Donacia tuberculata
Tuberculate reed beetle
Donacia tuberculata is a species of aquatic leaf beetle in the subfamily Donaciinae. It occurs in North America and is associated with aquatic and semi-aquatic habitats. Like other members of its genus, it is linked to wetland vegetation and has aquatic larvae that develop on submerged plant roots.
Draeculacephala inscripta
Waterlettuce Leafhopper
Draeculacephala inscripta is a leafhopper species in the family Cicadellidae, first described by Van Duzee in 1915. It is commonly known as the Waterlettuce Leafhopper due to its association with aquatic and semi-aquatic host plants. The species has been recorded from multiple U.S. states including Arkansas, Connecticut, Florida, Georgia, and Hawaii, as well as being present in California where it has been studied as a host for egg parasitoids. Its eggs serve as hosts for several parasitoid wasp species in the families Mymaridae and Trichogrammatidae.
Dromogomphus
Spinylegs, Spiny-legged Clubtails
Dromogomphus is a genus of dragonflies in the family Gomphidae, commonly known as spinylegs. The common name refers to the prominent spines on their legs, which function in prey capture. The genus includes three North American species: southeastern spinyleg, black-shouldered spinyleg, and flag-tailed spinyleg. These dragonflies are medium to large-sized clubtails with distinctive morphological features.
Drosophila deflecta
Drosophila deflecta is a fruit fly species in the Drosophila quinaria species group, first described by Malloch & McAtee in 1924. The species is notable for its specialized larval ecology: larvae function as scavengers on Nuphar water lilies. Unlike the well-studied invasive relative Drosophila suzukii, D. deflecta has received limited research attention and is known from few observations.
Dryopidae
long-toed water beetles
Dryopidae is a family of aquatic beetles in the superfamily Byrrhoidea, commonly known as long-toed water beetles. Adults possess dense hydrofuge hairs that enable underwater respiration and extended claws that give the family its common name. Despite being aquatic, adults cannot swim and instead cling to floating detritus. The family is distinguished from similar riffle beetles (Elmidae) by antennae that are not clubbed. Most species have terrestrial larvae, though the genus Stygoparnus has fully aquatic larvae and adults.
Dryops
long-toed water beetles
Dryops is a genus of long-toed water beetles in the family Dryopidae, with at least 20 described species and a worldwide distribution. The genus is particularly diverse in the Palaearctic region, where 34 species have been recorded. Members of this genus are aquatic beetles associated with freshwater habitats. Species such as Dryops rufipes and Dryops renateae have been documented from Turkey and Tajikistan respectively.
Dubiraphia giulianii
Giuliani's Dubiraphian Riffle Beetle
Dubiraphia giulianii is a species of riffle beetle (family Elmidae) described by Van Dyke in 1949. Like other elmid beetles, it is adapted to life in flowing water environments. The species is known from North America, though detailed biological information remains limited.
Dubiraphia harleyi
Dubiraphia harleyi is a species of riffle beetle in the family Elmidae, described by Barr in 1984. Riffle beetles are aquatic beetles that inhabit flowing water environments. The species is known from North America.
Dytiscidae
predaceous diving beetles, diving beetles
Dytiscidae, the predaceous diving beetles, is a family of aquatic beetles containing over 4,000 described species. Adults range from 0.9 mm to 4.75 cm in length, with most species between 1–2.5 cm. They are characterized by enlarged, flattened hind legs adapted for swimming and a streamlined, oval body shape that reduces drag underwater. The family exhibits remarkable chemical ecology, producing unique defensive steroids found nowhere else in the animal kingdom. Larvae, known as "water tigers," are elongate predators with sharp, pincer-like mandibles that inject digestive enzymes into prey.
Dytiscus
predaceous diving beetles
Dytiscus is a Holarctic genus of large predaceous diving beetles comprising 26 species. Adults range 2.2–4.4 cm in length and are fully aquatic, breathing air collected under their elytra. Both adults and larvae are active predators. The genus has been observed to reduce mosquito larva populations in wetland habitats.
Dytiscus fasciventris
Understriped Diving Beetle
Dytiscus fasciventris, the Understriped Diving Beetle, is a predaceous diving beetle in the family Dytiscidae. The species is found in North America, with records from Canada (Alberta, British Columbia) and the United States. Like other members of the genus Dytiscus, it is an aquatic predator inhabiting freshwater environments.
Dytiscus harrisii
Harris's diving beetle
Dytiscus harrisii, commonly known as Harris's diving beetle, is a predaceous diving beetle in the family Dytiscidae. It is one of the larger diving beetle species in North America. As a member of the genus Dytiscus, it is an aquatic predator inhabiting freshwater systems.
Dytiscus hybridus
Hybrid Predaceous Diving Beetle
Dytiscus hybridus is a species of predaceous diving beetle in the family Dytiscidae. It is found in North America, with confirmed records from Alberta and Manitoba in Canada. As a member of the genus Dytiscus, it is an aquatic predator, though specific details of its biology remain poorly documented in published sources.
Dytiscus marginicollis
giant green water beetle
Dytiscus marginicollis, commonly known as the giant green water beetle, is a predaceous diving beetle in the family Dytiscidae. It is among the larger species in the genus Dytiscus and is distributed across North America. Like other members of its genus, it is an aquatic predator inhabiting freshwater systems.
Elmidae
Riffle beetles
Elmidae, commonly known as riffle beetles, is a family of aquatic beetles in the superfamily Byrrhoidea containing over 150 genera and 1,500 described species. Both adults and larvae are primarily aquatic, inhabiting fast-flowing shallow areas of streams and rivers such as riffles. Adults breathe underwater using a plastron—a microfilm of air held by microscopic hairs on their body surface that enables oxygen diffusion from the water. The family has significant value in freshwater biomonitoring due to their sensitivity to water pollution.
Elminae
riffle beetles
Elminae is a subfamily of riffle beetles (family Elmidae) comprising at least 120 genera and over 1,300 described species. Members are primarily aquatic, inhabiting running water habitats across all major biogeographic regions. The subfamily exhibits considerable morphological diversity, with some genera showing pronounced secondary sexual dimorphism and miniaturization traits.
Elmini
riffle beetles
Elmini is a tribe of riffle beetles within the family Elmidae, comprising over 90 genera and approximately 1,200 described species in North America. These beetles are strictly aquatic as adults and larvae, inhabiting flowing water systems. The tribe represents a major radiation within the Elmidae, with species diversity concentrated in freshwater habitats.
Enallagma carunculatum
Tule Bluet
Enallagma carunculatum, commonly known as the tule bluet, is a species of damselfly in the family Coenagrionidae. It is native to North America and belongs to a genus that has been intensively studied by odonatologists, including taxonomic revisions by Rosser Garrison in 1984. As a member of the narrow-winged damselflies, it shares characteristics typical of the genus Enallagma, including slender bodies and aquatic larval stages. The species name 'carunculatum' refers to a small fleshy protuberance, likely describing a distinctive morphological feature.
Estelloxus
Estelloxus is a genus of mites in the family Lebertiidae, order Trombidiformes. The genus was established by Habeeb in 1963 and belongs to the superfamily Lebertioidea. Members of this genus are predatory or parasitic mites associated with freshwater or semi-aquatic habitats. The genus is poorly known, with limited published information on species diversity and biology.
Eubriinae
Eubriinae is a cosmopolitan subfamily of water penny beetles (Psephenidae) comprising approximately 15 genera and 144 species, with greatest diversity in Asia. Adults are distinguished by a convex dorsal body form, truncate or emarginate anterior pronotum with exposed head, non-tapering maxillary palpus apex, and five abdominal ventrites. Larvae are aquatic and exhibit a metapneustic respiratory system with spiracles positioned near the bases of paratergites VIII. The subfamily exhibits nearly worldwide distribution except Antarctica and some oceanic islands including New Zealand.
Eugaurax
frit flies
Eugaurax is a genus of frit flies (Chloropidae) established by Malloch in 1913. The genus contains 11 New World species, with North American representatives including leaf miners of aquatic plants. Larvae of E. floridensis mine tissues of arrowheads (Sagittaria spp.), while E. hydrocotyles is a leaf and stem miner of floating marshpennywort (Hydrocotyle ranunculoides). A species complex centered on E. quadrilineatus occurs throughout the Americas but its larval biology remains largely unknown.
Eurypterida
sea scorpions, eurypterids
Eurypterida is an extinct order of aquatic chelicerate arthropods commonly known as sea scorpions. They ranged from the Cambrian through the Permian periods, with their main diversification occurring during the Paleozoic. The group includes both marine and freshwater species, with body sizes ranging from a few centimeters to over two meters in some species. Eurypterids are characterized by a segmented body with a prosoma and opisthosoma, paired swimming paddles, and a terminal telson spine. They represent one of the largest arthropod groups in the fossil record and are significant for understanding early chelicerate evolution.
Eylais
Eylais is a genus of water mites in the family Eylaidae. The genus has cosmopolitan distribution. Larvae are ectoparasites of diving beetles (family Dytiscidae), attaching primarily to the ventral surface of the hindwings beneath the elytra.
Globia oblonga
Oblong Sedge Borer Moth, oblong sedge borer
Globia oblonga is a noctuid moth commonly known as the oblong sedge borer. First described by Grote in 1882, this species was formerly placed in the genus Capsula before that name was abandoned due to a homonym conflict with a mollusk genus. The species is notable for its specialized larval biology: early instars are leaf miners on emergent aquatic plants, while later instars transition to stem-boring below the water line. Adults are medium-sized moths with wingspans of 35–50 mm.
Gomphurus hybridus
Cocoa Clubtail
A medium-sized clubtail dragonfly endemic to eastern North America. Adults are active in late spring through mid-summer, with peak emergence typically in June. The species is named for the cocoa-brown coloration of mature males. Larvae develop in streams and rivers, requiring clean, flowing water with sandy or gravelly substrates. Like other gomphids, it is a predator both as aquatic nymph and aerial adult.
Gonielmis
Gonielmis is a genus of riffle beetles (family Elmidae) described by Sanderson in 1954. These beetles are aquatic insects associated with flowing water habitats. The genus belongs to the tribe Elmini within the subfamily Elminae. Taxonomic status varies between sources, with some treating it as a synonym of Heterlimnius while others recognize it as valid.
Graptocorixa serrulata
Graptocorixa serrulata is a species of water boatman (family Corixidae) in the order Hemiptera. It is a small aquatic true bug distributed across parts of North America and Middle America. As a member of the Graptocorixini tribe, it inhabits freshwater environments where it likely occupies the benthic zone typical of corixid beetles. The species was described by Philip Reese Uhler in 1897.
Gyrinidae
Whirligig Beetles
Whirligig beetles (family Gyrinidae) are aquatic beetles that inhabit the surface film of freshwater habitats worldwide. The family comprises approximately 700 extant species in 15 genera. These beetles are instantly recognizable by their habit of swimming rapidly in circles on the water surface when alarmed, a behavior that gives them their common name. They possess divided compound eyes—upper portions adapted for vision above water and lower portions for underwater viewing—a unique adaptation among insects. Members of this family are active predators and scavengers that form conspicuous aggregations or "rafts" on the water surface, which serve defensive functions through enhanced predator detection and possible aposematic signaling.
Gyrinus
whirligig beetle
Gyrinus is a genus of small aquatic beetles in the family Gyrinidae, commonly known as whirligig beetles. Members are strictly aquatic and live almost exclusively on the water surface, where they form conspicuous aggregations called rafts. They are native to multiple biogeographic regions including the Palearctic, Nearctic, North Africa, Asia, and Australia. The genus is distinguished from the larger Dineutus by its smaller size, exposed scutellum, and more pointed elytral apices.
Gyrinus gehringi
whirligig beetle
Gyrinus gehringi is a species of whirligig beetle in the family Gyrinidae, described by Chamberlain in 1929. The species is known from North America, with records from Canada including New Brunswick, Newfoundland, and Nova Scotia. Whirligig beetles in this genus are small to medium-sized aquatic beetles that live on the water surface, where they form conspicuous aggregations and exhibit rapid, erratic swimming behavior.
Gyrinus gibber
whirligig beetle
Gyrinus gibber is a species of whirligig beetle (family Gyrinidae) described by LeConte in 1868. Like other members of its family, it inhabits the water surface of freshwater habitats. The species occurs in North America, with records from Canada (New Brunswick, Ontario, Québec) and the United States. Whirligig beetles are named for their characteristic rapid, circling swimming patterns on water surfaces.
Gyrinus maculiventris
whirligig beetle
Gyrinus maculiventris is a species of whirligig beetle in the family Gyrinidae. It inhabits freshwater surface habitats in North America. Like other members of its genus, it exhibits the characteristic rapid, erratic circling behavior on water surfaces that gives whirligig beetles their common name.
Gyrinus minutus
whirligig beetle
Gyrinus minutus is a small whirligig beetle with a Holarctic distribution spanning northern North America and Eurasia. It is distinguished from congeners by its weak swimming ability, solitary behavior among emergent vegetation, and near-absence of volatile defensive compounds. The species exhibits a shortened life cycle of approximately seven weeks from egg to adult and is capable of flight, allowing colonization of new water bodies throughout the summer.
Gyrinus pectoralis
whirligig beetle
Gyrinus pectoralis is a species of whirligig beetle in the family Gyrinidae, first described by LeConte in 1868. It is found across North America, with records from Canada (Alberta, British Columbia, Labrador) and presumably throughout much of the United States. Like other whirligig beetles, it inhabits the surface waters of freshwater systems.
Gyrinus picipes
whirligig beetle
Gyrinus picipes is a species of whirligig beetle in the family Gyrinidae. It is found in North America, with records from multiple Canadian provinces including Alberta, British Columbia, and Manitoba. Whirligig beetles in this genus are characterized by their distinctive habit of swimming in erratic, circling patterns on the water surface.
Gyrinus pleuralis
whirligig beetle
Gyrinus pleuralis is a species of whirligig beetle described by Fall in 1922. It is found in North America, with records from Canada (Alberta, British Columbia, Saskatchewan) and presumably the United States. As a member of the family Gyrinidae, it is an aquatic beetle that lives on the water surface, though specific details of its biology are poorly documented.
Gyrinus plicifer
whirligig beetle
Gyrinus plicifer is a species of whirligig beetle in the family Gyrinidae. It occurs in North America, with records from British Columbia, Canada. Like other members of the genus, it inhabits the surface of freshwater bodies where it forms aggregations. The species was described by LeConte in 1852.
Gyrinus wallisi
whirligig beetle
Gyrinus wallisi is a species of whirligig beetle described by Fall in 1922. Members of the genus Gyrinus are small, surface-dwelling aquatic beetles known for their rapid, circular swimming patterns. This species occurs in North America, with records from Canada including Alberta, British Columbia, and Labrador.
Gyrinus woodruffi
whirligig beetle
Gyrinus woodruffi is a species of whirligig beetle in the family Gyrinidae, described by Fall in 1922. It is found in North America. Like other members of the genus Gyrinus, it is a small, aquatic beetle that lives on the water surface in aggregations known as "rafts."
Haliplidae
Crawling Water Beetles, haliplids
Haliplidae is a family of small aquatic beetles comprising approximately 200 species in five genera, representing the sole extant member of the superfamily Haliploidea. These beetles are characterized by their distinctive swimming behavior using alternating leg movements, which renders them relatively clumsy swimmers compared to other aquatic beetles. Consequently, they predominantly move by crawling along substrates rather than swimming. The family exhibits a cosmopolitan distribution in freshwater habitats worldwide.
Haliplus borealis
Haliplus borealis is a small aquatic beetle in the family Haliplidae, commonly known as crawling water beetles. The species occurs in North America with confirmed records from Alberta and Manitoba in Canada. Like other members of its genus, it inhabits freshwater environments. The species was described by LeConte in 1850. Available information on this species is limited, with only five observations documented on iNaturalist.
Haliplus cylindricus
Haliplus cylindricus is a species of crawling water beetle in the family Haliplidae, described by Roberts in 1913. The species belongs to a genus of small aquatic beetles commonly found in freshwater habitats. Very few observations of this species have been documented, with only two records in iNaturalist as of the available data. Like other haliplids, it is presumed to inhabit shallow, vegetated waters where it feeds on algae and small aquatic organisms.
Haliplus gracilis
crawling water beetle
Haliplus gracilis is a species of crawling water beetle in the family Haliplidae, described by Roberts in 1913. The species belongs to a genus of small aquatic beetles commonly found in freshwater habitats. Like other haliplids, it is adapted to life in and around water, with morphology suited for crawling rather than swimming.
Haliplus longulus
crawling water beetle
Haliplus longulus is a species of crawling water beetle in the family Haliplidae. It occurs in North America with confirmed records from western Canada including Alberta and British Columbia. Like other members of Haliplidae, it inhabits aquatic environments. The species was described by LeConte in 1850. Observations remain sparse with only three documented records in iNaturalist.
Haliplus mimeticus
Warm Springs Crawling Water Beetle
Haliplus mimeticus is a small aquatic beetle in the family Haliplidae, commonly known as crawling water beetles. This species was described by Matheson in 1912 and is known from North and Middle America. It is associated with warm spring habitats, as suggested by its common name. The genus Haliplus comprises small beetles that inhabit aquatic environments, often found in vegetation at the margins of ponds, lakes, and slow-moving waters.
Haliplus parvulus
Haliplus parvulus is a species of crawling water beetle in the family Haliplidae. Members of this genus are small, semi-aquatic beetles associated with freshwater habitats. Very few observations of this species exist in public databases.
Helichus
Helichus is a genus of small beetles in the family Dryopidae, found worldwide except Australia and Antarctica. Adults measure 1–8 mm and inhabit aquatic or riparian environments. The genus is notable for having terrestrial larvae despite the aquatic lifestyle of adults, a trait that may be unique among water-associated insects.
Helichus lithophilus
Contrasted Dryopid
Helichus lithophilus is a species of long-toed water beetle in the family Dryopidae, found in North America. It is commonly known as the Contrasted Dryopid. The species was originally described as Elmis lithophila by Germar in 1824. It belongs to a family of beetles adapted to aquatic and riparian habitats.
Helichus triangularis
Helichus triangularis is a species of long-toed water beetle in the family Dryopidae, described by Musgrave in 1935. The genus Helichus comprises aquatic beetles commonly associated with stream and river habitats. Records indicate presence in North America and Middle America, though specific details about this species' biology remain sparse.
Helophoridae
water scavenger beetles, crawling water beetles
Helophoridae is a family of small aquatic beetles within the superfamily Hydrophiloidea, containing the single genus Helophorus. The family comprises approximately 191 species distributed primarily across the Holarctic region, with 150 species in the Palearctic and 41 in North America. A limited number of species occur in the Afrotropical region (six species), Central America, and the Indomalayan region (one species in northern India). Species inhabit diverse freshwater habitats including springs, ponds, temporary pools, and running waters, with many showing strong associations with specific hydrological regimes.
Hemerodromia
dance flies
Hemerodromia is a genus of dance flies (family Empididae) comprising at least 170 described species. Species are primarily associated with lotic freshwater habitats, particularly streams and rivers. Several species show obligate associations with calcareous water systems and tufa formations in limestone karst regions. The genus exhibits high diversity in tropical Southeast Asia, with significant undescribed species richness anticipated.
Hesperocorixa atopodonta
water boatman
Hesperocorixa atopodonta is a species of water boatman described by Hungerford in 1927. It belongs to the family Corixidae, a group of aquatic true bugs known for their oar-like hind legs adapted for swimming. The species is native to North America and is part of the genus Hesperocorixa, which includes several similar aquatic insect species.
Hesperocorixa kennicottii
Hesperocorixa kennicottii is a species of water boatman in the family Corixidae. It is a small aquatic true bug found in North American freshwater habitats. The species was described by Philip R. Uhler in 1897. Like other members of the genus Hesperocorixa, it is adapted to life in lentic and slow-moving waters.
Hesperocorixa laevigata
water boatman
Hesperocorixa laevigata is a species of water boatman (family Corixidae) described by Uhler in 1893. It is distributed across Central America and North America. As a member of the Nepomorpha, it is an aquatic true bug adapted to life in freshwater environments. The species is one of approximately 47 observed records on iNaturalist, indicating it is documented but not extensively studied.
Hesperocorixa minor
Hesperocorixa minor is a species of water boatman in the family Corixidae. It is a true bug in the order Hemiptera, characterized by aquatic habits typical of the Nepomorpha infraorder. The species was described by Abbott in 1913. Available records indicate a North American distribution, though detailed ecological and biological information remains limited.
Hesperocorixa vulgaris
water boatman
Hesperocorixa vulgaris is a species of water boatman in the family Corixidae, first described by Hungerford in 1925. This aquatic true bug is found in North America and is most active during spring through autumn. It feeds primarily on algae, which it sometimes stirs up from waterbeds. When threatened, it can deliver a painful bite.
Heterelmis
riffle beetles
Heterelmis is a genus of aquatic beetles in the family Elmidae, commonly known as riffle beetles. The genus includes several described species, with Heterelmis comalensis being particularly notable as an endangered species endemic to spring systems in Texas. Members of this genus are fully aquatic, inhabiting oxygen-rich flowing water environments where they feed on microbial biofilms. Some species have reduced or non-functional wings, limiting their dispersal ability and making them vulnerable to habitat degradation.
Heterelmis comalensis
Comal Springs riffle beetle
Heterelmis comalensis is a federally endangered aquatic beetle endemic to two spring systems in Texas. Described in 1988, this flightless species measures approximately 2 mm and inhabits shallow gravel substrates in spring riffles fed by the Edwards Aquifer. The species has been the focus of intensive conservation research, including captive breeding programs and studies on sexual dimorphism identification.
Huleechius marroni
Huleechius marroni is a species of riffle beetle (family Elmidae) described by Brown in 1981. It belongs to the genus Huleechius, a group of aquatic beetles adapted to flowing water habitats. The species is known from Middle America and North America. Like other elmids, it is presumed to inhabit freshwater streams and rivers, though specific ecological details remain poorly documented.
Hydaticus
Hydaticus is a genus of predatory diving beetles in the family Dytiscidae, comprising approximately 150 described species and 12 subspecies arranged in two subgenera: Hydaticus and Prodaticus. The genus has a cosmopolitan distribution spanning the Palearctic, Afrotropical, Nearctic, and Oriental regions. Members are aquatic predators found in freshwater habitats including streams, pools, and lentic environments.
Hydaticus rimosus
Hydaticus rimosus is a diving beetle species in the family Dytiscidae, first described by Aubé in 1838. The species belongs to the genus Hydaticus, a group of predatory aquatic beetles commonly found in freshwater habitats. It occurs in the Caribbean and Middle America based on documented distribution records.
Hydrachnoidea
water mites
Hydrachnoidea is a superfamily of aquatic mites commonly known as water mites. Members are exclusively freshwater dwellers, representing one of the most diverse radiations of arachnids into aquatic environments. The group includes several families with distinct morphological adaptations for underwater life.
Hydrobiomorpha casta
Hydrobiomorpha casta is a water scavenger beetle in the family Hydrophilidae. The species is distributed across the Caribbean, Central America, and southeastern North America. It is one of approximately 125 species in the genus Hydrobiomorpha, a group of aquatic beetles adapted to freshwater habitats.
Hydrochara
water scavenger beetles
Hydrochara is a genus of water scavenger beetles (Hydrophilidae) containing 23 species distributed across North America, Europe, Asia, and Africa. The genus was revised taxonomically in 1980, with 21 species recognized at that time and multiple synonymies established. Species in this genus are aquatic and have been studied for their potential role in biological control of mosquito larvae. At least one species, Hydrochara rickseckeri, is endemic to California.
Hydrochara rickseckeri
Ricksecker's Water Scavenger Beetle
Hydrochara rickseckeri is a rare, California-endemic water scavenger beetle in the family Hydrophilidae. It is one of the few Hydrochara species native to North America. The species was described by Horn in 1895 and remains poorly known due to its limited distribution and scarcity of observations.
Hydrochus granulatus
Hydrochus granulatus is a species of water scavenger beetle in the family Hydrochidae. The species was described by Blatchley in 1910 and is known from North America, with records from both the United States and Canada. It is sometimes treated as a member of the family Hydrophilidae in alternative classifications.
Hydrometra australis
water measurer
Hydrometra australis is a species of water measurer bug in the family Hydrometridae. It inhabits aquatic environments across the Caribbean, Central America, and North America. The species has been documented as a host for parasitic water mites, which can affect its population dynamics.
Hydrophorus chrysologus
Hydrophorus chrysologus is a species of long-legged fly in the family Dolichopodidae, subfamily Hydrophorinae. It belongs to a genus specialized for life on water surfaces, where adults skate across puddles, intermittent streams, and lake margins using surface tension. Like other dolichopodids, it is predatory on small insects and other arthropods. The species was described by Walker in 1849.
Hydroporus lapponum
Hydroporus lapponum is a predaceous diving beetle in the family Dytiscidae. The species has a broad Holarctic distribution, occurring across North America and the Palearctic region. It belongs to the genus Hydroporus, a large and diverse group of small diving beetles. The species was first described by Gyllenhal in 1808.
Hydrozetidae
Hydrozetidae is a family of aquatic oribatid mites (water mites) within the order Oribatida, established by Grandjean in 1954. The family contains at least one genus, Hydrozetes, with at least 20 described species. These mites are distinguished by adaptations to freshwater habitats, including a reduced bothridium lacking a sensillus, monodactylous legs, and the absence of pteromorphae. Fossil evidence from the Late Paleocene of Canada demonstrates the family's ancient aquatic association with submerged vegetation.
Hydryphantoidea
Hydryphantoidea is a superfamily of prostigmatan mites within Trombidiformes, comprising approximately 250 described species across six families. Members are predominantly aquatic or semiaquatic, inhabiting freshwater habitats worldwide. The group includes the families Hydryphantidae, Hydrodromidae, Rhynchohydracaridae, Thermacaridae, Ctenothyadidae, and Teratothyadidae.
Hygrobatidae
water mites
Hygrobatidae is a family of prostigmatan mites in the order Trombidiformes, commonly known as water mites. The family comprises approximately 78 genera and over 100 described species, though estimates vary. Members are primarily aquatic, inhabiting springs, streams, and running waters across multiple continents. The genus Hygrobates includes a well-studied species complex (fluviatilis-complex) with ten or more morphologically similar species. Some species exhibit parasitic associations with amphibians.
Lemnaphila
duckweed miner flies
Lemnaphila is a genus of shore flies in the family Ephydridae, commonly known as duckweed miner flies. The genus was established by Cresson in 1933 and contains seven described species distributed in the Neotropical region. At least one species, L. neotropica, is known to mine duckweeds (Lemna species), creating distinctive feeding damage on these aquatic plants. The genus has attracted attention due to the economic and ecological significance of its association with duckweed communities.
Lethocerinae
giant water bugs
Lethocerinae is a subfamily of giant water bugs within Belostomatidae, comprising large aquatic predatory insects distributed across the Palearctic, Oriental, and Australian regions. The subfamily includes two genera: Lethocerus and Kirkaldyia. Members are characterized by their large size—among the largest in Hemiptera—and are distinguished from other belostomatid subfamilies by morphological and genetic differences. Species within this group face conservation concerns in parts of their range due to harvesting for human consumption.
Lethocerus americanus
American giant water bug, electric light bug, toe biter, fish killer
Lethocerus americanus is a large aquatic true bug native to southern Canada and the northern United States. Adults measure 5–6 cm in length, with females averaging larger than males. The species is a generalist predator in freshwater habitats and is known for its powerful bite, which it uses to subdue prey and defend itself. It is attracted to electric lights at night, earning it the common name 'electric light bug.'
Lethocerus medius
giant water bug
Lethocerus medius is a giant water bug in the family Belostomatidae, distinguished by its substantial size and unique parental care behavior. Males reach 49–58 mm, while females are larger at 62–63 mm. Unlike related species in the subfamily Belostomatinae that brood eggs on their backs, males of this species exhibit postzygotic parental care by guarding eggs deposited on objects above the water surface, moistening them, shading them, and protecting them from predators. This species occurs across a broad range from the southern United States through Mexico to Central America and throughout the Caribbean.
Libellula jesseana
Purple Skimmer
Libellula jesseana, the purple skimmer, is a threatened dragonfly endemic to Florida, United States. Adults measure approximately 5 centimeters in length. The species is restricted to infertile, clear, sandy-bottomed lakes with specific shoreline vegetation, and is critically imperiled due to habitat degradation from human development.
Limnochares americana
Red Freshwater Mite
Limnochares americana is a species of freshwater mite in the family Limnocharidae, first described by Lundblad in 1941. It belongs to the order Trombidiformes, a group that includes many aquatic and terrestrial mites. As a member of the infraorder Hydrachnidia (water mites), this species is adapted to life in freshwater habitats. The common name "Red Freshwater Mite" reflects its aquatic ecology and likely coloration.
Limnohydrobius melaenus
Limnohydrobius melaenus is a species of water scavenger beetle in the family Hydrophilidae. It is found in eastern North America, ranging from the Canadian Maritime provinces west to the upper Midwest and south to the southeastern United States. The species is associated with aquatic and semi-aquatic habitats. Like other members of the genus, it is presumed to have aquatic larvae and terrestrial adults.
Liodessus
Liodessus is a genus of small diving beetles in the family Dytiscidae, tribe Bidessini. Species occur across the Americas from North America through the Andes to southern South America, with notable diversity in high-altitude wetlands and páramo ecosystems. Many species inhabit shallow, exposed pools and temporary water bodies, showing phenotypic plasticity in body form correlated with habitat permanence. The genus contains numerous species, with several new species described from the high Andes of Peru and Colombia in recent years. Taxonomic resolution relies heavily on male genital morphology.
diving-beetleaquatic-insecthigh-altitudepáramoAndesphenotypic-plasticityBidessiniHydroporinaeDytiscidaeColeopterawetlandtemporary-poolpeatlandsubantarctictaxonomymale-genitaliaCOI-barcodecryptic-speciesincipient-speciationhybridizationgeometric-morphometricsenvironmental-sentineldrought-indicatorSouth-AmericaNorth-AmericaFijiNavarino-IslandMagallanesPeruColombiaChileArgentinaBoliviaBogotáJunínCuscoHuánucoCundinamarcaSumapazPunasteppepeat-bogshallow-poolpuddlealtitude-3400-4900mL.-affinis-complexL.-bogotensis-complexL.-chilensisL.-affinisL.-obscurellusL.-noviaffinisL.-saratogaeL.-picinusL.-santarositaL.-alpinusL.-hauthiL.-rhigosL.-thespesiosGuignot-1939ZimmermannSharpLeConteSaySteinheilRégimbartBiströmMillerBalkeFranciscoloSanfilippoPederzaniNilssonYoungClarkHatchMannerheimCaseyAubéBalfour-BrowneBrinckPeschetSolierWhiteZootaxaZooKeysInsect-Systematics-&-EvolutionAnales-del-Instituto-de-la-PatagoniaPLoS-ONEGBIFiNaturalistCatalogue-of-LifeNCBI-TaxonomyBOLDBarcode-of-Life-Data-SystemCOImitochondrial-DNAphylogeographypopulation-structurerange-expansionlineagegenetic-structuresubspecieslectotypeneotypesynonymynew-speciesnew-subspeciesrevisionmorphometricsallometrybody-sizebody-shapeelytrapronotumstriaecolorationpunctationhabitusoccipital-linegenital-morphologyaedeagusparamerediagnosisidentification-keytype-localitytype-specimenetymologybionomicsnatural-historygeographic-distributiondistribution-mapNearcticNeotropicalPatagoniaCape-Horn-Biosphere-Reservedroughtclimate-changeenvironmental-changesentinel-speciesplasticityadaptationphenotypic-variationhydrological-landscapewater-permanencepoolbogAltiplanoAndeanhigh-elevationmontanealpineaquaticfreshwaterlenticloticinsectbeetleAdephagaLiodessusLiodessus affinis
Liodessus affinis is a small predaceous diving beetle in the family Dytiscidae, first described by Say in 1823. The species belongs to a taxonomically challenging species complex that was revised in 1998, during which several former synonyms were elevated to species status. It is found in aquatic habitats across North America, with records from Canada and the United States. Like other members of the genus, it is a predator in freshwater ecosystems.
Listronotus anthracinus
underwater weevil
Listronotus anthracinus is a species of underwater weevil in the beetle family Curculionidae. It is distinguished from congeners by its aquatic habits. The species was originally described under the genus Macrops. Very little is known about its biology compared to better-studied relatives such as L. oregonensis (carrot weevil) and L. maculicollis (annual bluegrass weevil).
Listronotus appendiculatus
underwater weevil
Listronotus appendiculatus is an underwater weevil in the family Curculionidae. It is found in North America, with distribution records from Canada including Alberta, British Columbia, Manitoba, and New Brunswick. The species inhabits aquatic environments, distinguishing it from many congeneric species that are agricultural pests of terrestrial crops.
Listronotus caudatus
Listronotus caudatus is a species of underwater weevil in the beetle family Curculionidae. It is found in North America. The species has been recorded in multiple Canadian provinces including British Columbia, New Brunswick, Ontario, and Québec. Very little is known about its specific biology or ecology compared to better-studied congeners such as the carrot weevil (L. oregonensis) and annual bluegrass weevil (L. maculicollis).
Listronotus delumbis
underwater weevil
Listronotus delumbis is a species of underwater weevil in the beetle family Curculionidae. It is one of relatively few weevil species adapted to aquatic or semi-aquatic habitats. The species was described by Gyllenhal in 1834 and has been historically classified under the genus Listroderes as a synonym. Knowledge of its biology and ecology remains limited compared to better-studied Listronotus species.
Listronotus dietzi
Listronotus dietzi is a species of underwater weevil in the beetle family Curculionidae. It was described by O'Brien in 1979. The species is known from a very limited geographic range, with records restricted to Louisiana in the United States. Very little is known about its biology or ecology.
Listronotus frontalis
underwater weevil
Listronotus frontalis is a species of underwater weevil in the beetle family Curculionidae. It is found in North America, with records from Canada (Alberta, British Columbia, Ontario, Québec). The species was described by J.L. LeConte in 1876. Like other members of the genus Listronotus, it is associated with aquatic or semi-aquatic habitats.
Listronotus lodingi
Listronotus lodingi is a species of underwater weevil in the beetle family Curculionidae. It was originally described as Hyperodes lodingi by Blatchley in 1920. The species is currently recognized as a synonym of Hyperodes lodingi in some taxonomic databases, though it is listed under the genus Listronotus in other sources. Very little specific information is available about its biology or ecology.
Listronotus marshalli
Listronotus marshalli is a species of underwater weevil in the beetle family Curculionidae, described by O'Brien in 1981. It is found in North America and belongs to a genus that includes several aquatic or semi-aquatic species. The species is provisionally accepted in taxonomic databases.
Listronotus pallidus
Listronotus pallidus is an underwater weevil in the family Curculionidae, described by O'Brien in 1981. It is distinguished from other weevils by its aquatic habitat. The species is known from North America, though specific details about its biology and ecology remain poorly documented. Like other members of the genus Listronotus, it likely inhabits submerged vegetation in freshwater environments.
Listronotus porcellus
underwater weevil
Listronotus porcellus is an aquatic weevil in the family Curculionidae, known for its underwater lifestyle. It is one of several species in the genus Listronotus, which includes both aquatic and terrestrial weevils. The species has been documented in North America, including British Columbia and Québec in Canada. Relatively little is known about its specific biology compared to better-studied congeners such as L. maculicollis and L. oregonensis.
Listronotus squamiger
Listronotus squamiger is a species of underwater weevil in the family Curculionidae. It has been documented in aquatic habitats across North America, including the Canadian provinces of Alberta, British Columbia, Manitoba, and New Brunswick. The species belongs to a genus that includes several economically significant agricultural pests, though specific information about the biology and ecology of L. squamiger itself remains limited.
Listronotus turbatus
Listronotus turbatus is a species of underwater weevil in the family Curculionidae. It was described by O'Brien in 1981. The species is found in North America. Unlike several congeners, it is not known to be an agricultural pest.
Maccaffertium mediopunctatum mediopunctatum
Maccaffertium mediopunctatum mediopunctatum is a subspecies of mayfly in the family Heptageniidae. The genus Maccaffertium was established to accommodate species formerly placed in Heptagenia and related genera, distinguished by particular features of the male genitalia and wing venation. This subspecies is part of a group of flatheaded mayflies commonly found in North American streams and rivers. Subimagos (subadults) of this species exhibit the characteristic pale, milky-white coloration and opaque wings typical of the genus, with sexual dimorphism in eye size and abdominal claspers.
Maccaffertium terminatum terminatum
Maccaffertium terminatum terminatum is a subspecies of mayfly in the family Heptageniidae. Like other members of its genus, it exhibits the unique mayfly developmental trait of having a winged sub-adult stage (subimago) that requires one final molt to reach full adulthood. The subspecies is part of the Nearctic fauna, with records from North America. As an aquatic insect, its life cycle is tied to freshwater habitats, with nymphs developing in water before emerging as winged adults.
Macronychini
riffle beetles
Macronychini is a tribe of riffle beetles within the family Elmidae, first described by Gistel in 1848. The tribe comprises more than 20 genera and over 80 described species. Members of this tribe are aquatic beetles associated with running water habitats. The tribe is taxonomically well-established but individual species continue to be described, particularly from underexplored regions such as China.
Matus bicarinatus
Matus bicarinatus is a predaceous diving beetle in the family Dytiscidae. It belongs to the subfamily Matinae, a group of specialized diving beetles. The species is documented from North America, with records from Ontario, Canada. Knowledge of its biology remains limited due to few observations.
Megamelus flavus
Megamelus flavus is a species of delphacid planthopper in the family Delphacidae, described by Crawford in 1914. It belongs to the genus Megamelus, a group of planthoppers associated with aquatic and semi-aquatic habitats. Distribution records indicate presence in northern North America and parts of Asia.
Microvelia
small water striders, riffle bugs
Microvelia is a large genus of small semiaquatic bugs comprising at least 230 described species distributed worldwide. These insects inhabit nearshore areas of stagnant or slow-flowing freshwater, where they move on the water surface using surface tension. The genus exhibits wing dimorphism, with populations containing both fully winged (macropterous) and short-winged (brachypterous) individuals. Several subgenera have been recognized including Austromicrovelia, Barbivelia, Pacificovelia, and Picaultia, based on phylogenetic and morphological analyses.
Microvelia californiensis
Microvelia californiensis is a species of small water strider in the family Veliidae, described by McKinstry in 1937. The species is known from California and adjacent regions of western North America. Like other members of the genus, it inhabits the surface film of freshwater habitats. Very few observations of this species have been documented in public databases.
Myxophaga
Minute Bog and Skiff Beetles
Myxophaga is a small suborder of Coleoptera comprising approximately 65 described species across four extant families: Lepiceridae, Hydroscaphidae, Sphaeriusidae, and Torridincolidae. These beetles are among the smallest in the order, ranging from small to minute in size. All members are aquatic or semiaquatic, with many species inhabiting hygropetric environments—thin water films on rock surfaces in running water. The suborder is characterized by distinctive morphological features including clubbed antennae with fewer than nine segments, open mesocoxal cavities, and apically rolled hind wings in resting position.
Neargyractis
Neargyractis is a genus of aquatic moths in the family Crambidae, subfamily Acentropinae. The genus was established by W.H. Lange in 1956 during a revision of North American aquatic moths. It contains at least eight described species distributed primarily in the Americas.
Nectoporus
Nectoporus is a genus of diving beetles (family Dytiscidae) established by Guignot in 1950. It belongs to the subfamily Hydroporinae, a diverse group of small to medium-sized predatory aquatic beetles. Members of this genus are found in freshwater habitats, primarily in the Southern Hemisphere based on available distribution records. As with other dytiscids, they are aquatic predators with adaptations for underwater life including modified hind legs for swimming.
Neoelmis caesa
riffle beetle
Neoelmis caesa is a species of riffle beetle in the family Elmidae. It is found in North America. As a member of the Elmidae, it inhabits aquatic environments, specifically riffle habitats in streams and rivers. The species was first described by LeConte in 1874.
Neohermes
Gray Fishflies
Neohermes is a genus of fishflies in the family Corydalidae containing approximately five described species. Species in this genus are aquatic insects associated with freshwater habitats, particularly intermittent streams. The genus is known for including long-lived aquatic species with specialized burrowing behaviors.
Neophylax fuscus
Neophylax fuscus is a species of caddisfly in the family Uenoidae (formerly placed in Thremmatidae). It occurs in North America. Like other members of the genus Neophylax, the larvae are case-building caddisflies that construct protective cases using silk and materials gathered from their aquatic environment.
Neoplea
pygmy backswimmers
Neoplea is a genus of minute aquatic predatory true bugs in the family Pleidae, commonly known as pygmy backswimmers. The genus contains at least 25 described species distributed across North and South America, with numerous undescribed species. Members are small-bodied (approximately 1.5 mm), active predators in lentic freshwater habitats. The best-studied species, Neoplea striola, has been investigated for its potential role in trophic cascades in fishless subtropical ponds.
Neoporus vitiosus
Neoporus vitiosus is a small predaceous diving beetle in the family Dytiscidae, subfamily Hydroporinae. The species was described by LeConte in 1855 and is documented from North America, with specific records from Ontario and Québec in Canada. As a member of the Hydroporinae, it likely inhabits freshwater environments, though detailed ecological studies are limited.
Nepidae
waterscorpions, water scorpions, water stick insects, needle bugs
Nepidae is a family of exclusively aquatic predatory true bugs containing approximately 250 species in 14 genera. Members are commonly called waterscorpions due to their superficial resemblance to scorpions, possessing raptorial forelegs and a long caudal respiratory siphon. The family is divided into two subfamilies: Nepinae (broad, flat-bodied 'water scorpions') and Ranatrinae (slender 'water stick insects'). They occur on all continents except Antarctica, inhabiting stagnant or slow-moving freshwater habitats.
Nigronia serricornis
Serrate Dark Fishfly, Saw-combed Fishfly, Hellgrammite
Nigronia serricornis is a large aquatic insect in the family Corydalidae, commonly known as the serrate dark fishfly or saw-combed fishfly. The aquatic larvae, often called hellgrammites, are sit-and-wait ambush predators and frequently the largest insect predators in small woodland streams. Adults are short-lived, weak fliers that do not disperse far from their emergence sites. The species has a broad distribution across eastern North America with documented genetic structure shaped by the Appalachian Mountains.
Notomicrus
Notomicrus is a genus of small water beetles in the family Noteridae, first described by Sharp in 1882. Species occur in freshwater habitats across the New World, from North America through Central America to South America and the Caribbean. The genus includes at least 13 described species, with several recent descriptions from Guadeloupe and Suriname. Some species exhibit unusual reproductive traits, including suspected obligate parthenogenesis.
Notonecta
backswimmers, common backswimmer, Milky Backswimmers
Notonecta is a genus of aquatic predatory insects commonly known as backswimmers, comprising approximately 70 species. They are distinguished by their habit of swimming upside down, using long hind legs modified as oars. The genus exhibits distinct morphological and behavioral traits that separate it from superficially similar water boatmen (family Corixidae), including triangular cross-section body shape, bold color patterns, and diurnal activity patterns. Notonecta species occupy diverse freshwater habitats and function as active predators of small aquatic organisms.
Notonecta borealis
backswimmer
Notonecta borealis is a species of backswimmer (family Notonectidae) found in North America. As a member of the genus Notonecta, it is an aquatic predatory insect that swims upside down. The species was described by Hussey in 1919. Very little specific biological information has been published for this particular species.
Notonecta hoffmani
Notonecta hoffmani is an aquatic predatory bug in the family Notonectidae, commonly known as backswimmers. The species has been studied for its functional response to prey availability, showing measurable preferences among alternative prey types. Like other Notonecta species, it swims inverted using its keel-shaped dorsum and modified hind legs. Research indicates its predatory efficiency on mosquito larvae is influenced by the presence of competing prey sources.
Notonecta insulata
backswimmer
Notonecta insulata is a species of backswimmer, an aquatic true bug in the family Notonectidae. It is found in North America, where it inhabits freshwater environments. Like other backswimmers, it swims upside down and is an active predator of small aquatic invertebrates. The species was described by Kirby in 1837.
Notonecta irrorata
backswimmer
Notonecta irrorata is a North American backswimmer species first described in 1879. It is a predatory aquatic insect known for its distinctive upside-down swimming posture. Research has demonstrated that its presence in ponds repels oviposition by Culex mosquitoes, and that its colonization patterns show a hump-shaped relationship with habitat isolation.
Notonecta kirbyi
Kirby's Backswimmer
Notonecta kirbyi, commonly known as Kirby's backswimmer, is a species of aquatic true bug in the family Notonectidae. Like other backswimmers, it swims inverted on the water surface using its long, oar-like hind legs. The species is found in North America and shares the general habits of the genus Notonecta, including predatory feeding on small aquatic organisms and the ability to fly between water bodies.
Notonecta montezuma
Notonecta montezuma is a species of backswimmer in the family Notonectidae. It is an aquatic true bug found in North and Middle America. Like other backswimmers, it swims upside down using its long, oar-like hind legs. The species is part of the genus Notonecta, which contains backswimmers that are generally larger than water boatmen and more commonly encountered in open water habitats.
Notonecta petrunkevitchi
Notonecta petrunkevitchi is a species of backswimmer in the family Notonectidae, described by Hutchinson in 1945. As a member of the genus Notonecta, it shares the characteristic habit of swimming upside down in aquatic environments. The species is known from North America, though specific details about its ecology and distribution within this range remain poorly documented. Like other backswimmers, it is presumably predatory and possesses hydrophobic hairs for underwater respiration.
Notonecta unifasciata andersoni
Notonecta unifasciata andersoni is a subspecies of backswimmer (family Notonectidae) described by Hungerford in 1933. It belongs to the genus Notonecta, a group of aquatic true bugs known for their upside-down swimming posture. The subspecies is part of the widespread N. unifasciata species complex, which occurs across the Americas. As with other notonectids, it is presumed to be an aquatic predator, though specific biological details for this subspecies remain poorly documented in available literature.
Nymphomyia
Nymphomyia is a genus of minute, delicate flies in the family Nymphomyiidae. Adults are characterized by reduced wing venation and reduced mouthparts. Larvae are aquatic and inhabit fast-flowing streams.
Nymphulini
Nymphulini is a tribe of small moths in the subfamily Acentropinae (Crambidae), characterized by aquatic or semi-aquatic larval habits. The group includes numerous species whose larvae feed on submerged or emergent aquatic plants, making them among the few truly aquatic Lepidoptera. Adults are generally small with narrow wings and reduced mouthparts. The tribe is most diverse in tropical and subtropical regions, with some species extending into temperate zones.
Ochthebius lineatus
Ochthebius lineatus is a species of minute moss beetle in the family Hydraenidae. The species was described by J.L. LeConte in 1852. It has a broad distribution across the Americas, with records from North America, Central America, and South America. As a member of the Hydraenidae, it belongs to a family of small aquatic or semiaquatic beetles commonly known as minute moss beetles or water scavenger beetles.
Ochthera
Mantid Shoreflies
Ochthera is a genus of predatory shore flies (Ephydridae) distinguished by raptorial forelegs modified for seizing prey. Adults and larvae are exclusively predatory, feeding on midge and mosquito larvae. The genus has a global distribution with approximately 37-55 species, including 13 species in North America. Some species have documented significance as predators of disease vectors, including African malaria vectors.
Ochthera anatolikos
Ochthera anatolikos is a small predatory shore fly in the family Ephydridae. Adults and larvae are both predatory, with raptorial forelegs adapted for seizing prey. The species occurs in shallow aquatic habitats across North America, where it feeds on small insects and their larvae. It was described by Clausen in 1977.
Ochthera lauta
Ochthera lauta is a species of shore fly in the family Ephydridae, characterized by raptorial forelegs adapted for capturing prey. The species belongs to a genus known for predatory behavior on small insects near aquatic habitats. It has been documented in the United States, West Indies, and Central America.
Oligostigmoides
A genus of aquatic moths in the family Crambidae, described by William Harry Lange in 1956. The genus contains five described species distributed from the southwestern United States through Central America to Peru. As members of the subfamily Acentropinae, these moths are associated with freshwater habitats during their larval stages.
Optioservus canus
pinnacles optioservus riffle beetle
Optioservus canus is a species of riffle beetle in the family Elmidae, first described by Chandler in 1954. Riffle beetles are aquatic insects that inhabit running water environments. The species is known from North America.
Optioservus trivittatus
Optioservus trivittatus is a species of riffle beetle in the family Elmidae. It is found in North America, with records from Canada including New Brunswick, Newfoundland, Nova Scotia, and Ontario. Riffle beetles in this family are aquatic insects typically associated with running water habitats. The species was described by Brown in 1930 and has been subject to taxonomic revision based on male genitalia, elytral patterns, and geographic distribution.
Ostracoda
Ostracods, Seed Shrimp
Ostracoda are small bivalved crustaceans, typically 0.5-2 mm in length, characterized by a hinged carapace that completely encloses the body. They exhibit remarkable diversity with over 33,000 described species and an estimated 70,000 total species when fossil forms are included. The group occupies virtually all aquatic environments from deep ocean trenches to temporary freshwater pools, with some species adapted to moist terrestrial microhabitats. Their calcified carapaces provide excellent fossil preservation, making them valuable for paleoecological and biostratigraphic studies.
Oulimnius nitidulus
riffle beetle
Oulimnius nitidulus is a species of riffle beetle in the family Elmidae, described by LeConte in 1866. It is distributed across North America. Riffle beetles in this genus are aquatic insects adapted to flowing water environments. The species was originally described under the basionym Elmis nitidula.
Parapoynx allionealis
Watermilfoil Leafcutter Moth
Parapoynx allionealis, commonly known as the watermilfoil leafcutter moth, is a crambid moth native to North America. The species is notable for its aquatic larval stage and its specialized feeding on submerged and floating aquatic plants. It has been documented across a broad geographic range spanning eastern and central North America, from southern Canada to the Gulf Coast. The common name reflects its association with Myriophyllum (watermilfoil) species, though larvae feed on multiple aquatic plant genera.
Parapoynx badiusalis
Chestnut-marked Pondweed Moth
Parapoynx badiusalis, the chestnut-marked pondweed moth, is a small crambid moth found in freshwater wetland habitats across north-central and northeastern North America. The species was described by Francis Walker in 1859. Adults are active from late spring through early autumn. Larvae are aquatic and feed on pondweed species.
Parapoynx maculalis
polymorphic pondweed moth
Parapoynx maculalis, commonly known as the polymorphic pondweed moth, is a small aquatic moth in the family Crambidae. It inhabits freshwater environments across eastern North America, from Florida and Texas north to Canada. The species is distinguished by its small size, white forewings with fuscous markings, and fully aquatic larval stage that feeds on submerged plants.
Parapoynx obscuralis
obscure pondweed moth, American china-mark, vallisneria leafcutter
Parapoynx obscuralis is an aquatic moth in the family Crambidae, first described by Augustus Radcliffe Grote in 1881. The species is native to North America, with a range extending from Nova Scotia to Florida and west to Texas, Wisconsin, and Ontario. It has been introduced to Great Britain, where it is naturalised in aquatic nurseries. The larvae are aquatic and feed on submerged aquatic plants.
Parapoynx seminealis
floating-heart waterlily moth
Parapoynx seminealis is a small crambid moth in the subfamily Acentropinae, commonly known as the floating-heart waterlily moth. It was described by Francis Walker in 1859. The species is associated with aquatic habitats and has been recorded from multiple states in eastern North America.
Pelocoris balius
Pelocoris balius is a species of saucer bug (family Naucoridae) endemic to Florida. Originally described as a subspecies of P. femoratus in 1970, it was elevated to full species status in 2015 based on sympatric occurrence with P. femoratus and distinct morphological characters. It is much less commonly encountered than its congener P. femoratus. In the Everglades, it shows a preference for shorter hydroperiod sites.
Pelocoris carolinensis
Carolina creeping water bug
Pelocoris carolinensis is a species of creeping water bug in the family Naucoridae, described by Torre-Bueno in 1907. It is found in the southeastern United States. Creeping water bugs are aquatic predators that inhabit freshwater environments.
Peltodytes callosus
Peltodytes callosus is a species of crawling water beetle in the family Haliplidae, first described by LeConte in 1852. It belongs to a genus of small aquatic beetles adapted to life in shallow water environments. The species is documented from North America, with distribution records extending into Middle America.
Peltodytes oppositus
Peltodytes oppositus is a species of crawling water beetle in the family Haliplidae, first described by Roberts in 1913. The genus Peltodytes contains approximately 20 species distributed across North America. Members of this genus are aquatic beetles associated with freshwater habitats. The specific epithet "oppositus" distinguishes this species from congeners, though detailed biological studies remain limited.
Peltodytes shermani
Peltodytes shermani is a species of crawling water beetle in the family Haliplidae, first described by Roberts in 1913. It is a small aquatic beetle known from North America.
Peltodytes tortulosus
Peltodytes tortulosus is a species of crawling water beetle in the family Haliplidae, described by Roberts in 1913. The species occurs in North America, with records from western Canada including Alberta and British Columbia. As a member of the Haliplidae, it inhabits aquatic environments and exhibits the characteristic crawling behavior of this beetle family.
Petrophila bifascialis
two-banded petrophila moth
Petrophila bifascialis is a small crambid moth found across eastern and central North America. Adults are active from late spring through early autumn. The species is notable for its aquatic larval stage, which feeds on algae and diatoms in freshwater habitats. Adults have a wingspan of 11–24 mm and display two distinct dark bands across the forewings.
Petrophila canadensis
Canadian petrophila moth, Canadian Petrophila
A small aquatic moth in the family Crambidae, described by Eugene G. Munroe in 1972. Adults have a wingspan of 11–18 mm and are active from May to September with two generations per year. The species is notable for its fully aquatic larval stage, which feeds on diatoms and algae in freshwater habitats. It overwinters as an adult.
Petrophila heppneri
Heppner's Petrophila
Petrophila heppneri is a small moth species in the family Crambidae, described in 1983 by André Blanchard and Edward C. Knudson. The species is known from Texas, where it inhabits riparian and aquatic-associated habitats typical of the genus Petrophila. Like other members of this genus, the larvae are presumed to be aquatic and feed on algae. Adults are nocturnal and can be attracted to light. The specific epithet honors Dr. J. B. Heppner, a lepidopterist who contributed significantly to the study of North American microlepidoptera.
Petrophila jaliscalis
Jalisco Petrophila
Petrophila jaliscalis is a small aquatic moth in the family Crambidae, described by Schaus in 1906. The species has a broad North American distribution spanning from Alberta, Canada through the western United States to Mexico. Adults are active nearly year-round and have forewings measuring 7–11 mm. The larvae are aquatic and feed on algae.
Philorus
Philorus is a genus of net-winged midges in the family Blephariceridae, established by Kellogg in 1903. Members of this genus are aquatic flies whose larvae inhabit fast-flowing streams and rivers, where they attach to rocks using specialized sucking discs. The genus is part of a family notable for having some of the most highly specialized larval adaptations among Diptera for life in torrential waters.
Phytobius
Phytobius is a genus of weevils in the family Curculionidae. At least one species, Phytobius vestitus, has been investigated as a biological control agent for invasive aquatic vegetation. The genus is recorded from northern Europe including Denmark, Norway, and Sweden.
Platambus semivittatus
Platambus semivittatus is a predaceous diving beetle in the family Dytiscidae. It is native to North America with a broad geographic range spanning from the Great Lakes region to the southwestern United States. The species belongs to the genus Platambus, a group of medium-sized diving beetles commonly found in freshwater habitats.
Plateumaris dubia
long-horned leaf beetle, Dubious spade beetle
Plateumaris dubia is a species of aquatic leaf beetle in the family Chrysomelidae. It is found in North America, with records from western Canada and Alaska. The species is commonly known as the long-horned leaf beetle or the dubious spade beetle. Like other members of the genus Plateumaris, it is associated with wetland habitats.
Plateumaris nitida
Lustrous reed beetle
Plateumaris nitida is an aquatic leaf beetle in the subfamily Donaciinae, commonly known as the Lustrous reed beetle. It is found in North America, with records from Canada (Alberta, British Columbia, Labrador) and the United States. As a member of the Donaciinae, it is associated with aquatic or semi-aquatic habitats, particularly wetlands with emergent vegetation.
Plauditus cestus
Plauditus cestus is a small mayfly in the family Baetidae, widespread across eastern and central North America. Recent records have extended its known range substantially, including a 1900 km northwestward expansion to the Northwest Territories. The species exhibits notable larval color variation, with some populations showing pronounced abdominal segment coloration.
Plectrocnemia
tube maker caddisflies
Plectrocnemia is a genus of tube maker caddisflies in the family Polycentropodidae comprising more than 120 described species. Larvae are aquatic predators that construct silken capture nets to intercept prey. The genus has been extensively studied for its larval silk production, vibration-mediated predatory behavior, and population genetics. Species occur across Europe and into western Asia, with detailed biological information available for several well-studied species including P. conspersa and P. brevis.
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aquatica
water springtail
Podura aquatica, commonly known as the water springtail, is one of only four described species in the family Poduridae. It is exclusively aquatic, living its entire life on the surface of still water bodies where it scavenges. The species exhibits specialized sensory adaptations for locating water surfaces through polarized light detection. Adults possess a large, flattened furcula that enables jumping without breaking water surface tension. The species has a Holarctic distribution and is considered abundant across its range.
Poduridae
Water Springtails
Poduridae is a small family of springtails (Collembola) containing a single genus, Podura. Members are stout-bodied and commonly known as water springtails due to their association with aquatic or semi-aquatic habitats. The family constitutes the monotypic superfamily Poduroidea and includes four described species. These springtails are part of the order Poduromorpha, one of the three major lineages of Collembola.
Poecilocera
Poecilocera is a monotypic genus of aquatic leaf beetles in the family Chrysomelidae, containing the single species Poecilocera harrisii. The genus was established by Schaeffer in 1919. The name has been used historically in both Coleoptera and Orthoptera, creating taxonomic confusion.
Psephenus herricki
Herrick's Water Penny Beetle
Psephenus herricki is a water penny beetle found in eastern North America. Larvae are highly flattened and inhabit stream riffles, where they occupy interstitial crevices and undersides of stones. The species exhibits specific behavioral and morphological adaptations to flowing water environments, including thigmokinesis that inhibits movement when wedged in substrate. Adults are associated with wave-splashed rocks that serve as mating sites and oviposition substrates.
Pseudochironomus
nonbiting midge
Pseudochironomus is a genus of nonbiting midges in the family Chironomidae, containing more than 20 described species distributed across the Americas and Europe. The genus belongs to the tribe Pseudochironomini within subfamily Chironominae. Larvae are aquatic and have been studied for their growth responses to diet quality and thermal conditions.
Ranatra
water stick-insect, water scorpion, water mantis
Ranatra is a genus of slender aquatic predatory insects in the family Nepidae, commonly called water stick-insects or water scorpions. The genus contains over 140 species distributed across freshwater habitats worldwide, with highest diversity in South America and Asia. These insects are sit-and-wait predators that use raptorial forelegs to capture prey. They possess a distinctive long caudal siphon for breathing while submerged.
Ranatra drakei
Ranatra drakei is a species of water scorpion in the family Nepidae, described by Hungerford in 1922. Like other members of the genus Ranatra, it is an aquatic predatory insect with raptorial forelegs adapted for capturing prey. The species is known from North America, though specific ecological details remain poorly documented in published literature.
Ranatra fusca
brown water scorpion, brown waterscorpion
Ranatra fusca is a predatory aquatic bug in the family Nepidae, commonly called the brown water scorpion. Native to North America, it inhabits freshwater environments where it hunts small invertebrates using raptorial forelegs. Adults reach 3.2–4.2 cm in length and possess a long respiratory siphon for breathing at the water surface. The species is most active from spring through autumn.
Ranatra montezuma
Montezuma waterscorpion
Ranatra montezuma is a waterscorpion species endemic to Montezuma Well, a thermally constant, fishless spring in Yavapai County, Arizona. It was described by J. Polhemus in 1976 and represents a highly localized population with specific dietary adaptations to its unique aquatic environment. The species exhibits nocturnal foraging behavior and maintains high population densities supported by abundant endemic prey.
Ranatra quadridentata
waterscorpion
Ranatra quadridentata is a species of waterscorpion described by Stål in 1862. As a member of the family Nepidae, it is an aquatic predatory insect that inhabits freshwater environments. The species has been documented in Central America and North America, with 188 observations recorded on iNaturalist.
Rhagovelia obesa
riffle bug
Rhagovelia obesa is a small aquatic true bug in the family Veliidae, commonly known as a riffle bug or smaller water strider. It inhabits flowing water environments where it uses a unique swimming plume to move underwater. The species has an unusual life history among aquatic Hemiptera, with four nymphal instars rather than five and eggs as the overwintering stage. It has been successfully reared in laboratory conditions from first instar to adult.
Rhantus
Rhantus is a genus of predaceous diving beetles (Dytiscidae) comprising approximately 100 species with worldwide distribution. Members of this genus are primarily associated with lentic freshwater habitats, particularly pools and marshy wetlands. The genus is currently recognized as paraphyletic; taxonomic revision is anticipated following 2017 research that transferred 17 species to newly established genera (Nartus, Meridiorhantus, Caperhantus, and Carabdytes). Several species have independently colonized oceanic islands and undergone endemic diversification.
Rhantus wallisi
Rhantus wallisi is a predaceous diving beetle in the family Dytiscidae, described by Hatch in 1953. The species is endemic to North America, with documented occurrences in western Canada including Alberta and British Columbia. Like other members of the genus Rhantus, it is an aquatic predator inhabiting freshwater environments.
Rhizelmis
Rhizelmis is a genus of riffle beetles (family Elmidae) established by Chandler in 1954. These beetles are aquatic insects inhabiting running water environments. The genus is part of the tribe Elmini within the subfamily Elminae.
Sanfilippodytes pacificus
Sanfilippodytes pacificus is a predaceous diving beetle in the family Dytiscidae. The species was described by Fall in 1923 and is known from western North America, including Alaska and British Columbia. Like other members of Dytiscidae, it is an aquatic predator.
Sepedon americana
American snail-killing fly
Sepedon americana is a species of marsh fly in the family Sciomyzidae, commonly known as snail-killing flies. The species was described by Steyskal in 1951. Like all members of the genus Sepedon, its larvae are obligate predators of aquatic pulmonate snails. The species occurs in North America and has been documented in regions including Colorado. Adults are active early in the year, with observations in February recorded.
Sepedon armipes
marsh fly
Sepedon armipes is a North American species of marsh fly in the family Sciomyzidae, commonly known as snail-killing flies. Like other members of its genus, its larvae are aquatic predators of pulmonate snails. The species exhibits early-season activity, with adults observed active as early as February.
Sepedon floridensis
Florida snail-killing fly
A species of snail-killing marsh fly in the family Sciomyzidae, endemic to Florida. Adults are active in early spring and are associated with aquatic habitats where their predatory larvae feed on pulmonate snails. The species was described by Steyskal in 1951 and belongs to a genus with 20 recognized species in North America, many of which have been investigated for potential biological control of snail vectors of parasitic diseases.
Sepedon fuscipennis
marsh fly
Sepedon fuscipennis is a species of marsh fly in the family Sciomyzidae. The larvae are aquatic predators of pulmonate snails, living just beneath the water surface suspended by water-resistant hairs surrounding their posterior spiracles. Adults are slender flies with a concave face and jutting antennae; males have distinctive bowed hind tibiae and swollen, toothed hind femora for grasping females during mating. The species exhibits overlapping generations during summer with unimodal population peaks, and has been studied for its potential in biological control of snail hosts of parasitic diseases.
Sialis rotunda
Alderfly
Sialis rotunda is a species of alderfly in the family Sialidae, order Megaloptera. It is a small to medium-sized aquatic insect whose larvae develop in freshwater habitats. Adults are short-lived and primarily focus on reproduction. The species was described by Nathan Banks in 1920 and is native to western North America.
Sigara
water boatmen
Sigara is a genus of water boatmen (family Corixidae) comprising approximately 100 species of aquatic true bugs. The genus is notable for containing halophilic species capable of inhabiting saline and hypersaline waters, including extreme environments such as the Makgadikgadi Pans in Botswana. Members of this genus are aquatic insects found in various freshwater and saline habitats across multiple continents.
Sigara trilineata
Sigara trilineata is a species of water boatman in the family Corixidae, found in North America. Water boatmen are aquatic insects that inhabit freshwater environments. The species name "trilineata" refers to three lines, likely describing a distinctive marking pattern on the body. As a member of the Corixidae, it possesses the family's characteristic oar-like hind legs adapted for swimming.
Sisyra
spongillafly, spongilla-fly
Sisyra is a genus of spongillaflies (Neuroptera: Sisyridae), a small family of aquatic insects. The genus comprises at least eight described species distributed across multiple continents, including Brazil, Panama, Australia, and Asia. Sisyra species have been documented in freshwater habitats, with larvae associated with freshwater sponges. The family holds a basal phylogenetic position within Neuroptera, and at least one species (S. nigra) exhibits an unusual achiasmatic sex chromosome segregation mechanism not observed in other neuropterans.
Sperchonopsis
Sperchonopsis is a genus of water mites in the family Sperchontidae. The genus was established by Piersig in 1896 and contains species found in Europe and North America. Sperchonopsis phreaticus is the only formally described species currently associated with this genus. These mites belong to the diverse group of prostigmatic mites within the order Trombidiformes.
Sperchopsis tessellata
Checkered Water Beetle
Sperchopsis tessellata, commonly known as the Checkered Water Beetle, is a species of water scavenger beetle in the family Hydrophilidae. It is distributed across eastern North America, from Canada through the eastern and central United States. As a member of the Hydrophilidae family, it is adapted to aquatic environments. The species was first described by Ziegler in 1844 under the basionym Spercheus tessellatus.
Sphaerius
Sphaerius is a genus of minute beetles in the family Sphaeriusidae, comprising 18 extant species. These beetles are among the smallest in Coleoptera, measuring 0.5–1.2 mm in length. They inhabit damp environments near water bodies across all continents except Antarctica. The genus has undergone significant taxonomic revision, with its family name changing from Sphaeriidae to Microsporidae and finally to Sphaeriusidae.
Stenelmis
riffle beetle
Stenelmis is the largest and most widespread genus of beetles in the family Elmidae. Members are commonly known as riffle beetles due to their association with fast-flowing stream habitats. The genus contains numerous species distributed across multiple continents, with documented presence in North America, Europe, and the Caucasus region.
Stenelmis bicarinata
riffle beetle
Stenelmis bicarinata is a species of riffle beetle in the family Elmidae, first described by LeConte in 1852. Riffle beetles are aquatic insects associated with running water habitats. The species occurs in Central and North America, though detailed ecological studies remain limited.
Stenocolus
Stenocolus is a monotypic genus of beetles in the family Eulichadidae. The sole described species, S. scutellaris, is a forest stream beetle found in western North America. The genus belongs to a small family of aquatic or semi-aquatic beetles associated with running water habitats.
Stygoparnus
Comal Springs dryopid beetle (for sole species S. comalensis)
Stygoparnus is a monotypic genus of dryopid beetles containing the single species Stygoparnus comalensis, known as the Comal Springs dryopid beetle. The genus is endemic to central Texas spring systems fed by the Edwards Aquifer. Its sole member is a federally endangered, subterranean-obligate aquatic beetle with vestigial eyes and reduced pigmentation. The genus represents the only known aquatic lineage within the family Dryopidae.
Stygoparnus comalensis
Comal Springs dryopid beetle
Stygoparnus comalensis is a federally endangered, subterranean-obligate dryopid beetle endemic to three spring systems in the Edwards Aquifer region of central Texas. Adults are approximately 2 mm long with vestigial eyes and undeveloped wings, reflecting their troglobitic ancestry. The species exhibits extreme population genetic structure with no gene flow among localities, indicating evolutionary isolation. Captive studies reveal strong habitat preferences for wood substrates and specific phototactic responses to light wavelength. The beetle serves as a surrogate species for monitoring aquifer health and faces threats from groundwater extraction, pollution, and drought-induced springflow reduction.
Suphis
Suphis is a genus of small aquatic beetles in the family Noteridae, first described by Aubé in 1836. The genus contains approximately eleven described species distributed primarily in the Americas, with records from North, Central, and South America. Members of this genus are associated with freshwater habitats, particularly wetlands and slow-moving waters. As part of the Noteridae family, they represent one of several genera within the diverse group of predaceous diving beetles.
Suphisellus puncticollis
Suphisellus puncticollis is a burrowing water beetle described by Crotch in 1873. It belongs to the subfamily Noterinae within the family Noteridae, a group of small aquatic beetles adapted to living in muddy substrates. The species has been recorded in Canada (Ontario, Québec) and the United States.
Synaptonecta issa
Synaptonecta issa is a minute water bug species in the family Micronectidae, first described by Distant in 1910. The species belongs to a genus of tiny aquatic true bugs that inhabit freshwater environments. Limited observational data exists, with only six iNaturalist records documented. The species has a disjunct distribution spanning North America and Southern Asia.
Tabanus atratus
Black Horse Fly
Tabanus atratus, commonly known as the black horse fly, is a large, all-black species of horse fly in the family Tabanidae. It is one of the largest horse flies in North America, with body lengths ranging from 20–25 mm. Females are blood-feeders with scissor-like mouthparts used to cut skin and lap blood, while males have reduced mouthparts and do not feed on blood. The species is recognizable by its uniform black coloration and distinctively hooked antennae. Females lay eggs in large masses of 100–1000 eggs near water's edge, with each female capable of producing three to four egg masses.
Tanysphyrus lemnae
Duckweed Weevil
Tanysphyrus lemnae is a small weevil species native to Europe with established populations in North America. The larvae are leaf-miners that develop within duckweed plants of the genus Lemna. The species is commonly known as the Duckweed Weevil due to this specialized host relationship.
Tetanocera ferruginea
common buff snailkiller
Tetanocera ferruginea is a medium to large sciomyzid fly with light brown coloration and no obvious wing markings. It is an obligate malacophage, with larvae preying on non-operculate snails in aquatic and semi-aquatic habitats. The species has a Holarctic distribution and is multivoltine, overwintering in the puparial stage. Despite forming large populations, individuals exhibit very limited movement within habitats.
Tetragnatha
Stretch Spiders, Long-jawed Orb Weavers
Tetragnatha is a large genus of long-jawed orb-weaving spiders with hundreds of species distributed worldwide, though most occur in tropical and subtropical regions. These spiders are characterized by their elongated bodies, extremely long legs, and notably elongated chelicerae (jaws) with well-developed teeth. They construct horizontal orb webs, often near water, and many species can run across water surfaces. The genus exhibits remarkable variation in dispersal ability, with some species being excellent dispersers capable of colonizing islands, while others appear to have secondarily lost this capacity. Hawaiian Tetragnatha species have undergone adaptive radiation, with some lineages evolving cursorial (running) behavior and abandoning web-building entirely.
spiderorb-weaverlong-jawedaquaticstretch-spideradaptive-radiationHawaiicursorial-evolutionhorizontal-webwater-walkingcamouflagemultivoltinesocial-spiderTetragnathidaeAraneaeArachnidapredatorinsectivoreriparianwetlandmeadowtropicalsubtropicalcosmopolitandispersalisland-colonizationphylogeographychemical-species-recognitioncommunal-webbingkleptoparasitism-targetThermonectus nigrofasciatus nigrofasciatus
Thermonectus nigrofasciatus nigrofasciatus is a subspecies of predatory diving beetle in the family Dytiscidae. The nominate subspecies occurs in North and Middle America. Members of the genus Thermonectus are recognized for their distinctive color patterns and aquatic lifestyle.
Tipula abdominalis
giant crane fly
Tipula abdominalis, commonly known as the giant crane fly, is a large species of crane fly in the family Tipulidae. The larvae are aquatic detritivores found in riparian habitats, where they feed on decomposing leaf litter. Their hindgut harbors a dense, diverse bacterial community that facilitates digestion of lignocellulosic material. The species has been studied for its potential applications in biofuel production due to its efficient natural biorefinery system. Adults are among the largest crane flies in North America but do not feed.
Trichoclinocera
Trichoclinocera is a genus of aquatic dance flies in the family Empididae, subfamily Clinocerinae. The genus contains approximately 30 described species distributed across the Holarctic and eastern Asia, with 16 species recognized from the Nearctic region and nine from Japan. Species are organized into six species-groups based on morphological characteristics, primarily of the male terminalia. The genus was established by Collin in 1941 and has undergone significant taxonomic revision, with Seguyella and Acanthoclinocera recently synonymized under Trichoclinocera.
Xylotopus par
Xylotopus par is an aquatic xylophagous midge in the family Chironomidae. The larvae colonize submerged wood substrates in freshwater streams, feeding on decaying wood. Populations exhibit plastic phenology, with development rates varying based on wood substrate quality—larvae in fresh, introduced wood can complete development in a single summer, while those in indigenous logs normally require a full year. The species has been studied for its unique bacterial associations in the larval midgut, where morphotypically uniform bacteria form a well-defined band in the posterior midgut's ectoperitrophic space.
Zapada frigida
Frigid Forestfly
Zapada frigida is a species of spring stonefly in the family Nemouridae, commonly known as the Frigid Forestfly. Like other members of the genus Zapada, adults retain diagnostic branched cervical gills from the nymphal stage, with two sets of branched gills on each side of the body. The species has been documented from western North America including Alberta, Alaska, British Columbia, California, and Colorado.