High-altitude
Guides
Aedes epactius
Western Rock Pool mosquito
Aedes epactius is a North American mosquito species in the subgenus Georgecraigius, first described in 1908. It exhibits remarkable altitudinal plasticity, occurring from sea level to over 2,500 meters elevation. The species breeds in both natural rock pools and artificial containers including tires, water tanks, and flower pots. It has been studied for potential arbovirus transmission, with laboratory evidence of competence for St. Louis encephalitis and Jamestown Canyon viruses, and field detection of West Nile and Zika virus RNA. Recent research suggests it may compete with invasive Aedes aegypti, potentially limiting its spread in some habitats.
Bombus cockerelli
Cockerell's Bumble Bee, Cockerell's Bumblebee
Bombus cockerelli is a bumble bee species with the smallest known range of any bumble bee species worldwide, restricted to high-elevation localities in the White Mountains of New Mexico. First described in 1913 from six specimens, it remained unobserved between 1956 and 2011. Fewer than 30 specimens have ever been collected. The species was once considered a potential subspecies, but fresh specimens now allow for genetic analysis to resolve its taxonomic status. Its entire range falls within protected U.S. National Forest and tribal lands.
Bombus flavifrons
yellow-fronted bumble bee, yellowhead bumblebee
Bombus flavifrons is a robust bumble bee native to North America, distributed across much of Canada, Alaska, and the western contiguous United States. The species exhibits intraspecific color pattern variation across its range due to Müllerian mimicry. It inhabits high-altitude and high-latitude environments including tundra, taiga, and mountain forests and meadows. The species is host to the parasitic cuckoo bumble bee Bombus insularis.
Boreidae
snow scorpionflies, snow fleas
Boreidae, commonly known as snow scorpionflies or snow fleas, are a small family of approximately 30 species of scorpionflies in the order Mecoptera. These tiny, wingless insects are notable for their unusual winter activity period, emerging on snow surfaces during cold months when most other insects are dormant. Recent molecular research indicates Boreidae are more closely related to fleas (Siphonaptera) than to other scorpionflies, a finding that has significant implications for understanding the evolutionary relationships within Mecoptera. The family has a boreal and high-altitude distribution in the Northern Hemisphere, with species documented across North America, Europe, and Asia.
Branchinecta
fairy shrimp
Branchinecta is a genus of fairy shrimp (Anostraca) comprising approximately 50 species distributed across all continents except Australia. The genus includes the largest anostracan, Branchinecta gigas, reaching up to 10 cm in length, and the highest-altitude crustacean, B. brushi, recorded at 5,930 m elevation. Species occupy diverse temporary freshwater habitats from Arctic and Antarctic regions to high-elevation Andean pools. Two species, B. gaini and B. granulosa, have been synonymized based on molecular evidence showing insufficient genetic differentiation.
Coleomyia alticola
Coleomyia alticola is a species of robber fly (family Asilidae) described by James in 1941. The specific epithet 'alticola' suggests a high-altitude habitat association. As a member of the Asilidae, it belongs to a family of predatory flies known for their aerial hunting behavior. Very few observations of this species have been recorded.
Cosmiina
Cosmiina is a subtribe of owlet moths (Noctuidae) within the tribe Xylenini. It comprises multiple genera including Ipimorpha, which contains species with broad geographic distributions spanning Europe, Asia, and North America. Members of this subtribe have been documented in high-altitude wetland habitats in China.
Euophrys
Euophrys is a genus of jumping spiders (family Salticidae) first described by Carl Ludwig Koch in 1834. The genus contains approximately 99 species distributed across Eurasia, Africa, and the Americas. Notable members include E. omnisuperstes, which inhabits Mount Everest at elevations up to 6,700 meters, and E. monadnock, a colorful North American species with distinctive male courtship displays. Species in this genus exhibit typical salticid characteristics including excellent vision and complex visual communication.
Hydatostega
Hydatostega is a genus of long-legged flies in the family Dolichopodidae. The genus was restored from synonymy with Hydrophorus and is now recognized as distinct. It contains approximately 11 described species with a disjunct distribution pattern.
Liodessus
Liodessus is a genus of small diving beetles in the family Dytiscidae, tribe Bidessini. Species occur across the Americas from North America through the Andes to southern South America, with notable diversity in high-altitude wetlands and páramo ecosystems. Many species inhabit shallow, exposed pools and temporary water bodies, showing phenotypic plasticity in body form correlated with habitat permanence. The genus contains numerous species, with several new species described from the high Andes of Peru and Colombia in recent years. Taxonomic resolution relies heavily on male genital morphology.
diving-beetleaquatic-insecthigh-altitudepáramoAndesphenotypic-plasticityBidessiniHydroporinaeDytiscidaeColeopterawetlandtemporary-poolpeatlandsubantarctictaxonomymale-genitaliaCOI-barcodecryptic-speciesincipient-speciationhybridizationgeometric-morphometricsenvironmental-sentineldrought-indicatorSouth-AmericaNorth-AmericaFijiNavarino-IslandMagallanesPeruColombiaChileArgentinaBoliviaBogotáJunínCuscoHuánucoCundinamarcaSumapazPunasteppepeat-bogshallow-poolpuddlealtitude-3400-4900mL.-affinis-complexL.-bogotensis-complexL.-chilensisL.-affinisL.-obscurellusL.-noviaffinisL.-saratogaeL.-picinusL.-santarositaL.-alpinusL.-hauthiL.-rhigosL.-thespesiosGuignot-1939ZimmermannSharpLeConteSaySteinheilRégimbartBiströmMillerBalkeFranciscoloSanfilippoPederzaniNilssonYoungClarkHatchMannerheimCaseyAubéBalfour-BrowneBrinckPeschetSolierWhiteZootaxaZooKeysInsect-Systematics-&-EvolutionAnales-del-Instituto-de-la-PatagoniaPLoS-ONEGBIFiNaturalistCatalogue-of-LifeNCBI-TaxonomyBOLDBarcode-of-Life-Data-SystemCOImitochondrial-DNAphylogeographypopulation-structurerange-expansionlineagegenetic-structuresubspecieslectotypeneotypesynonymynew-speciesnew-subspeciesrevisionmorphometricsallometrybody-sizebody-shapeelytrapronotumstriaecolorationpunctationhabitusoccipital-linegenital-morphologyaedeagusparamerediagnosisidentification-keytype-localitytype-specimenetymologybionomicsnatural-historygeographic-distributiondistribution-mapNearcticNeotropicalPatagoniaCape-Horn-Biosphere-Reservedroughtclimate-changeenvironmental-changesentinel-speciesplasticityadaptationphenotypic-variationhydrological-landscapewater-permanencepoolbogAltiplanoAndeanhigh-elevationmontanealpineaquaticfreshwaterlenticloticinsectbeetleAdephagaLiodessusOreoneta
A genus of dwarf spiders (family Linyphiidae) established in 1894, comprising approximately 30 species distributed across cold-temperate and Arctic regions of the Northern Hemisphere. Most species inhabit high-latitude or high-altitude environments. The genus was revised extensively by Saaristo & Marusik in 2004, resulting in numerous new species descriptions.
ArachnidaAraneaeLinyphiidaedwarf-spiderArcticborealHolarctictundramontanepredatorcold-adaptedground-dwellinghigh-latitudehigh-altitudeSaaristoMarusikChyzerKulczyński18942004-revisiongenitalic-charactersepigynumembolusconductorpalaearcticnearcticEurasiaNorth-AmericaScandinaviaSiberiaAlaskaCanadaGreenlandKazakhstanMongoliaChinaKurile-IslandsFinlandSwedenNorwayGermanySwitzerlandItalySlovakiaBulgariaCentral-Europetype-species:-Oreoneta-frigidathirty-species30-speciessmall-spidersheet-web-weavermoney-spiderLinyphiinaeErigoninaemicrohabitatlitter-dwellingcrypticunderstudiedtaxonomic-revisionmorphological-taxonomylight-microscopyspecies-descriptionbiogeographyglacial-refugiapost-glacial-colonizationendemicregional-endemicalpine-endemicArctic-endemicboreal-foresttaigarocky-habitatmeadowwetlandstream-marginsnowbedfellfieldheathmosslichenvegetationsoil-surfacestonerockboulderscreetalusdebrishumusorganic-layerpermafrostpolarsubpolarcold-stenothermstenothermstenotopichabitat-specialistconservationraredata-deficientIUCNred-listnatural-historysystematicsphylogenymolecular-systematicsDNA-barcodingCOIcytochrome-oxidase28S18SITSmorphologytaxonomyalpha-taxonomyrevisionmonographfaunisticschorologyzoogeographydispersalvicariancespeciationcryptic-speciesspecies-complexspecies-groupspecies-aggregatesubspeciesinfraspecificvariationpolymorphismsexual-dimorphismmalefemalejuvenileinstarecdysismoltingdevelopmentgrowthreproductionmatingcourtshipsperm-transferspermatophorepedipalppalpal-organcopulatory-organgenitaliaepigynevulvaspermathecafertilization-ductcopulatory-ductatriumscapemacerationdissectionmicroscopestereomicroscopecompound-microscopeillustrationdrawingphotographySEMscanning-electron-microscopyhabitusdorsalventrallateralprosomaopisthosomaabdomencephalothoraxcarapacesternumchelicerafanglegtarsusmetatarsustibiapatellafemurcoxatrochanterspinneretcolulustracheal-spiraclebook-lungcuticlesetahairspinemacrosetasensillumchaetotaxyeyeoculusocellianterior-median-eyeposterior-median-eyeanterior-lateral-eyeposterior-lateral-eyeeye-tubercleclypeusfoveapedicelwaistconstrictioncolorpatternpigmentationmelanismalbinismsizemeasurementtotal-lengthcarapace-lengthcarapace-widthabdomen-lengthleg-lengthleg-formulaindexratioproportionallometryontogenysubadultadultmaturitysexual-maturityreproductive-maturitylongevitylife-spangeneration-timevoltinismunivoltinesemivoltinebiennialperennialoverwinteringdiapausequiescencecold-hardinessfreeze-tolerancefreeze-avoidancesupercoolingcryoprotectantglycerolsorbitoltrehalosethermal-biologythermoregulationbehavioral-thermoregulationpostural-thermoregulationactivitydiurnalnocturnalcrepuscularmatutinalvespertinecircadianrhythmseasonalphenologycalendardatemonthactivity-periodbreeding-seasonovipositioneggegg-saccocoonsilkwebsheet-webtangle-webspace-webplatform-webdome-webbowl-webfunnelretreatsignal-linedraglineattachment-discampullate-glandcylindriform-glandaciniform-glandaggregate-glandflagelliform-glandtubuliform-glandpiriform-glandmajor-ampullateminor-ampullatesilk-proteinspidroinfibroinsericingluevisciddrycapture-silkdragline-silksafety-lineanchor-lineframe-lineradial-linespiral-linesticky-spiralnon-sticky-spiralhubperimetersubstratenicheguildfunctional-grouptrophic-levelcarnivorezoophagearthropodivoreinsectivorepreycaptureattacksubdueimmobilizekillfeeddigestextraoral-digestionsucking-stomachgutmidguthindgutMalpighian-tubuleexcretionosmoregulationmetabolismrespirationtracheaspiraclegas-exchangecirculationhemolymphheartaortasinusnervous-systembrainganglionnervesensory-organmechanoreceptorchemoreceptorthermoreceptorhygroreceptorphotoreceptorproprioceptortrichobothriumtarsal-organslit-sensillumlyriform-organpedipalp-organHaller's-organcontact-chemoreceptorgustatoryolfactorypheromonesemiochemicalkairomoneallomonesynomonecommunicationsignalvibrationseismicairbornevisualchemicaltactilecopulationinseminationspermspermatozoonmedian-apophysisterminal-apophysisparacymbiumcymbiumbulbustegulumsubtegulumradixlamellaprocessapophysisflangehookcoilspiralductreservoirscleriteplatefieldfossapocketreceptaculumspermathecaeglandinsemination-ductuterusuterus-externusuterus-internusvaginagonoporeovipositorovaryoviductfolliclevitellogenesisoogenesisspermatogenesistestisvas-deferensejaculatory-ductseminal-vesiclefecundityfertilityclutch-sizeegg-numberhatchingemergenceballooningaerial-dispersalsilk-draglinegossamermigrationmovementwalkingrunningjumpingclimbingburrowinghidingrefugeshelterdefensepredationpredator-avoidanceantipredatorcrypsiscamouflagemimicryaposematismdeimaticthanatosisautotomyleg-lossregenerationinjurywoundhealingimmune-responsehemocytemelanizationencapsulationnoduleparasiteparasitoidpathogenfungusbacteriumvirusnematodemitegroomingcleaninghygieneself-maintenancesilk-useweb-useprey-captureforaginghuntingambushsit-and-waitactive-huntingsearchingprobingtestingexplorationlearningmemorycognitionbehaviorethologyecologypopulationcommunityecosystembiomebiospherehabitatenvironmentabioticbioticfactortemperaturehumiditymoistureprecipitationrainsnowwindlightphotoperiodday-lengthradiationUVsoilcovercomplexityheterogeneityproductivityenergynutrientcarbonnitrogenphosphorusfood-webtrophic-interactionpredator-preycompetitionintraspecificinterspecificresourcepartitioningcoexistencediversityrichnessevennessabundancedensityfrequencyoccurrencepresenceabsencedetectionsamplingcollectionpitfall-trapquadrattransecthand-collectingbeatingsiftingextractionBerleseTullgrenWinklerflotationsortingpreservationethanolformalinmountslidelabelcatalogdatabaserepositorymuseumvouchertypeholotypeparatypesyntypelectotypeparalectotypeneotypetopotypeoriginaldatalocalitygeoreferencecoordinatelatitudelongitudeelevationaltitudedepthcollectordeterminationidentificationexpertspecialistliteraturepublicationjournalarticlepaperbookthesisdissertationreportchecklistonlinedigitalarchivebibliographyreferencecitationauthoryearvolumepagefiguremaptableappendixsupplementarymaterialmethodsresultsdiscussionconclusionabstractkeywordindex-termsubjecttopicdisciplinesciencebiologyzoologyarachnologyentomologyevolutionphylogeneticsbiologicalnatureprotectionmanagementpolicylegislationrestorationmonitoringassessmentindicatorflagshipumbrellakeystoneengineerservicefunctiondynamicsstabilityresilienceresistancerecoverysuccessiondisturbanceclimatechangewarmingglacierrangeshiftexpansioncontractionfragmentationisolationconnectivitycorridornetworkreserveparkprotectedareawildernessurbanruralagriculturalforestryminingimpactthreatriskvulnerabilitysensitivityexposureadaptivecapacitymitigationadaptationsustainableuseharvesttradeeconomymedicineresearcheducationoutreachcitizenengagementawarenessappreciationaestheticculturalspiritualexistencevalueintrinsicinstrumentalutilitariananthropocentricbiocentricecocentricethicphilosophyhistorydiscoveryexpeditionvoyagesurveyinventoryfaunabiotabiodiversitylifeearthplanetwildlifeanimalinvertebratearthropodcheliceratearachnidspideraraneomorphentelegynemicryphantidOreoneta2004frigidatype-speciesalpinefeedingtype-specimenclimate-changeprotected-areaParnassius eversmanni
Eversmann's parnassian
Parnassius eversmanni, or Eversmann's parnassian, is a high-altitude butterfly in the swallowtail family Papilionidae. It occurs across a broad transcontinental range spanning eastern Russia, Mongolia, Japan, Alaska, and the Yukon. The species was named in honor of Eduard Friedrich Eversmann. Like other Parnassius species, it exhibits complex wing pattern variation with numerous described subspecies and forms, making identification challenging without examination of structural characters.
Puto
giant mealybugs, Puto scale insects
Puto is a genus of scale insects in the family Putoidae, commonly known as giant mealybugs. The genus was established by Signoret in 1875 and is distinguished by its large body size relative to other mealybugs, dense white wax coating, and distinctive morphological features including cerarii and multilocular disc-pores. Putoidae is recognized as a distinct family separate from Pseudococcidae, representing a transitional evolutionary group between archaeococcoids and neococcoids. Species in this genus are primarily associated with woody host plants in temperate and montane regions.
Titanoecidae
Rock Weavers
Titanoecidae is a family of cribellate spiders comprising five genera and approximately 67 species. Members are primarily dark-colored web-builders that construct woolly, cribellate silk webs. The family occurs across the New World and Eurasia, with several species inhabiting high-altitude mountain environments where they can achieve high local abundance. The family was elevated from Amaurobiidae by Lehtinen in 1967.
Trechini
Trechini is a large and diverse tribe of small ground beetles in the family Carabidae, comprising over 270 genera and more than 2,400 described species worldwide. Members are typically diminutive, often less than 5 mm in length, with many species measuring 1 mm or less. The tribe exhibits remarkable habitat breadth, occurring in terrestrial, subterranean, and high-altitude mountain environments across all continents except Antarctica, though fossil evidence indicates former presence there. Trechini includes numerous cave-adapted lineages, making it the most diverse and cave-specialized group among Carabidae tribes in subterranean habitats.