Endangered

Guides

Microhexura montivaga
Spruce-fir moss spider
A miniature mygalomorph spider endemic to the highest peaks of the southern Appalachian Mountains. First described in 1925, this species inhabits moss growing on rocks beneath spruce-fir forest canopy. Listed as federally endangered in 1995 due to severe declines in its montane forest habitat. Genetic studies reveal significant population fragmentation across six allopatric populations, though morphological differences remain minimal.
endangered mygalomorph sky-island bryophyte-associated montane-endemic federally-listed
Nehalennia
Sprites
Nehalennia is a genus of very small damselflies in the family Coenagrionidae, commonly known as Sprites. Most species occur in North and South America, with one species, N. speciosa, distributed in Eurasia. Members of this genus are among the smallest odonates and are associated with specific wetland microhabitats.
damselfly odonata wetland peatland endangered microrefugium tyrphophilous conservation
Neonympha
satyr butterflies
Neonympha is a genus of satyrid butterflies occurring in the Nearctic and Neotropical regions. The genus contains several recognized species, including the endangered Mitchell's satyr (Neonympha mitchellii) with its two subspecies: the nominate form in Michigan prairie fens and the Saint Francis satyr (N. m. francisci) restricted to North Carolina. These butterflies exhibit strong habitat specificity to wetland environments, particularly those with sedge vegetation.
butterflies satyrinae wetland-dependent endangered Nearctic Neotropical
Neonympha mitchellii
Mitchell's Satyr, Mitchell's Marsh Satyr, Saint Francis' Satyr
Neonympha mitchellii is a federally endangered nymphalid butterfly of the eastern United States. The species comprises two recognized subspecies: N. m. mitchellii (Mitchell's satyr) in Michigan and Indiana, and N. m. francisci (Saint Francis' satyr) restricted to a single metapopulation at Fort Bragg, North Carolina. Additional populations discovered in Alabama, Mississippi, and Virginia since 1998 remain under taxonomic study. All populations are protected under the Endangered Species Act.
endangered federally-protected wetland-specialist prairie-fen invasive-plant-association
Nomotettix cristatus
crested pygmy grasshopper, crested grouse locust, northern crested grouse locust
Nomotettix cristatus is a small pygmy grasshopper in the family Tetrigidae, commonly known as the crested pygmy grasshopper or crested grouse locust. It is one of approximately 35 Nearctic species of Tetrigidae. The species exhibits three recognized subspecies with distinct geographic distributions across North America. Like other members of its family, it is characterized by an elongated pronotum that extends over the abdomen, a trait distinguishing pygmy grasshoppers from typical grasshoppers in Acrididae.
pygmy-grasshopper groundhopper Tetrigidae Nearctic moist-habitat pronotum cristate North-America leaf-litter aquatic-margin Orthoptera Caelifera jumping-insect small-grasshopper crested-pronotum subspecies-variation Alberta Connecticut Florida Georgia Illinois lake-shore pond-edge stream-margin ground-dwelling Scudder-1862 Batrachidea-cristata Nomotettix-cristatus-cristatus Nomotettix-cristatus-compressus Nomotettix-cristatus-floridanus Tetriginae Acrididea Tetrigoidea Hexapoda Pterygota insect arthropod animal eukaryote metazoa animalia arthropoda insecta nomotettix cristatus crested pygmy grasshopper grouse-locust northern-crested-grouse-locust crested-grouse-locust crested-pygmy-grasshopper iNaturalist GBIF Catalogue-of-Life NCBI-Taxonomy Wikipedia Skejo Tumbrinck Devriese Hochkirch Morse-1895 Hancock-1902 Scudder-1863 Batrachidea cristata compressus floridanus Nearctic-region 35-species 2000-species 230-million-years dinosaur-contemporaries water-dependent-lifecycle Peruća-lake Croatia Balkans Europe systematics taxonomy faunistics natural-history biodiversity conservation endangered imperiled ZooKeys research amateur-naturalist citizen-science observation 53-observations rare neglected-diversity tropics Madagascar Australia New-Guinea Borneo Brazil Atlantic-Forest leaf-like-morphology spines warts undulations horns bizarre grotesque Hancock-1907 Genera-Insectorum Acididae pronotal-projection Holocerus Notocerus Holocerus-lucifer Holocerus-devriesei Notocerus-formidabilis Metopomystrum-muriciense Ophiotettix-pulcherrima Paraselina-brunneri Selivinga-tribulata Criotettix-bispinosus Tetrix-subulata Tettigidea-lateralis Tetrix-undulata Tetrix-tenuicornis Paratettix-meridionalis Paratettix-mexicanus Tetrix-depressa Tetrix-arenosa Tetrix-bipunctata Tetrix-japonica Paratettix-aztecus Hyperyboella-orphania Scelimena-producta Eurymorphopus-bolivariensis Discotettix-belzebuth Tetrix-ceperoi Tetrix-transsylvanica Arulenus Rhacocleis-buchichii Barbitistes-kaltenbachi Adžić Deranja Pavlović Fran-Rebrina Ivan-Budinski Vrlika HPM Hrvatski-Prirodoslovni-Muzej Zagreb Kitonić Miko Franjević Medak Mathieu Silva Pereira De-Domenico Sperber Connors Hendriksen Lambert Chong McMaster Monaghan Rentz Richter Rose Telnov Barclay Pauwels Entomological-Society-of-Latvia Riga Latvia Wallacea Volume-III biogeography nature-conservation Trinity-Audio guest-blog Josip-Skejo Orthoptera-of-Croatia Balkan-endemic endemic-species natural-history-museum museum-collections field-research career-determination serendipitous-discovery new-species systematic-research understudied-group regular-publishing 2021 2017 2020 2014 2016 2013 2012 2011 primary-school high-school biology-student snakes Uropeltidae Scolecophidia shield-tailed-snakes blind-snakes grasshoppers crickets Croatian-endemic first-encounter water-dependency lifecycle leading-European-orthopterists rare-species new-Arulenus-species understudied publishing Tetrigidae-research encyclopedia Elsevier digital-taxonomy open-access innovations journal growth trending social-media Flickr unidentified-rare-pygmy-grasshoppers students amateurs new-records not-rare Tribulation-helmed-groundhopper Kuranda Tully-Range Redlynch Kingfisher-park Speewah selected-awesome-places selected-amazing-taxa rainforests humid-forests extremely-rare weird-looking largest most-colourful ZooKeys-studies photographs identified-to-species-level variability pronotal-projection-morphology holotype Maroantsentra Antongil-Bay Tamatave Andasibe Vohimana Ranomafana Analamazaotra Traveler's-Palm Ravenala-madagascariensis natural-habitat skilful-flier flight large-back-spines Formidable-Pygmy-Grasshopper Sava-region true-colours small interesting Pygmy-unicorns Southern-America Murici description male-holotype head sternum frontal-view dorsal-view lateral-view sternomentum scale-bars Giraffehoppers unique antennal-shape face-coloration visual-animals courtship mating-pair Yapen-Island Cenderawasih-Bay W-New-Guinea young-entomologists decision study-pygmy-grasshoppers direction primary-and-high-school compare-data first-idea familiar-animals field order-Orthoptera friend fellow-student first-systematic-research two-Croatian-endemic-species first-years 2011-2012 never-saw single-pygmy-grasshopper BIOM Sinj Tetrigidae-location around-water first-time-ever lifecycle-water-dependent researching contacting Hendrik-Devriese Axel-Hochkirch Josef-Tumbrinck Croatian-Natural-History-Museum Skejo-et-al.-2014 Skejo-&-Caballero-2016 regularly-publishing Adžić-et-al.-2021 Endangered-Pygmy-Grasshoppers Encyclopaedia-of-Conservation Mathieu-et-al.-2021 ZooKeys-1042 Silva-et-al.-2017 comments recent-changes Metopomystrum Cleostratini Miriatrini ZooKeys-702 Skejo-et-al.-2020 online-social-media Anaselina Paraselina Selivinga rare-Australian-pygmy-grasshoppers ZooKeys-948 Malagasy-devils north south ZooKeys-957 Tumbrinck-&-Skejo-2017 taxonomic-and-biogeographic-revision New-Guinean-genus Ophiotettix-Walker-1871 Metrodorinae Ophiotettigini-trib.-nov.33-new-species nature's-creations encouraged U.S.-species spectacular closed Halloween orange-and-black Argus-tortoise-beetle Chelymorpha-cassidea oleander-caterpillar Syntomeida-epilais wheel-bug Arilus-cristatus morning-glory Convolvulaceae sweet-potato toxic-compounds alkaloids nerve-poisons monarch-butterfly milkweed-leaf-beetle oleander-aphid predator-protection polka-dot-wasp-moth vermillion-wings iridescent-blue-body white-spots wasp-mimicry ultrasonic-song mate-attraction egg-deposition oleander-leaf caterpillars cluster-feeding cardiac-glycosides heart-poisons dogbane-beetles natural-enemies barrel-shaped-eggs pale-orange exoskeleton-hardening orange-black-orange antennae-pale-orange body-and-legs-jet-black rear-end-brilliant-orange stout-beak proboscis impalement digestive-enzymes liquefaction medieval-torture-wheel Hemiptera true-bugs sucking-mouthparts incomplete-metamorphosis egg-nymph-adult plant-feeders harlequin-bugs squash-bugs stink-bugs fierce-predators pest-control Reduviidae assassin-bugs Zelus-longipes milkweed-assassin-bug sticky-forelegs prey-snaring proteolytic-enzymes predigestion muscular-pump black-wing-buds goldenrods meadow-plants Pselliopus-barberi orange-assassin-bug black-stripes stealth leafhopper brown-marmorated-stink-bug Halyomorpha-halys biological-control-agent native-protein-sources autumn-egg-laying spring-hatching May-June red-abdomens orange-antennae sawfly-larvae beetles other-bugs painful-poke caution handling pet-keeping Mad-Magazine Spy-vs-Spy pointy-nosed-secret-agents entomological-equivalent Bug-vs-Bug Heteroptera clan awesome-predator native-to-North-America record-numbers nursery function-of-wheel speculation cautious-approach embrace liquefied-meal protein-for-growth egg-production cluster-size tree-bark magnificent-creatures wide-variety scores stealthy-stalking assassination complementary-outbreak benefit naturally-occurring-predators parasites stem-onslaught no-interest-in-humans retreat memorable-painful-poke firsthand-experience clever-assassin USDA-NIFA-SCRI-Award research-support action-video website predator-prey-relationship natural-enemy ultimate-comeuppance Chinese-praying-mantis yellow-and-black-garden-spider arachnid close-encounter invasive-pest Asia Allentown-PA 1990s millions-of-dollars-damage crops home-invasion 33-states winter-refuge rising-crescendo concern farm-fields landscapes creature's-bane bounty awesome-six-legged-predator medieval-torture-device source-of-speculation little-fact tasty-morsel mark hapless-victim slurp protein-for-development autumn well-fed-female several-to-more-than-one-hundred sycamore-tree attendance irony nocent triggered-outbreak remain-to-be-seen word-of-caution Assasi pet laboratory learned-firsthand try-not-to-handle victim Shrewsbury Martinson Sargent fascination inspiration archive creatures trees UF extension integrated-pest-management NC-State AG295 html tortoise-beetles IFAS orn ole-cpillar UKY Agriculture CritterFiles casefile insects bugs assassin suggested-common-name scientific-name Linnaeus Megha-Kalsi Dakshina-R.-Seal publication preparation happy-safe-Halloween eat-stink-bug part-3 meets-wheel-bug mid-Atlantic-region inimitable-pest apples vegetables soybeans homes discovery damage distress homeowners wicked-winter previous-episodes give-stink-bugs Baby-boomers outwit undo sucking-insects Mother-Nature liquefied-tissues egg-conversion clusters spring bright-red-abdomens lucky half-dozen stealthily-stalked assassinating stinky-marmorated-cousins no-particular-interest stalking-humans too-large taste-bad firsthand beware handling-directly Ashley Nancy Chris Ryan Erik Caroline wrangling death-of-stink-bug Part-2 vandalizing-vegetables sullying-soybeans invading-homes wide-variety-crops curious-reunion arachnid-kind stalking capturing assassinating-stink-bugs game gorgeous-wheel-bug-nymphs cluster-near-eggs venturing-off first-victim true-bug-clan bed-bugs leaf-footed-bugs nefarious-brown-marmorated-stink-bug predicted struggle protect-crops prevent-stinky-invaders overlook eating-mechanism business-end powerful-beak front-legs cautiously-approaches embraces impales pumps-strong-digestive-enzymes liquefy-body-tissues slurps larvae tens-to-more-than-one-hundred plant-hoppers aphids increase-in-populations stink-bug-laden-nursery curious-irony outbreak beneficial-wheel-bugs iNaturalist-taxon rank preferred-common-name observations-count 399 Wikipedia-summary kingdom phylum class order family genus GBIF-taxonomy-match matched-scientific-name canonical-name status accepted match-type exact distribution-records authorship specific-epithet classification eukaryota nomotettix-cristatus authority basionym group grasshoppers-crickets-katydids species instructions fill-all-fields available-knowledge field-cannot-be-supported return-null NOT-repeat-information each-section-focused useful-detail schema taxon-record entomology-guide accurate conservative informative factual-correctness completeness clarity verbosity usefulness CRITICAL-RULES information-not-clearly-supported NOT-infer-species-level-traits higher-taxa explicitly-justified NOT-repeat-same-information multiple-fields unique-non-overlapping-content vague-generalizations like-most-insects typically-feeds-on-plants cautious-language has-been-observed is-known-to NOT-fabricate behaviors diet life-cycle-details host-relationships FIELD-INTENT summary high-level-overview 3-5-sentences appearance physical-description-only identification distinguish-from-similar-taxa habitat environment-conditions distribution geographic-range-only seasonality timing-of-activity feeding-habits null-if-unknown developmental-stages behavior notable-actions-or-habits ecologicalRole role-in-ecosystem humanRelevance interaction-with-humans similarTaxa must-include-reason misconceptions only-if-meaningful extraDetails important-additional-context STYLE-RULES clear-direct-sentences avoid-fluff-filler-language repeating-taxonomy-in-prose overly-technical-jargon concrete-statements abstract-descriptions QUALITY-RULES completeness-high most-fields-well-supported completeness-medium partial-but-reliable completeness-low sparse-data hasInferredContent true-ONLY-if-generalization-used otherwise-false OUTPUT-FORMAT strictly-match-JSON-schema no-extra-fields no-commentary-outside-JSON water-associated three-subspecies N.-c.-cristatus N.-c.-compressus N.-c.-floridanus small-size 399-observations exact-match medium-completeness no-inferred-content factual-correctness-prioritized conservative-approach informative-content no-fluff no-vague-generalizations cautious-language-where-needed no-fabrication unique-field-content focused-sections JSON-schema-compliance no-commentary
Oarisma poweshiek
Poweshiek Skipperling
The Poweshiek skipperling is a small, critically endangered North American grass skipper butterfly. Historically common across tallgrass prairie systems of the upper Midwest United States and southern Manitoba, Canada, it has experienced catastrophic population declines since 2005. The species now persists at only six verified extant sites: four prairie fens in Michigan, one mesic prairie in Wisconsin, and one tallgrass prairie in Manitoba. Intensive conservation efforts include captive breeding programs at the Minnesota Zoo and Assiniboine Park Conservancy, federal endangered species protection in both countries, and habitat restoration initiatives.
endangered prairie-obligate univoltine grass-skipper captive-breeding-subject indicator-species
Ormosia
Ormosias, horse-eye beans, chocho
Ormosia is a genus of woody legumes (Fabaceae) comprising approximately 130 species of trees and large shrubs distributed across tropical regions of the Americas, Asia, and New Guinea. The genus is characterized by distinctive, often brightly colored seeds with decorative patterns that resemble eyes—commonly called "horse-eye beans"—which are poisonous if ingested but valued for ornamental use. Several species are threatened by habitat destruction, with Ormosia howii likely extinct. The genus forms nitrogen-fixing root nodules with rhizobia and exhibits strong adaptability to nutrient-poor soils.
Fabaceae legume nitrogen-fixation rhizobia-symbiosis ornamental-seeds timber endangered phosphate-solubilization tropical-trees China-endemic-species
Osmia calaminthae
blue calamintha bee
Osmia calaminthae is a rare mason bee endemic to a restricted area of Florida scrub habitat. The species was described in 2011 and is known from only eleven sites concentrated in the southern Lake Wales Ridge and Ocala National Forest. It exhibits striking blue coloration and shows extreme host specialization, depending primarily on two mint-family plants for pollen. The species is considered Critically Imperiled due to its extremely limited range, specific habitat requirements, and ongoing threats from development and pesticide exposure. It was rediscovered in March 2020 after concerns about its continued existence.
bee mason-bee endemic endangered pollinator-specialist Florida-endemic ground-nesting oligolectic Critically-Imperiled host-specific rediscovered-2020
Papaipema eryngii
Rattlesnake-master Borer Moth
Papaipema eryngii is a noctuid moth whose larvae are specialized stem borers of the rattlesnake master (Eryngium yuccifolium). The species has a restricted range in the central United States, tied to the distribution of its host plant. It is the only member of its genus known to have adapted to the defensive chemistry of this particular host. The species is listed as endangered in Illinois and was the subject of a declined federal protection petition in 2020.
endangered specialist-herbivore stem-borer prairie conservation
Papaipema marginidens
brick-red borer moth, Wild Yam and Rattlesnake-master Borer Moths
Papaipema marginidens, known as the brick-red borer moth, is a noctuid moth native to eastern North America. It inhabits open oak woodlands and rocky barrens with herbaceous undergrowth, particularly near streams. The species is listed as endangered in Connecticut, reflecting habitat vulnerability in parts of its range.
endangered eastern-North-America oak-woodland herbaceous-borer streamside
Papilio aristodemus
Schaus' swallowtail, island swallowtail
Papilio aristodemus, commonly known as Schaus' swallowtail or island swallowtail, is a butterfly in the family Papilionidae endemic to southern Florida, the West Indies, and the Bahamas. It is one of North America's most endangered butterflies, with the Florida subspecies (P. a. ponceana, now Heraclides ponceana) listed as endangered since 1984. The species produces a single generation annually with adults active primarily in May and June. Habitat loss from urban development, hurricanes, and mosquito control spraying has caused dramatic population declines, including near-extinction from Hurricane Andrew in 1992.
endangered flagship-species tropical-hardwood-hammock single-generation captivity-breeding West-Indies Florida-endemic Rutaceae-specialist territorial-behavior cross-water-dispersal
Parvindela celeripes
Swift Tiger Beetle
Parvindela celeripes, the Swift Tiger Beetle, is a tiny (6–8 mm), flightless tiger beetle species native to the Great Plains of North America. Once abundant in native grasslands of Iowa, Nebraska, and Kansas during the late 1800s and early 1900s, the species has experienced severe population declines due to habitat loss. It was unknown from Missouri until its discovery in 2010, where it persists in small, isolated loess hilltop prairie remnants. The species remains robust in northwestern Oklahoma's red clay/gypsum exposures, representing its most secure population. Its flightless nature makes recolonization of disturbed or burned sites extremely difficult.
tiger-beetle flightless prairie-specialist conservation-concern loess-hills gypsum endangered Great-Plains Cicindelidae
Pentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetles Dynastinae Scarabaeidae Coleoptera tribe global-distribution morphological-diversity sexual-dimorphism generic-diversity mouthpart-morphology secondary-sexual-characters horns biogeographic-restriction taxonomic-revision dichotomous-keys new-species-description new-genus-description lectotype-designation synonymy new-combination distribution-mapping female-description habitat-data behavioral-observations Australia Colombia Bolivia India Western-Australia New-South-Wales Neotropical Australian Afrotropical Oriental Palaearctic Cheiroplatina Dipelicina Pentodontina Pseudoryctina Bothynus Heteronychus Epironastes Philcarneum Constricticollis Carneiola Anomalomorpha Enracius Erbmahcedius Cavonus Pericoptus Pentodon Calicnemis Metanastes Neometanastes Pimelopus Podalgus Pseudoryctes Cheiroplatys Dipelicus Denhezia Euetheola Hylobothynus Oxyligyrus Parapucaya Pucaya Tomarus Adoryphorus Carneoryctes Teinogenys Ligyrus Allsopp Hutchinson Arrow Carne Endrödi Dechambre Prell Ohaus Bates Hope Laporte-de-Castelnau Erichson Burmeister Sharp Mulsant Blackburn Dupuis Özdikmen Yamaya Fairmaire Redtenbacher Steinheil Ratcliffe Cave Fabricius Dejean iNaturalist Wikipedia Catalogue-of-Life Zootaxa Journal-of-Insect-Biodiversity Records-of-the-Zoological-Survey-of-India The-Coleopterists-Bulletin BioLib.cz Wikimedia-Commons DOI 10.11646/zootaxa.4048.4.1 10.11646/zootaxa.4852.4.2 10.11646/zootaxa.5351.3.2 10.26515/rzsi/v125/i2s/2025/172964 10.11646/zootaxa.5716.4.7 10.11646/zootaxa.5072.5.2 10.11646/zootaxa.4852.4.3 10.12976/jib/2024.54.2.2 10.1649/1186.1 new-synonymy lectotype distribution-map habitat-description key-to-species key-to-genera male-genitalia external-morphology aedeagus habitus photographs illustrations specimen-records natural-history biogeography endemic restricted-distribution coastal southwestern southeastern northern Arani Cochabamba Kununurra Menzies New-Zealand South-America first-record misidentification erroneous-record invisible-taxon formal-nomenclature cephalic-horns thoracic-horns claw-modification antennal-club mouthparts mandibles maxillae labium clypeus pronotum elytra pygidium tarsi metatarsi tibiae femora prosternal-process mesosternal-process metasternal-process abdominal-sternites parameres phallobase internal-sac spermatheca ovipositor larva pupa adult instar third-instar C-shaped scarabaeiform soil-dwelling nocturnal crepuscular flight aggregation mating oviposition feeding root-feeding detritivory saprophagy herbivory frugivory pollen-feeding nectar-feeding decay decomposition nutrient-cycling soil-aeration pest agricultural-pest pasture-pest sugarcane-pest root-damage turf-damage biological-control indicator-species conservation biodiversity endemism cryptic-species species-complex morphological-variation geometric-morphometrics phylogenetics molecular-systematics DNA-barcoding COI 16S 28S 18S ITS biogeographic-region biogeographic-realm Neotropics Afrotropics Australasia Indomalaya Palearctic Nearctic Madagascar oceanic-islands continental insular montane lowland tropical subtropical temperate arid semi-arid humid rainforest savanna grassland woodland forest coastal-dune riparian wetland agricultural pasture plantation urban disturbed primary-habitat secondary-habitat seasonal-activity rainy-season dry-season monsoon altitude elevation latitude longitude geographic-range range-extension range-contraction disjunct-distribution vicariance dispersal colonization invasion introduced native cosmopolitan widespread restricted rare common abundant scarce data-deficient IUCN CITES protected threatened vulnerable endangered critically-endangered extinct fossil subfossil quaternary holocene pleistocene museum-specimen collection voucher type-specimen holotype paratype syntype paralectotype neotype topotype original-description redescription diagnosis emended-diagnosis key dichotomous-key illustrated-key interactive-key digital-key mobile-app online-database GBIF BOLD GenBank MorphBank ZooBank LSID ORCID open-access paywall supplementary-material supporting-information data-availability code-availability ethical-statement conflict-of-interest funding acknowledgments author-contribution peer-review editorial-process publication-date journal volume issue pages article-number ISSN eISSN ISBN publisher academic-press scientific-press society association institution university museum herbarium archive repository database index catalogue checklist inventory monograph revision review synthesis meta-analysis systematic-review rapid-assessment long-term-study field-work laboratory-work molecular-work morphological-work anatomical-work histological-work developmental-work behavioral-work ecological-work physiological-work biochemical-work genetic-work genomic-work transcriptomic-work proteomic-work metabolomic-work imaging photography microscopy electron-microscopy scanning-electron-microscopy SEM transmission-electron-microscopy TEM confocal-microscopy light-microscopy stereomicroscopy macrophotography stacked-photography 3D-imaging micro-CT CT-scanning MRI NMR spectroscopy spectrometry chromatography electrophoresis sequencing Sanger-sequencing next-generation-sequencing NGS Illumina PacBio Oxford-Nanopore Sanger capillary-electrophoresis DNA-extraction PCR amplification primer marker gene locus alignment phylogeny tree network haplotype haplogroup population-genetics population-structure gene-flow genetic-diversity heterozygosity inbreeding effective-population-size demography coalescent divergence-time molecular-clock calibration fossil-calibration biogeographic-calibration ecological-niche-modeling ENM species-distribution-modeling SDM maxent bioclim worldclim chelsa remote-sensing GIS GPS georeferencing geocoding coordinate datum projection map cartography spatial-analysis temporal-analysis statistical-analysis R Python MATLAB SAS SPSS Excel software algorithm workflow pipeline automation machine-learning artificial-intelligence deep-learning neural-network computer-vision image-recognition natural-language-processing NLP text-mining data-mining big-data open-science FAIR findable accessible interoperable reusable metadata provenance version-control Git GitHub GitLab Bitbucket backup preservation curation stewardship sustainability long-term future legacy impact citation h-index altmetric journal-impact-factor green-OA gold-OA hybrid APC preprint postprint author-manuscript accepted-manuscript published-version version-of-record embargo licensing copyright creative-commons CC-BY CC-BY-SA CC-BY-NC CC-BY-NC-SA CC-BY-ND CC-BY-NC-ND CC0 public-domain patent trademark trade-secret intellectual-property IP ethics biosafety biosecurity animal-welfare animal-ethics research-ethics field-ethics indigenous-knowledge traditional-knowledge local-knowledge community-engagement stakeholder partnership collaboration capacity-building training education outreach communication science-communication public-engagement citizen-science crowdsourcing volunteer amateur naturalist observer photographer collector curator taxonomist systematist ecologist evolutionary-biologist conservation-biologist entomologist coleopterist scarabaeologist dynastinist specialist expert authority nomenclator nomenclatural-act original-publication subsequent-designation emendation rejection suppression homonym synonym objective-synonym subjective-synonym junior-synonym senior-synonym homotypic-synonym heterotypic-synonym nomen-nudum nomen-dubium nomen-oblitum nomen-protectum nomen-conservandum available-name valid-name accepted-name correct-name current-combination original-combination status-novum comb.-nov.syn.-nov.sp.-nov.gen.-nov.subgen.-nov.subsp.-nov.nom.-nov.nom.-nud.nom.-dub.incertae-sedis unplaced unassigned subtribe genus subgenus species subspecies variety form morph ecotype population stock strain breed cultivar clone lineage biotype karyotype chromosome genome transcriptome proteome metabolome phenome morphome anatomical morphological meristic morphometric allometric ontogenetic developmental life-history reproductive behavioral ecological physiological biochemical genetic genomic evolutionary phylogenetic phylogeographic biogeographic historical paleontological archaeological geological climatological environmental applied economic medical veterinary forestry horticultural industrial pollution climate-change global-warming habitat-loss fragmentation deforestation agricultural-intensification urbanization invasive-species pathogen parasite predator competition mutualism symbiosis commensalism amensalism neutralism facilitation inhibition trophic-cascade food-web food-chain energy-flow ecosystem-function ecosystem-service provisioning regulating supporting cultural alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance rarefaction extrapolation asymptotic non-asymptotic sample sampling survey monitoring assessment evaluation indicator bioindicator flagship umbrella keystone ecosystem-engineer foundation-species dominant subordinate frequent occasional accidental vagrant migrant resident breeding non-breeding wintering summering migration nomadism irruption establishment naturalization spreading range-expansion range-shift extirpation extinction local-extinction global-extinction functional-extinction ecological-extinction pseudextinction Lazarus-taxon Elvis-taxon zombie-taxon living-fossil relict indigenous autochthonous allochthonous exotic alien invasive naturalized casual escaped cultivated ornamental pet food medicine fiber fuel timber fodder pollinator pest-control seed-dispersal soil-formation erosion-control water-purification air-purification carbon-sequestration climate-regulation disease-regulation pest-regulation flood-regulation storm-protection recreation tourism aesthetic spiritual cultural-heritage research inspiration existence-value option-value bequest-value intrinsic-value anthropocentric biocentric ecocentric utilitarian instrumental relational intrinsic inherent absolute conditional responsibility care respect reverence wonder curiosity knowledge understanding wisdom science natural-philosophy biology zoology entomology coleopterology scarabaeology dynastinology systematics taxonomy nomenclature classification evolution ecology behavior paleontology conservation-biology environmental-science agricultural-science forestry-science veterinary-science medical-science public-health one-health planetary-health ecosystem-health biodiversity-health species-health population-health individual-health genetic-health environmental-health social-health economic-health political-health cultural-health spiritual-health holistic-health integrated-health sustainable-development sustainable-use sustainable-management adaptive-management precautionary-principle ecosystem-approach landscape-approach seascape-approach connectivity-conservation corridor buffer-zone protected-area national-park nature-reserve wildlife-refuge wilderness-area world-heritage-site biosphere-reserve Ramsar-site Important-Bird-Area Key-Biodiversity-Area Alliance-for-Zero-Extinction-site conservation-priority hotspot crisis-ecoregion global-200 last-of-the-wild human-footprint cumulative-impact threat-index vulnerability-index adaptive-capacity exposure sensitivity resilience resistance recovery restoration rehabilitation reintroduction translocation ex-situ in-situ captive-breeding botanic-garden zoo gene-bank seed-bank tissue-bank sperm-bank oocyte-bank embryo-bank DNA-bank frozen-zoo ark insurance safety-net de-extinction genetic-rescue genetic-restoration genetic-augmentation genetic-management population-management metapopulation source-sink patch matrix landscape seascape ecosystem biome ecoregion province zone region district site locality habitat microhabitat niche ecological-niche fundamental-niche realized-niche trophic-niche spatial-niche temporal-niche biotic-niche abiotic-niche multidimensional-niche n-dimensional-niche Hutchinsonian-niche Grinnellian-niche Eltonian-niche resource requirement limitation stress disturbance perturbation fluctuation variability heterogeneity complexity diversity redundancy stability persistence adaptation acclimation plasticity evolvability heritability selection drift flow mutation recombination speciation coalescence divergence convergence parallelism homoplasy analogy homology synapomorphy symplesiomorphy autapomorphy apomorphy plesiomorphy derived ancestral primitive advanced basal crown stem node branch clade grade subfamily family superfamily infraorder suborder order superorder infraclass subclass class superclass subphylum phylum superphylum kingdom domain life organism individual etc.
Phalangodidae
Phalangodid Harvestmen
Phalangodidae is a family of small harvestmen in the suborder Laniatores, containing approximately 30 genera and more than 100 described species. Members are characterized by pedipalps armed with prominent spines and body lengths generally under 3 mm. The family is primarily distributed in the Holarctic region, with exceptional diversity in the western Nearctic, particularly California. Several species, especially in the genus Texella, are obligate cave-dwellers exhibiting troglomorphic traits such as depigmentation and reduced eyes.
harvestmen Opiliones Laniatores cave-dwelling troglobitic endangered karst Holarctic California Texas
Phyllonorycter ledella
Labrador Tea Leaf Miner
Phyllonorycter ledella is a micro-moth in the family Gracillariidae, commonly known as the Labrador Tea Leaf Miner. It is a small leaf-mining species with a wingspan of 9–10 mm. The species has a disjunct distribution across North America and is listed as endangered in Connecticut.
micro-moth leaf-miner monophagous endangered North-America
Plebejus icarioides missionensis
Plebejus icarioides missionensis is a subspecies of the Mission blue butterfly, a federally endangered lycaenid butterfly endemic to a restricted range in coastal California. The subspecies is part of a complex that has been subject to taxonomic revision, with some authorities placing it in the genus Icaricia. Its habitat consists of coastal grasslands and scrub where its larval host plant, silver lupine (Lupinus albicaulis), occurs. The subspecies has been impacted by urban development, invasive plants, and fire suppression.
endangered conservation California-endemic lycaenidae host-plant-specialist urban-wildlife prescribed-fire-management
Plebejus melissa samuelis
Karner blue butterfly
Plebejus melissa samuelis, commonly known as the Karner blue butterfly, is a federally endangered subspecies first described by Vladimir Nabokov in 1944 from Karner, New York. It is highly specialized, with larvae feeding exclusively on wild lupine (Lupinus perennis) and inhabiting dry sandy barrens ecosystems. The subspecies produces two generations annually and has become a flagship for conservation efforts, particularly in Wisconsin where a pioneering statewide Habitat Conservation Plan was developed.
endangered conservation pollinator barrens-ecosystem habitat-specialist flagship-species Nabokov
Pogonomyrmex comanche
Comanche harvester ant
Pogonomyrmex comanche, the Comanche harvester ant, is an endangered harvester ant species endemic to the western Gulf Coastal Plain of North America. Genetic studies indicate central Texas serves as the center of genetic diversity, consistent with Pleistocene refugia patterns observed in many regional species. Northern populations in Arkansas and Oklahoma exhibit lower genetic diversity and evidence of population contraction rather than recent range expansion. The species shows signs of dispersal limitation, with high genetic diversity and few identical sequences among sampled individuals. Its decline appears driven by a combination of habitat loss and limited dispersal capacity, though exact mechanisms remain under investigation.
endangered harvester-ant Pogonomyrmex western-Gulf-Coastal-Plain phylogeography Pleistocene-refugia dispersal-limitation conservation Formicidae North-America Texas Oklahoma Arkansas sandy-soils prairie savanna
Polites mardon
Mardon skipper
Polites mardon, the Mardon skipper, is a small butterfly in the family Hesperiidae endemic to the Pacific Northwest of the United States. It inhabits native grassland prairies, where it has been observed using native fescue grasses as oviposition sites. The species is listed as endangered in Washington state and was petitioned for federal endangered species listing in 2002. Conservation efforts initiated by the Xerces Society have focused on habitat protection and research into its ecological requirements.
endangered Pacific-Northwest-endemic prairie-specialist conservation-flagship non-migratory
Polyphylla barbata
Mount Hermon June Beetle
Polyphylla barbata is a federally endangered scarab beetle endemic to a single restricted locality in the Zayante sandhills of Santa Cruz County, California. The species occupies less than 1,500 acres of habitat, making it one of the most geographically limited beetles in North America. Adults are active in mid-summer, with males flying at dusk to locate flightless females using pheromone-detecting antennae. The species faces severe habitat loss from sand mining and development, with over 40% of its sandhills habitat already destroyed.
endangered endemic sand-dune California Scarabaeidae Melolonthinae Zayante-sandhills nocturnal flightless-female pheromone-mediated-mating
Pseudocopaeodes eunus
alkali skipper, Carson wandering skipper (for ssp. obscurus)
Pseudocopaeodes eunus is a rare skipper butterfly endemic to alkali flat habitats in the western United States and Mexico. The species comprises five recognized subspecies, with P. e. obscurus (Carson wandering skipper) federally listed as endangered due to habitat loss and extremely limited distribution. As of 2007, only four populations were known to persist. The species is highly specialized, with larvae dependent on saltgrass as their sole documented host plant.
alkali-flats endangered saltgrass specialist western-North-America wetland-dependent
Pycnomerus
ironclad beetles
Pycnomerus is a genus of ironclad beetles (family Zopheridae) comprising more than 20 described species. Members are small, elongated beetles with hardened, often sculptured exoskeletons typical of the family. The genus includes both widespread and highly restricted species, with some island endemics now considered extinct due to habitat loss. Several species are associated with dead wood habitats and are classified as saproxylic.
ironclad-beetles saproxylic endangered introduced-species extinct forest-dependent Zopheridae Coleoptera dead-wood-habitat conservation-concern
Pyractomena ecostata
Keel-necked firefly
Pyractomena ecostata, the keel-necked firefly, is an endangered firefly species endemic to the United States with a highly disjunct distribution. It occurs in two widely separated coastal regions: the Mobile Bay area of Alabama east through peninsular Florida, and saltmarshes surrounding Delaware Bay and Cape May in Delaware and New Jersey. This species is restricted to brackish habitats, particularly those receiving saltwater intrusion, and faces severe threats from coastal development, sea level rise, invasive vegetation, and light pollution.
endangered firefly coastal salt-marsh brackish disjunct-distribution climate-change-vulnerable
Pyrgus centaureae wyandot
Appalachian Grizzled Skipper
Pyrgus centaureae wyandot is a subspecies of northern grizzled skipper endemic to the Appalachian Highlands and Northern Michigan. It is a small butterfly with distinctive checkered wing patterns and low, erratic flight. The subspecies is listed as federally threatened and state endangered in Ohio and New Jersey due to habitat loss and pesticide exposure. Populations have declined significantly from insecticide spraying targeting spongy moth outbreaks.
skipper butterfly Appalachian-endemic endangered univoltine disturbance-dependent
Rhadine
Rhadine is a genus of ground beetles (Carabidae: Platyninae) described by LeConte in 1846. Many species are troglobitic, restricted to cave habitats in North America, particularly in Texas. Several species, including R. exilis and R. infernalis, are federally listed as endangered due to their restricted ranges and vulnerability to habitat disturbance. The genus is named after a figure from Greek mythology.
cave-beetle troglobite endangered Texas karst ground-beetle Platyninae
Rhaphiomidas
flower-loving flies
Rhaphiomidas is a genus of mydid flies comprising fewer than 30 species and subspecies restricted to desert regions of the southwestern United States and northwestern Mexico. Adults are active for brief periods in spring or fall, with multiple species in the same dune system often separated by seasonal timing. Several taxa are critically endangered due to habitat loss from development and recreational disturbance.
desert sand-dune endangered mydidae allochronic ant-association habitat-specialist
Satyrium edwardsii
Edwards' Hairstreak
Satyrium edwardsii, the Edwards' Hairstreak, is a butterfly in the family Lycaenidae. It occurs in eastern North America from the United States into southern Canada. Larvae feed on Quercus species, particularly scrub oak (Q. ilicifolia). The species has been documented as endangered in some regions and exhibits myrmecophilous associations with ants.
butterfly lycaenid hairstreak oak-associated myrmecophilous endangered North-America
Schinia bimatris
White Flower Moth
Schinia bimatris, commonly known as the White Flower Moth, is a small noctuid moth species found in the south-central and south-eastern United States and in a disjunct population in Manitoba, Canada. The species is designated as Endangered under Canada's Species At Risk Act. Adults are distinctive in their entirely white wings, thorax, and abdomen with an orange head, and have a wingspan of approximately 30 mm. The larval stage remains undescribed, and the host plant relationship is poorly understood, though Nuttall's Evening-primrose (Oenothera nuttallii) has been suggested as a possible food source.
endangered conservation-concern habitat-specialist undescribed-larva disjunct-distribution
Schinia gracilenta
Slender Flower Moth, Iva Flower Moth
Schinia gracilenta is a small noctuid moth native to North America, first described by Jacob Hübner in 1818. The species has a wingspan of approximately 28 mm and produces one generation annually. It is listed as endangered in Connecticut, reflecting localized population concerns in parts of its range.
moth Noctuidae flower-moth North-America prairie specialist-herbivore endangered Iva Asteraceae
Somatochlora hineana
Hine's Emerald, Hine's Emerald Dragonfly
Somatochlora hineana is a rare dragonfly species endemic to North America, listed as endangered in Canada since 2011 and of conservation concern in the United States. The species is highly specialized, requiring groundwater discharge areas in wetland habitats to complete its life cycle. It exhibits habitat segregation between sexes and has been documented with biased sex ratios, factors that complicate conservation efforts. The Canadian population is restricted to the Minesing Wetlands in Ontario, while U.S. populations occur in Michigan and other states with suitable calcareous wetland habitats.
endangered groundwater-dependent fen-specialist conservation sex-ratio-bias habitat-segregation calcareous-wetland Somatochlora Corduliidae Odonata Canada Ontario Michigan wetland-ecology hydrology marl-deposits rich-fen
Strymon acis
Bartram's scrub-hairstreak, Caribbean Scrub-Hairstreak
Strymon acis is a small lycaenid butterfly with a wingspan of 22–29 mm, occurring in southern Florida and the West Indies. The subspecies S. a. bartrami, restricted to southern Florida, is federally listed as Endangered under the United States Endangered Species Act. The species is tightly associated with pine rockland habitats and depends on Croton linearis as its larval host plant.
endangered pine-rockland fire-dependent-ecosystem specialist-herbivore Florida-endemic-subspecies
Stygobromus pecki
Peck's cave amphipod
Stygobromus pecki is a small, eyeless, unpigmented cave-dwelling amphipod endemic to four spring systems in Comal County, Texas. It is a federally listed endangered species in the United States and classified as Endangered on the IUCN Red List due to its extremely limited geographic distribution. The species inhabits subterranean limestone aquifers and exhibits adaptations typical of stygobitic organisms, including light sensitivity and starvation resistance. Very few individuals have been documented since its listing, and no formal recovery plan or comprehensive population assessment exists as of 2022.
stygobite cave-dwelling endangered aquifer Texas-endemic Edwards-Aquifer crustacean amphipod
Stygoparnus
Comal Springs dryopid beetle (for sole species S. comalensis)
Stygoparnus is a monotypic genus of dryopid beetles containing the single species Stygoparnus comalensis, known as the Comal Springs dryopid beetle. The genus is endemic to central Texas spring systems fed by the Edwards Aquifer. Its sole member is a federally endangered, subterranean-obligate aquatic beetle with vestigial eyes and reduced pigmentation. The genus represents the only known aquatic lineage within the family Dryopidae.
endangered stygobitic subterranean aquatic endemic Edwards-Aquifer Texas monotypic Dryopidae cave-adapted phototaxis conservation captive-breeding
Stygoparnus comalensis
Comal Springs dryopid beetle
Stygoparnus comalensis is a federally endangered, subterranean-obligate dryopid beetle endemic to three spring systems in the Edwards Aquifer region of central Texas. Adults are approximately 2 mm long with vestigial eyes and undeveloped wings, reflecting their troglobitic ancestry. The species exhibits extreme population genetic structure with no gene flow among localities, indicating evolutionary isolation. Captive studies reveal strong habitat preferences for wood substrates and specific phototactic responses to light wavelength. The beetle serves as a surrogate species for monitoring aquifer health and faces threats from groundwater extraction, pollution, and drought-induced springflow reduction.
endangered subterranean aquatic spring-endemic Edwards-Aquifer troglobitic conservation
Texamaurops
Texamaurops is a genus of small rove beetles (Staphylinidae: Pselaphinae) established by Barr & Steeves in 1963. The genus contains at least one described species, Texamaurops reddelli, commonly known as the Kretschmarr Cave mold beetle. Members of this genus are associated with cave environments and belong to the diverse tribe Batrisini within the ant-loving beetle subfamily Pselaphinae.
cave-beetle subterranean endangered Texas-endemic Pselaphinae Staphylinidae rove-beetle
Texella
cave harvestmen, Texella harvesters
Texella is a genus of armored harvestmen (Opiliones: Phalangodidae) containing more than 20 described species. The genus is endemic to cave and karst systems of central Texas, where species exhibit varying degrees of troglomorphy. Several species, including T. reyesi and T. reddelli, are federally endangered due to extreme habitat restriction. Genetic studies reveal strong population structuring aligned with geologic karst fauna regions.
cave-dwelling troglomorphic endemic endangered karst central-Texas conservation-genetics
Thermosphaeroma
Thermosphaeroma is a genus of isopod crustaceans in the family Sphaeromatidae, endemic to thermal springs of the southwestern United States and central Mexico. The genus contains nine described species, most of which are critically endangered or extinct in the wild due to groundwater extraction and habitat alteration. These isopods exhibit specialized adaptations to thermo-mineral spring environments, including temperature-dependent life histories and pronounced sexual dimorphism in uropod morphology. Several species have been studied for their complex social behaviors, including precopulatory mate-guarding and cannibalism.
isopod crustacean thermal-spring endemic endangered cannibalism sexual-conflict conservation
Thermosphaeroma thermophilum
Socorro isopod, Socorro sowbug
Thermosphaeroma thermophilum is a small, thermophilic isopod endemic to thermal spring habitats in Socorro County, New Mexico. It is one of the most endangered crustaceans in North America, having been driven to extinction in the wild in 1988 when a tree root burst the water pipeline that formed its only remaining habitat following spring diversion in 1947. The species persists through captive breeding programs and has been reintroduced to artificial habitats. It exhibits pronounced sexual dimorphism in body size and complex social behaviors including cannibalism and mate-guarding.
endemic endangered thermal-spring cannibalism sexual-dimorphism captive-breeding first-ESA-listed-crustacean
Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
Ephemeroptera Polymitarcyidae burrowing-mayflies Neotropical clay-bank-habitat U-shaped-tunnels mandibular-tusks Tortopsis Campsurinae Panamerican-distribution cladistic-revision synapomorphy parastyli-receptors flattened-penes furcasternal-protuberances subapical-tubercles limnological-sampling-bias nymph-morphology adult-morphology type-species-T.-igaranus new-species-T.-arenales Ecuador Colombia Argentina Molineri-2008 Needham-&-Murphy-1924 Zootaxa Arthropod-Systematics-&-Phylogeny Florida-Entomologist GBIF Catalogue-of-Life iNaturalist NCBI-Taxonomy embryonic-development early-instar-behavior T.-harrisi T.-bellus T.-circumfluus T.-zottai T.-igaranus T.-incertus T.-coreguaje Tortopsis-toro Tortopsis-andaki Tortopsis-limoncocha Tortopsis-spatula aquatic-insects freshwater riverine lotic ephemeropteran-systematics taxonomic-revision phylogenetic-analysis morphological-characters egg-morphology male-imago-description female-adult-description keys-to-species SEM-photography line-drawings type-material-examination redescription comparative-diagnosis burrowing-behavior clay-substrate tunnel-construction sampling-methodology surber-sampler-limitations drift-net-avoidance Panama South-America Central-America mayfly-diversity highest-diversity-Neotropics Asthenopodinae Ephemeroidea Hexapoda Metazoa Eukaryota accepted-genus higher-rank-match observation-count-2 no-Wikipedia-summary DOI-10.11646/zootaxa.2481.1.1 DOI-10.11646/zootaxa.2436.1.4 DOI-10.3897/asp.79.e62735 DOI-10.2307/3493493 Scott-et-al.-1959 Navás Ulmer Traver Lugo-Ortiz-&-McCafferty Domínguez Banks McDunnough Pictet Needham-&-Murphy freshwater-ecology aquatic-entomology insect-development nymphal-stages imago-description alate-adults wing-venation R-sector gonopore-structure parastyli-length pedestal-morphology lateral-view-curvature mandibular-morphology tusk-tubercles mesosternal-structure abdominal-sternum-notch furcasternum protuberance-configuration receptor-socket-orientation IR-vein R2-vein claw-like-structure gonopore-association penis-base-separation female-fore-wing additional-veins comparative-morphology type-specimen holotype allotype paratype neotropical-species new-species-description species-key identification-key male-adult-key female-adult-key nymph-key generic-key Polymitarcyidae-genera systematics phylogeny cladistics monophyly sister-group character-state transformation-series parsimony-analysis morphological-data-set external-morphology egg-chorion micropyle egg-burster hatching emergence behavioral-study early-instar embryology oviposition substrate-preference habitat-specificity endemic regional-diversity conservation biomonitoring water-quality-indicator macroinvertebrate benthic infaunal interstitial hyporheic river-bank stream-bank erosion sediment particle-size clay-content bank-stability root-mat vegetation-association flow-regime current-velocity depth width stream-order catchment basin ecoregion biogeography dispersal colonization vicariance speciation adaptive-radiation morphological-divergence convergent-evolution homoplasy character-conflict bootstrap-support Bremer-support consensus-tree strict-consensus majority-rule taxonomic-stability nomenclatural-act original-description subsequent-designation type-locality type-depository museum-collection specimen-preservation alcohol-preservation slide-mounting SEM-preparation imaging-techniques scientific-illustration photography measurement morphometrics ratios proportions allometry sexual-dimorphism geographic-variation intraspecific-variation interspecific-variation diagnostic-characters identification-confidence taxonomic-expertise literature-review bibliographic-search database-query online-repository open-access paywall abstract-only full-text-unavailable citation-analysis reference-list source-quality evidence-strength knowledge-gap research-need future-study collaboration field-work collection-trip sampling-effort specimen-availability type-examination revision-scope taxonomic-concept circumscription inclusion exclusion species-group species-complex cryptic-species morphospecies phylogenetic-species evolutionary-significant-unit operational-taxonomic-unit DNA-barcoding molecular-systematics integrative-taxonomy traditional-taxonomy alpha-taxonomy beta-taxonomy gamma-taxonomy classification-hierarchy Linnaean-system rank-based-nomenclature phylogenetic-nomenclature clade-name crown-group stem-group total-evidence congruence incongruence data-partition character-weighting implied-weighting equal-weighting ordered-characters unordered-characters multistate-characters binary-characters continuous-characters meristic-characters qualitative-characters quantitative-characters morphological-matrix data-matrix character-matrix taxon-sampling character-sampling missing-data inapplicable-data polymorphism intraspecific-polymorphism fixed-difference diagnostic-difference unique-derived-character autapomorphy shared-derived-character symplesiomorphy convergent-character parallel-evolution reversal character-optimization ACCTRAN DELTRAN unambiguous-optimization ambiguous-optimization most-parsimonious-tree tree-length consistency-index retention-index rescaled-consistency-index homoplasy-index character-fit character-congruence character-support branch-support node-support clade-stability taxonomic-stability-index phylogenetic-signal noise saturation substitution-model rate-heterogeneity among-site-rate-variation gamma-distribution invariant-sites model-selection Akaike-information-criterion Bayesian-information-criterion likelihood-ratio-test Bayesian-inference Markov-chain-Monte-Carlo posterior-probability prior-probability hyperprior convergence effective-sample-size burn-in stationarity trace-plot parameter-estimate credible-interval highest-posterior-density tree-topology tree-shape lineage-through-time diversification-rate extinction-rate speciation-rate net-diversification birth-death-model Yule-model coalescent-model population-size effective-population-size gene-flow migration-rate isolation-by-distance phylogeography historical-biogeography ancestral-area-reconstruction dispersal-vicariance-analysis decisive-weighting event-based-method taxon-area-cladogram area-cladogram biogeographic-homology track-analysis panbiogeography phylogenetic-biogeography ecological-biogeography dispersal-biogeography vicariance-biogeography center-of-origin area-of-endemism refugium Pleistocene glaciation climate-change habitat-fragmentation habitat-loss anthropogenic-impact pollution eutrophication sedimentation dam-construction water-abstraction flow-modification thermal-pollution toxic-substances heavy-metals pesticides urbanization agriculture deforestation land-use-change riparian-degradation channelization dredging mining invasive-species biological-invasion competition predation parasitism mutualism commensalism amensalism facilitation ecosystem-engineer keystone-species flagship-species umbrella-species indicator-species sentinel-species bioindicator ecological-indicator environmental-indicator water-quality-index biotic-index multimetric-index rapid-bioassessment protocol standardization quality-assurance quality-control reference-condition least-disturbed minimally-disturbed best-available type-specific region-specific stream-type river-type functional-feeding-group collector-gatherer collector-filterer scraper shredder predator parasite engulfer piercer burrower sprawler clinger swimmer skater diver surface-dweller benthic-dweller interstitial-dweller hyporheic-dweller phreatic-dweller troglobite stygobite cave-inhabitant spring-inhabitant headwater mid-order high-order lowland montane alpine subtropical tropical temperate boreal arctic antarctic palearctic nearctic afrotropical oriental australian oceanic cosmopolitan widespread restricted relict disjunct-distribution range-extension new-record first-record country-record state-record province-record locality-record elevation-record altitudinal-range latitudinal-range longitudinal-range distribution-pattern range-size range-fill extent-of-occurrence area-of-occupancy IUCN-category threat-status data-deficient not-evaluated least-concern near-threatened vulnerable endangered critically-endangered extinct-in-wild extinct population-trend population-density population-structure age-structure size-structure sex-ratio reproductive-output fecundity fertility mortality survivorship life-table cohort-analysis generation-time doubling-time intrinsic-rate-of-increase finite-rate-of-increase net-reproductive-rate stable-age-distribution stationary-population expanding-population declining-population source-sink-dynamics metapopulation population-viability-analysis minimum-viable-population genetic-diversity heterozygosity allelic-richness private-allele founder-effect bottleneck genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression hybridization introgression gene-flow-barrier reproductive-isolation prezygotic-isolation postzygotic-isolation mechanical-isolation temporal-isolation habitat-isolation behavioral-isolation gametic-isolation zygotic-mortality hybrid-inviability hybrid-sterility hybrid-breakdown reinforcement character-displacement ecological-speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation centrifugal-speciation divergence-with-gene-flow speciation-continuum anagenesis cladogenesis budding-speciation branching-speciation phyletic-gradualism punctuated-equilibrium stasis trend progression orthogenesis aristogenesis mosaic-evolution co-evolution arms-race Red-Queen-hypothesis evolutionary-stable-strategy frequency-dependent-selection density-dependent-selection r-selection K-selection bet-hedging iteroparity semelparity ploidy-level chromosome-number karyotype genome-size transposable-element gene-duplication neofunctionalization subfunctionalization pseudogene regulatory-evolution cis-regulatory trans-regulatory enhancer promoter silencer insulator transcription-factor signaling-pathway developmental-pathway gene-network epigenetic-modification DNA-methylation histone-modification chromatin-remodeling genomic-imprinting parental-effect maternal-effect paternal-effect cytoplasmic-inheritance mitochondrial-DNA chloroplast-DNA endosymbiont Wolbachia Rickettsia Spiroplasma Cardinium Arsenophonus Hamiltonella Regiella Serratia Buchnera Carsonella Portiera primary-endosymbiont secondary-endosymbiont facultative-symbiont obligate-symbiont mutualistic-symbiont parasitic-symbiont commensal-symbiont pathogen vector disease-transmission zoonosis emerging-infectious-disease biosecurity quarantine eradication control management conservation-biology restoration-ecology habitat-restoration river-restoration stream-restoration floodplain-restoration riparian-restoration wetland-restoration ecosystem-restoration passive-restoration active-restoration assisted-recovery managed-relocation assisted-colonization ex-situ-conservation in-situ-conservation protected-area national-park nature-reserve wildlife-refuge biosphere-reserve world-heritage-site ramsar-site important-bird-area key-biodiversity-area Alliance-for-Zero-Extinction EDGE-species evolutionarily-distinct globally-endangered conservation-prioritization systematic-conservation-planning Marxan Zonation conservation-opportunity conservation-cost conservation-benefit trade-off socioeconomic-factor stakeholder-engagement community-based-conservation adaptive-management monitoring-program long-term-study citizen-science public-participation environmental-education outreach science-communication knowledge-transfer capacity-building institutional-strengthening policy-development legislation regulation enforcement compliance incentive payment-for-ecosystem-services carbon-credit biodiversity-offset mitigation-banking conservation-easement land-trust private-land-conservation conservation-covenant stewardship sustainable-use sustainable-management ecosystem-based-management integrated-water-resources-management basin-management catchment-management landscape-management seascape-management connectivity-conservation corridor stepping-stone buffer-zone matrix-management countryside-biogeography reconciliation-ecology novel-ecosystem designer-ecosystem ecosystem-service provisioning-service regulating-service supporting-service cultural-service food-production water-supply raw-materials genetic-resources medicinal-resources ornamental-resources air-quality-regulation climate-regulation water-regulation erosion-regulation water-purification waste-treatment disease-regulation pest-regulation pollination seed-dispersal habitat-provision nutrient-cycling soil-formation primary-production recreation tourism aesthetic-value spiritual-value educational-value scientific-value inspiration sense-of-place cultural-heritage traditional-knowledge indigenous-knowledge local-knowledge ecosystem-service-valuation economic-valuation contingent-valuation choice-experiment benefit-transfer total-economic-value use-value non-use-value direct-use indirect-use option-value bequest-value existence-value willingness-to-pay willingness-to-accept cost-benefit-analysis cost-effectiveness-analysis multi-criteria-analysis decision-support uncertainty risk-assessment scenario-analysis sensitivity-analysis robustness-analysis adaptive-capacity resilience vulnerability exposure sensitivity coping-capacity transformation social-ecological-system complex-adaptive-system tipping-point regime-shift alternative-stable-state hysteresis path-dependence legacy-effect slow-variable fast-variable panarchy adaptive-cycle exploitation release reorganization foresight anticipatory-governance transformative-change sustainability-science future-earth interdisciplinary transdisciplinary problem-oriented solution-oriented action-oriented knowledge-co-production boundary-work boundary-object boundary-organization science-policy-interface science-society-interface knowledge-brokerage knowledge-exchange knowledge-integration mode-1-science mode-2-science triple-helix quadruple-helix responsible-innovation reflexive-governance adaptive-governance collaborative-governance polycentric-governance network-governance self-organization collective-action common-pool-resource tragedy-of-the-commons social-dilemma institutional-analysis institutional-design Ostrom-design-principles monitoring sanctioning conflict-resolution nested-enterprises proportionality collective-choice constitutional-choice operational-choice collective-action-theory game-theory prisoner's-dilemma chicken-game assurance-game coordination-game evolutionary-game-theory replicator-dynamics ESS Nash-equilibrium Pareto-optimality social-optimum market-failure government-failure voluntary-failure non-market-value public-good common-good club-good private-good rivalry excludability free-rider hold-up-problem moral-hazard adverse-selection information-asymmetry transaction-cost property-right common-property state-property private-property hybrid-property bundle-of-rights usufruct access withdrawal alienation tenure-security land-tenure water-tenure resource-tenure customary-tenure statutory-tenure legal-pluralism de-facto de-jure formal informal legality legitimacy recognition deviance corruption elite-capture social-capital bonding-capital bridging-capital linking-capital trust reciprocity reputation social-network social-embeddedness structural-hole brokerage closure cohesion density centrality betweenness degree eigenvector PageRank small-world scale-free random-network regular-network hierarchical-network modularity community-structure clustering assortativity homophily heterophily preferential-attachment fitness-model network-evolution network-dynamics temporal-network multilayer-network multiplex-network bipartite-network affiliation-network one-mode-network two-mode-network adjacency-matrix edge-list graph-theory vertex edge node link tie connection path cycle tree forest component giant-component percolation phase-transition criticality self-organized-criticality power-law heavy-tail Pareto-distribution lognormal-distribution exponential-distribution Poisson-distribution binomial-distribution normal-distribution Gaussian-distribution statistical-distribution probability-distribution cumulative-distribution probability-density-function cumulative-distribution-function survival-function hazard-function moment mean median mode variance standard-deviation coefficient-of-variation skewness kurtosis quantile percentile decile quartile interquartile-range range minimum maximum outlier robust-statistic parametric-test non-parametric-test hypothesis-testing null-hypothesis alternative-hypothesis significance-level p-value confidence-interval type-I-error type-II-error statistical-power effect-size sample-size sampling-design experimental-design observational-study quasi-experiment natural-experiment randomized-controlled-trial before-after-control-impact BACI impact-assessment environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint material-flow-analysis substance-flow-analysis energy-analysis exergy-analysis emergy-analysis input-output-analysis environmentally-extended-input-output social-accounting-matrix computable-general-equilibrium agent-based-model system-dynamics-model Bayesian-network structural-equation-model path-analysis factor-analysis principal-component-analysis cluster-analysis discriminant-analysis canonical-correspondence-analysis redundancy-analysis non-metric-multidimensional-scaling ordination classification regression-analysis linear-regression nonlinear-regression generalized-linear-model generalized-additive-model mixed-effects-model multilevel-model hierarchical-model Bayesian-hierarchical-model state-space-model hidden-Markov-model time-series-analysis spatial-analysis geostatistics kriging variogram spatial-autocorrelation Moran's-I Geary's-c Getis-Ord-Gi local-indicator-of-spatial-association LISA hot-spot cold-spot spatial-cluster spatial-outlier edge-effect boundary-effect modifiable-areal-unit-problem MAUP ecological-fallacy atomistic-fallacy cross-scale-inference upscaling downscaling scaling-law allometric-scaling fractal fractal-dimension self-similarity multi-fractal lacunarity roughness texture pattern process pattern-process-relationship neutral-model null-model randomization-test permutation-test bootstrap jackknife cross-validation model-averaging model-comparison information-criterion Bayesian-model-averaging ensemble-modeling consensus-model prediction forecast projection scenario narrative-scenario quantitative-scenario storyline shared-socioeconomic-pathway representative-concentration-pathway emission-scenario climate-scenario impact-scenario adaptation-scenario mitigation-scenario transformation-scenario backcasting forecasting trend-extrapolation expert-judgment Delphi-method participatory-modeling stakeholder-analysis influence-diagram conceptual-model mental-model system-archetype leverage-point intervention-point policy-instrument economic-instrument regulatory-instrument voluntary-instrument information-instrument market-based-instrument command-and-control-instrument subsidy tax charge fee levy deposit-refund tradable-permit emission-trading carbon-tax biodiversity-banking conditional-cash-transfer community-based-natural-resource-management joint-forest-management co-management adaptive-co-management transboundary-management international-regime global-governance multilateral-environmental-agreement convention treaty cartagena-protocol nagoya-protocol Aichi-target sustainable-development-goal SDG post-2020-global-biodiversity-framework Kunming-Montreal-global-biodiversity-framework 30-by-30-target nature-based-solution NbS ecosystem-based-adaptation ecosystem-based-mitigation green-infrastructure blue-infrastructure natural-infrastructure hybrid-infrastructure climate-smart-agriculture climate-smart-landscape climate-smart-development low-carbon-development green-economy circular-economy bioeconomy blue-economy sustainable-consumption sustainable-production life-cycle-thinking cradle-to-cradle industrial-ecology industrial-symbiosis urban-metabolism socio-ecological-metabolism material-intensity energy-intensity carbon-intensity decoupling absolute-decoupling relative-decoupling resource-efficiency eco-efficiency eco-innovation green-technology clean-technology environmental-technology appropriate-technology participatory-technology technology-transfer technology-diffusion innovation-system national-innovation-system sectoral-innovation-system technological-innovation-system socio-technical-system socio-technical-transition multi-level-perspective niche-innovation regime landscape transition-management strategic-niche-management transition-experiment living-lab real-world-laboratory urban-living-lab rural-living-lab sustainability-transition fundamental-change system-innovation directionality reflexivity responsiveness anticipation learning-by-doing doing-by-learning social-learning organizational-learning policy-learning transformative-capacity capacity-development institutional-learning evolutionary-governance institutional-change path-dependency lock-in carbon-lock-in technological-lock-in institutional-lock-in behavioral-lock-in cultural-lock-in unlocking destabilization phase-out divestment stranded-asset just-transition transition-pathway transition-scenario deep-decarbonization net-zero carbon-neutral climate-neutral nature-positive biodiversity-positive restoration-positive regenerative circular doughnut-economics planetary-boundary safe-operating-space environmental-ceiling social-foundation intergenerational-equity intragenerational-equity distributive-justice procedural-justice recognition-justice environmental-justice climate-justice energy-justice water-justice food-justice spatial-justice territorial-justice ecological-justice species-justice intersectionality adaptation mitigation sustainability sustainable-development well-being human-development capability-approach human-security environmental-security climate-security water-security food-security energy-security livelihood-security disaster-risk-reduction climate-change-adaptation climate-change-mitigation loss-and-damage climate-finance green-climate-fund adaptation-fund global-environment-facility biodiversity-finance nature-finance blended-finance impact-investment ESG environmental-social-governance sustainable-finance green-bond social-bond sustainability-bond blue-bond nature-bond biodiversity-credit offset inset nature-based-credit ecosystem-service-credit payment-for-results results-based-payment conditional-payment performance-based-payment outcome-based-payment environmental-service-payment PSA PES REDD REDD+AFOLU LULUCF NAMA NAMC NDC NAP NBSAP national-adaptation-plan national-biodiversity-strategy-and-action-plan nationally-determined-contribution nationally-appropriate-mitigation-action intended-nationally-determined-contribution long-term-low-emission-development-strategy net-zero-target carbon-budget remaining-carbon-budget emission-gap adaptation-gap finance-gap implementation-gap ambition-loop ratchet-mechanism global-stocktake transparency-framework enhanced-transparency-framework common-but-differentiated-responsibilities respective-capabilities climate-equity climate-litigation climate-governance climate-diplomacy climate-negotiation UNFCCC COP CMP CMA SBSTA SBI IPCC IPBES UNCCD CBD CITES Ramsar World-Heritage Man-and-the-Biosphere MAB UNESCO FAO WHO ILO UNDP UNEP UN-Habitat World-Bank IMF WTO OECD GEF GCF AF multilateral-development-bank regional-development-bank national-development-bank development-finance-institution export-credit-agency sovereign-wealth-fund pension-fund insurance-company asset-manager asset-owner corporate-investor retail-investor impact-investor philanthropic-investor development-partner bilateral-donor multilateral-donor private-donor foundation NGO CSO civil-society-organization non-governmental-organization community-based-organization indigenous-peoples-organization women's-organization youth-organization faith-based-organization research-organization think-tank university academic-institution knowledge-institution capacity-building-institution training-institution extension-service advisory-service consulting-firm engineering-firm technology-provider equipment-supplier service-provider contractor operator maintainer utility municipality local-government provincial-government national-government federal-government ministry department agency authority commission board council committee parliament congress legislature executive judiciary media journalist reporter editor publisher broadcaster platform social-media influencer celebrity activist advocate campaigner mobilizer organizer networker connector bridge-builder boundary-spanner knowledge-broker science-communicator extension-agent development-practitioner policy-maker decision-maker planner manager administrator leader champion entrepreneur innovator intrapreneur social-entrepreneur systems-entrepreneur institutional-entrepreneur policy-entrepreneur political-entrepreneur business-model business-model-innovation social-business inclusive-business base-of-pyramid fortune-at-bottom-of-pyramid shared-value creating-shared-value stakeholder-capitalism purpose-driven-business B-Corp benefit-corporation social-enterprise cooperative mutual association collective commons commoning peer-production open-source creative-commons knowledge-commons digital-commons data-commons genetic-commons seed-commons water-commons fisheries-commons forest-commons pasture-commons land-commons urban-commons atmospheric-commons global-commons common-heritage-of-mankind common-concern-of-humankind restorative-justice multispecies-justice more-than-human anthropocene capitalocene plantationocene chthulucene symbiocene good-anthropocene great-acceleration great-transformation great-decoupling great-convergence great-divergence great-fragmentation great-integration great-transition great-work long-now deep-time futures-thinking future-studies strategic-foresight scenario-planning trend-analysis weak-signal wild-card black-swan grey-rhino blue-sky green-swan systemic-change structural-change cultural-change behavioral-change individual-change collective-change social-change organizational-change technological-change economic-change political-change ecological-change planetary-change earth-system-change biodiversity-change ocean-change cryosphere-change atmospheric-change biogeochemical-change nitrogen-cycle-change phosphorus-cycle-change carbon-cycle-change water-cycle-change hydrological-change soil-change vegetation-change ecosystem-change landscape-change seascape-change biosphere-change noosphere-change technosphere-change anthroposphere-change sociosphere-change econosphere-change politosphere-change culturosphere-change semiosphere-change infosphere-change digisphere-change cybersphere-change urban-sphere-change agricultural-sphere-change industrial-sphere-change extractive-sphere-change manufacturing-sphere-change service-sphere-change financial-sphere-change knowledge-sphere-change innovation-sphere-change creativity-sphere-change culture-sphere-change identity-sphere-change meaning-sphere-change value-sphere-change purpose-sphere-change vision-sphere-change mission-sphere-change strategy-sphere-change tactic-sphere-change operation-sphere-change action-sphere-change practice-sphere-change habit-sphere-change routine-sphere-change ritual-sphere-change tradition-sphere-change custom-sphere-change convention-sphere-change norm-sphere-change rule-sphere-change law-sphere-change regulation-sphere-change standard-sphere-change protocol-sphere-change procedure-sphere-change process-sphere-change system-sphere-change structure-sphere-change function-sphere-change pattern-sphere-change dynamic-sphere-change state-sphere-change phase-sphere-change regime-sphere-change equilibrium-sphere-change stability-sphere-change instability-sphere-change chaos-sphere-change order-sphere-change complexity-sphere-change simplicity-sphere-change diversity-sphere-change uniformity-sphere-change heterogeneity-sphere-change homogeneity-sphere-change variation-sphere-change invariance-sphere-change change-sphere-change continuity-sphere-change discontinuity-sphere-change emergence-sphere-change submergence-sphere-change evolution-sphere-change devolution-sphere-change revolution-sphere-change involution-sphere-change convolution-sphere-change resolution-sphere-change dissolution-sphere-change absolution-sphere-change solution-sphere-change
Williamsonia fletcheri
Ebony Boghaunter
Williamsonia fletcheri, commonly known as the ebony boghaunter, is a small dragonfly in the emerald family Corduliidae. It is one of North America's earliest-flying dragonflies, active in May and June. The species is strongly associated with acidic bog habitats and has a restricted range in the northeastern United States and southeastern Canada. It is considered rare across most of its range and is classified as endangered in Massachusetts.
dragonfly Odonata Corduliidae bog-specialist early-season rare endangered acidic-habitat North-America
Zale curema
Black-eyed Zale Moth, Northeastern Pine Zale, Black-eyed Zale
Zale curema is a moth in the family Erebidae, first described by John Bernhard Smith in 1908. The species is strongly associated with pitch pine forests in eastern North America. It has one generation per year, with larvae feeding preferentially on young needles of pitch pine. The species is listed as endangered in Connecticut but considered globally secure.
moth Erebidae pitch-pine univoltine endangered Connecticut forest-specialist North-America herbivore conifer-feeder