Sphecidae
Guides
Agrilus obsoletoguttatus
Beech Borer
Agrilus obsoletoguttatus is a small metallic wood-boring beetle in the family Buprestidae, native to North America. It is among the smallest jewel beetle species utilized as prey by the specialist predatory wasp Cerceris fumipennis, which provisions its underground nests with paralyzed buprestid beetles. The species has been documented in nest caches containing up to 13 individuals, reflecting its small size relative to larger buprestid prey.
Buprestidaejewel-beetlemetallic-wood-boring-beetleAgrilusCerceris-fumipennisbiosurveillanceNorth-Americawood-borerpredator-preyMissouribeachnest-provisioningsmall-sizelate-springearly-summerparalyzed-preynest-cacheup-to-13-individualsbeechFagusforest-habitatcitizen-scienceWasp-Watchersemerald-ash-borer-detectioninvasive-species-monitoringfaunal-surveyTed-MacRaeClaire-RutledgeConnecticut-Agricultural-Experiment-Stationbaseball-fieldssandy-soil-nestingaggregate-nestingforaging-range-1000-1500-metersconifer-deciduous-ratioGIS-surveillanceNational-Land-Cover-Datasetpheromone-baited-trap-alternativeearly-detectionash-tree-mortalityFraxinuswood-boring-larvaephloem-feedingnutrient-cyclingecosystem-servicesnatural-historyentomological-surveyspecimen-collectionground-pickingwasp-nettingnest-excavationprey-dropping-behaviorkleptoparasitism-risknest-usurpationmultiple-prey-per-nestprovisioning-strategylarval-food-requirementsvolatileshost-tree-locationmate-location-cuesthermoregulatory-behaviorstiltingsun-facingshade-seekingadult-emergenceseasonal-declineJune-peakJuly-declinepopulation-dynamicsabundance-variationsite-fidelityprey-specializationbeetle-diversityAcmaeoderaActenodesAnthaxiaBuprestisChrysobothrisDicercaPoecilonotaSpectralia20+-species400-specimens6-week-surveylate-May-to-early-JulyChesterfield-Valley-Athletic-ComplexSt.-Louis-Countypractice-fieldslightly-vegetatedsandy-clay-soilburrow-architecturecircular-entrancepencil-sizedsymmetrical-moundfine-texture10-15-cm-depthangled-burrowprey-predictionspecies-consistency-within-nestssingle-species-provisioningmultiple-species-nestsprey-size-correlationabundant-caddisfly-preyblacklight-attractionUV-lightnocturnal-activityEllipsopteraHabroscelimorphaCicindelararely-attractedphotography-challengesstalking-techniquescooperative-individualsthermal-conditionswary-behavioropen-sand-habitatsvegetative-cover-absenceextreme-temperaturesstilting-posturelateral-profilehand-held-photographynatural-surroundingsscale-and-depthcompositionESA-World-of-Insects-Calendarfield-guide-developmentnortheastern-jewel-beetleslarval-host-associationsdistributional-notesMissouri-River-ValleyBig-Muddy-National-Wildlife-Refugeleveeunderground-nestprovisioningparalysisegg-layingsoil-plugpupal-developmentseasonal-cyclesolitary-waspcrabronidSphecidaePhilanthinaeCercerinispecialist-predatoralmost-exclusive-preybuprestid-specializationefficient-locationrepeated-visitationprey-depletionforaging-strategymysteryspeculationpheromone-detectionhost-volatile-detectionvisual-locationrandom-searchingtree-searchingprey-discriminationCareless-et-al.-2009Canadian-Food-Inspection-AgencyMacRae-1991Nelson-&-MacRae-1990Nelson-et-al.-1996MacRae-&-Nelson-2003MacRae-2004MacRae-2006Pearson-et-al.-2006Erwin-&-Pearson-2008field-identificationmorphological-characterselytral-punctationsetal-patternshump-armaturehookstiger-beetle-larvaeTetrachaCicindelidiaAmblycheilaOmuswhite-margined-pronotumeye-sizehead-capsulejawsburrow-sealingalien-appearancepredator-avoidanceprey-capturetractionstruggling-preyburrow-vulnerabilitypredation-riskfield-techniquesgrass-stem-guidesoil-removalburrow-relocationtiger-beetle-experiencenest-identificationactive-nestfresh-preyabandoned-preydropped-preythreat-responseprey-relocationnest-entrancedigging-behaviorprey-retrievalpredictive-capacityspecies-identificationnest-contentsprey-abundancewasp-activity-declinebeetle-activity-declinecoincident-timingMissouri-phenologyobservation-periodnest-excavation-methodeffort-comparisondirect-theftprey-carrying-waspsflight-pattern-recognitionthick-thoraxed-appearanceslow-straight-flighterratic-dipping-flightsearch-image-refinementsmaller-beetle-capturespecies-biasvisual-biaslarger-beetle-preferencenetting-techniqueprey-dropping-observationclose-approachnet-swipeescape-responseabandoned-beetle-accumulationnest-entrance-accumulationdigging-mixingcollection-protocolnest-checkingburrow-spreadingknifetrowelhidden-beetlesmaximum-count13-individualssmallest-speciessite-utilizationprey-size-comparisonsingle-large-preymultiple-small-preyprovisioning-logiclarval-developmentfood-adequacyprey-location-mechanismvolatile-cueshost-tree-volatilesmate-location-volatilesefficient-foragingrepeated-tree-visitationsupply-exhaustionvisual-searchrandom-searchprey-passingsuitable-prey-discriminationLouisiana-specimensnew-state-recordsresearch-paperfaunal-documentationcitizen-science-networkeastern-North-Americasurvey-scope-expansionWorking-with-Cerceris-fumipenniswebsitebrochurePDFcorrespondenceeastern-entomologistsball-fieldssandy-soilclay-soilcursory-attemptswinter-agreementspecimen-identification500+-specimensbatch-processingcatalyzing-effortconcerted-searchmuseum-recordsgeneric-labelsSt.-LouisColumbiamonthly-visitsregular-groomingheavy-claybarren-soilburrow-absencestroke-of-luckbike-routeknee-painflatter-routeregular-fieldsgrooming-evidencehuman-useimmediate-inspectionnumerous-burrowsoccupied-nestfemale-identificationyellow-facial-markingsburrow-occupancyactive-diggingsoil-pushi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Thread-waisted Sand Wasps
Ammophila is a large, cosmopolitan genus of solitary hunting wasps in the family Sphecidae, comprising over 200 species distributed across warmer regions of all continents except Antarctica. These thread-waisted wasps are characterized by their elongated, slender petiole connecting the thorax and abdomen, and their distinctive nesting behavior involving burrow excavation and caterpillar provisioning. Females construct underground nests in sandy or compact soils, paralyze caterpillars with their sting, and provide this food source for their developing larvae. The genus serves as a host for Strepsiptera endoparasites and is subject to nest parasitism by satellite flies and other organisms.
Ammophila cleopatra
Ammophila cleopatra is a species of thread-waisted wasp in the family Sphecidae, described by Arnold Menke in 1964. Like other members of the genus Ammophila, it is a solitary wasp that hunts caterpillars to provision nests for its offspring. The species occurs in North America and Middle America, though specific details about its biology remain poorly documented compared to better-studied congeners such as A. procera and A. pictipennis.
Chlorion
cricket hunter wasps, steel-blue cricket hunters
Chlorion is a genus of solitary sphecid wasps distributed across the Americas, Africa, and tropical Asia through Java. The genus contains approximately 20 described species, including the well-known Steel-blue Cricket Hunter Wasp (Chlorion aerarium). Females construct underground burrows provisioned with paralyzed orthopteran prey—primarily crickets for most species, though Chlorion cyaneum preys on cockroaches. The genus is characterized by metallic blue to violet coloration, large size, and distinctive nesting behavior often associated with hard-packed soil habitats.
Chlorion cyaneum
purplish-blue cricket hunter, Iridescent Cockroach-hunter Wasp
Chlorion cyaneum, commonly known as the purplish-blue cricket hunter or iridescent cockroach-hunter wasp, is a species of thread-waisted wasp in the family Sphecidae. It is a solitary predatory wasp that hunts cockroaches as prey. The species exhibits a distinctive metallic blue to purplish coloration and is found in North America and Middle America.
Eremnophila
thread-waisted wasps
Eremnophila is a genus of solitary thread-waisted wasps in the family Sphecidae, comprising approximately nine described species distributed primarily across the Americas. These wasps are specialized caterpillar hunters that provision subterranean nests with paralyzed prey for their larvae. Adults are frequent flower visitors, particularly to spotted horsemint (Monarda punctata), goldenrods, and other late-summer asters. A distinctive behavioral trait involves prolonged tandem mating, where males grasp females by the neck and accompany them during foraging activities.
Isodontia
Grass-carrying Wasps, Grass-carrier Wasps
Isodontia is a genus of solitary wasps in the family Sphecidae, commonly known as grass-carrying wasps. Females construct nests in pre-existing cavities such as hollow stems, window tracks, or abandoned beetle tunnels, lining them with dry grass or hay. They provision nests with paralyzed Orthoptera (tree crickets and katydids) as food for their larvae. Several species are invasive in Europe, with Isodontia mexicana being the most widespread non-native species.
Isodontia apicalis
grass-carrier wasp
Isodontia apicalis is a solitary thread-waisted wasp in the family Sphecidae, known as a 'grass-carrier' for its distinctive nesting behavior. The species occurs in North and Central America. Like other members of its genus, females construct nests using dry grass to partition cells and plug entrances.
Isodontia auripes
Brown-legged Grass-carrier Wasp, brown-legged grass-carrier
Isodontia auripes, the brown-legged grass-carrier wasp, is a solitary thread-waisted wasp in the family Sphecidae. The species is known for its distinctive nesting behavior: females construct nests in pre-existing cavities such as window tracks, wind chimes, or abandoned carpenter bee tunnels, filling them with dry grass and provisioning them with paralyzed tree crickets (Oecanthus spp.) as food for their larvae. The wasp is non-aggressive toward humans and has been observed using nests made by Xylocopa virginica or mining bees. Larvae feed for approximately three days before pupating for two days.
Isodontia elegans
Elegant Grass-carrying Wasp
Isodontia elegans is a solitary, nest-provisioning thread-waisted wasp in the family Sphecidae. Commonly known as the elegant grass-carrying wasp, it hunts orthopteran prey including tree crickets and katydids, which it paralyzes and stores in nest cells for its developing larvae. The species is notable for using pre-existing cavities—particularly window tracks and wind chime tubes in suburban settings—lining them with finely chewed grass fibers that serve as both partitions between cells and defensive barriers against parasites.
Isodontia mexicana
Mexican Grass-carrying Wasp, Grass-carrying Wasp
Isodontia mexicana is a solitary sphecid wasp native to North America that has become a successful invasive species in Europe since the 1960s. Females construct nests in pre-existing cavities such as hollow stems, abandoned beetle tunnels, or even window tracks, lining them with grass fragments. They provision nests with paralyzed orthopteran prey—primarily small katydids (Tettigoniidae) and tree crickets (Gryllidae, particularly Oecanthinae)—to feed their larvae. The species is notable for its splayed wing posture at rest and has been observed visiting diverse flowering plants for nectar.
Isodontia philadelphica
Grass-carrying Wasp
Isodontia philadelphica is a solitary wasp in the family Sphecidae, commonly known as a grass-carrying wasp. Females construct nests in pre-existing cavities such as hollow stems, abandoned insect burrows, or window tracks, using dry grass to partition cells and plug entrances. They provision nests with paralyzed tree crickets and katydids as food for their larvae. The species occurs in North America and is not aggressive toward humans.
Palmodes dimidiatus
Palmodes dimidiatus is a species of thread-waisted wasp in the family Sphecidae. It is a solitary, ground-nesting wasp that hunts katydids (Tettigoniidae) as prey for its larval offspring. The species is known to be parasitized by the kleptoparasitic wasp Stizoides renicinctus, which raids its food caches. Palmodes dimidiatus occurs in North America and Middle America.
Palmodes praestans
Palmodes praestans is a thread-waisted wasp in the family Sphecidae, characterized by its distinctive thread-like petiole connecting the thorax and abdomen. The species is known to be kleptoparasitized by Stizoides renicinctus, which exploits its food caches of katydid prey. Like other members of its genus, it hunts katydids (Tettigoniidae) to provision underground nest cells for its larvae.
Podium
Podium is a genus of solitary wasps in the family Sphecidae, subfamily Sceliphrinae, established by Fabricius in 1804. These thread-waisted wasps are part of the diverse sphecid wasp lineage, which includes many species that construct nests from mud or hunt prey to provision their offspring. The genus is classified within the tribe Podiini, which shares its root name. As with other sphecid wasps, members of this genus exhibit the characteristic narrow petiole (thread-waisted appearance) and are likely predatory, though specific ecological details for the genus as a whole remain limited in available sources.
Podium krombeini
Podium krombeini is a species of thread-waisted wasp in the family Sphecidae, described by Bohart and Menke in 1963. It belongs to the tribe Podiini within the subfamily Sceliphrinae. The genus Podium comprises predatory wasps that construct mud nests. This species is recorded from Middle America and North America, though specific details about its biology remain poorly documented in the accessible literature.
Prionyx
Prionyx is a genus of solitary thread-waisted wasps in the family Sphecidae, distributed across the Americas from southern Canada to Argentina. All species are specialist predators of grasshoppers (Acrididae), which they paralyze with venom and provision as food for their larvae in underground burrows. The genus is characterized by a globose abdomen, spiny legs with saw-like tarsal claws, and distinctive nesting behaviors. Females dig burrows after securing prey, transport paralyzed grasshoppers to single-celled nests, and seal the entrance upon completion. Some species exhibit facultative kleptoparasitism when nesting aggregations overlap with other wasp species.
Prionyx fervens
Prionyx fervens is a solitary thread-waisted wasp in the family Sphecidae. It is a ground-nesting predator that hunts grasshoppers, paralyzes them, and provisions its burrow with living prey for larval development. The species ranges from Argentina to the southern United States, with documented nesting behavior observed in coastal southeastern Brazil.
Prionyx parkeri
Parker's Thread-waisted Wasp
Prionyx parkeri is a species of thread-waisted wasp in the family Sphecidae. It is one of seven Prionyx species found in the United States. The species is distinguished from close relatives by two pairs of long palps that are part of its mouthparts. Like other members of the genus, it is a solitary parasitoid wasp that hunts grasshoppers as prey for its larvae.
Prionyx thomae
Prionyx thomae is a solitary, thread-waisted wasp in the family Sphecidae. It is one of seven Prionyx species in the United States, ranging from the southeastern and western U.S. south to Argentina. The species is a specialist predator of grasshoppers, paralyzing them with venom to provision underground nests for its larvae. It lacks a common English name, reflecting its limited economic importance.
Sceliphron curvatum
Asian mud-dauber wasp
Sceliphron curvatum is a solitary mud-daubing wasp native to Central Asia that has become invasive in Europe and North America. First recorded in Europe in 1979, it has since expanded across multiple European countries and was first documented in North America in 2013. The species is smaller and darker than the native North American Sceliphron caementarium, with a body length of 15–25 mm and black coloration with yellow and red ornaments. It exhibits strongly synanthropic behavior, frequently nesting indoors on clothing, books, and furniture rather than exclusively on building exteriors.
Sphecinae
digger wasps, thread-waisted wasps, grass-carrying wasps
Sphecinae is a subfamily of digger wasps (family Sphecidae) characterized by thread-waisted bodies and solitary nesting behavior. Members construct nests in soil, plant stems, or pre-existing cavities, provisioning cells with paralyzed prey for their larvae. The subfamily includes notable genera such as Sphex, Isodontia, and Podalonia, with diverse nesting strategies ranging from burrowing to grass-carrying.
Sphex flavovestitus
Sphex flavovestitus is a species of thread-waisted digger wasp in the family Sphecidae, first described by Frederick Smith in 1856. The species is distributed across the Americas, with records from North America, Middle America, and South America. Two subspecies are recognized: the nominate Sphex flavovestitus flavovestitus and Sphex flavovestitus saussurei. Like other members of the genus Sphex, it is a solitary wasp that constructs subterranean nests provisioned with paralyzed prey for its larvae.
Sphex habenus
Golden-reined Digger Wasp
Sphex habenus is a solitary thread-waisted wasp in the family Sphecidae. Like other members of the genus Sphex, females construct individual subterranean nests provisioned with paralyzed prey for their larvae. The species is distributed across North and Middle America.
Sphex nudus
Katydid Wasp
Sphex nudus, commonly known as the katydid wasp, is a solitary digger wasp in the family Sphecidae. Females construct subterranean nests in soil or dirt substrates, including the floors of old barns. The species is notable for its specialized hunting behavior, targeting leaf-rolling crickets as prey for its larvae. It occurs across the eastern United States.
Sphex pensylvanicus
Great Black Wasp, Great Black Digger Wasp
Sphex pensylvanicus is a large, solitary digger wasp native to North America. Females construct underground burrows and provision them with paralyzed orthopteran prey, primarily katydids, for their larvae. Adults feed on nectar from diverse flowering plants and serve as pollinators. The species exhibits protandry, with males emerging before females in the breeding season. Despite their formidable size and appearance, they are not aggressive toward humans and sting only when handled.
Sphex tepanecus
Sphex tepanecus is a species of thread-waisted digger wasp in the family Sphecidae. It is a large solitary wasp known from the western United States, with records from Arizona, Colorado, and potentially Idaho. Like other members of the genus Sphex, it constructs subterranean nests and provisions them with paralyzed prey for its larvae. The species was first described by de Saussure in 1867.