Fraxinus

Guides

Aceria fraxini
Ash Bead Gall Mite
Aceria fraxini, commonly known as the ash bead gall mite, is an eriophyid mite that induces distinctive capsule galls on ash trees (Fraxinus spp.). This species exhibits a complex life cycle involving alternation of generations, with an overwintering generation consisting solely of parthenogenetic females (deutogynes) and a summer generation containing both sexes (protogynes and males). The mite is distributed across North America, where it is considered a minor pest of ornamental and native ash species.
gall-mite eriophyid Fraxinus North-America alternation-of-generations deutogyne protogyne
Agrilus obsoletoguttatus
Beech Borer
Agrilus obsoletoguttatus is a small metallic wood-boring beetle in the family Buprestidae, native to North America. It is among the smallest jewel beetle species utilized as prey by the specialist predatory wasp Cerceris fumipennis, which provisions its underground nests with paralyzed buprestid beetles. The species has been documented in nest caches containing up to 13 individuals, reflecting its small size relative to larger buprestid prey.
Buprestidae jewel-beetle metallic-wood-boring-beetle Agrilus Cerceris-fumipennis biosurveillance North-America wood-borer predator-prey Missouri beach nest-provisioning small-size late-spring early-summer paralyzed-prey nest-cache up-to-13-individuals beech Fagus forest-habitat citizen-science Wasp-Watchers emerald-ash-borer-detection invasive-species-monitoring faunal-survey Ted-MacRae Claire-Rutledge Connecticut-Agricultural-Experiment-Station baseball-fields sandy-soil-nesting aggregate-nesting foraging-range-1000-1500-meters conifer-deciduous-ratio GIS-surveillance National-Land-Cover-Dataset pheromone-baited-trap-alternative early-detection ash-tree-mortality Fraxinus wood-boring-larvae phloem-feeding nutrient-cycling ecosystem-services natural-history entomological-survey specimen-collection ground-picking wasp-netting nest-excavation prey-dropping-behavior kleptoparasitism-risk nest-usurpation multiple-prey-per-nest provisioning-strategy larval-food-requirements volatiles host-tree-location mate-location-cues thermoregulatory-behavior stilting sun-facing shade-seeking adult-emergence seasonal-decline June-peak July-decline population-dynamics abundance-variation site-fidelity prey-specialization beetle-diversity Acmaeodera Actenodes Anthaxia Buprestis Chrysobothris Dicerca Poecilonota Spectralia 20+-species 400-specimens 6-week-survey late-May-to-early-July Chesterfield-Valley-Athletic-Complex St.-Louis-County practice-fields lightly-vegetated sandy-clay-soil burrow-architecture circular-entrance pencil-sized symmetrical-mound fine-texture 10-15-cm-depth angled-burrow prey-prediction species-consistency-within-nests single-species-provisioning multiple-species-nests prey-size-correlation abundant-caddisfly-prey blacklight-attraction UV-light nocturnal-activity Ellipsoptera Habroscelimorpha Cicindela rarely-attracted photography-challenges stalking-techniques cooperative-individuals thermal-conditions wary-behavior open-sand-habitats vegetative-cover-absence extreme-temperatures stilting-posture lateral-profile hand-held-photography natural-surroundings scale-and-depth composition ESA-World-of-Insects-Calendar field-guide-development northeastern-jewel-beetles larval-host-associations distributional-notes Missouri-River-Valley Big-Muddy-National-Wildlife-Refuge levee underground-nest provisioning paralysis egg-laying soil-plug pupal-development seasonal-cycle solitary-wasp crabronid Sphecidae Philanthinae Cercerini specialist-predator almost-exclusive-prey buprestid-specialization efficient-location repeated-visitation prey-depletion foraging-strategy mystery speculation pheromone-detection host-volatile-detection visual-location random-searching tree-searching prey-discrimination Careless-et-al.-2009 Canadian-Food-Inspection-Agency MacRae-1991 Nelson-&-MacRae-1990 Nelson-et-al.-1996 MacRae-&-Nelson-2003 MacRae-2004 MacRae-2006 Pearson-et-al.-2006 Erwin-&-Pearson-2008 field-identification morphological-characters elytral-punctation setal-patterns hump-armature hooks tiger-beetle-larvae Tetracha Cicindelidia Amblycheila Omus white-margined-pronotum eye-size head-capsule jaws burrow-sealing alien-appearance predator-avoidance prey-capture traction struggling-prey burrow-vulnerability predation-risk field-techniques grass-stem-guide soil-removal burrow-relocation tiger-beetle-experience nest-identification active-nest fresh-prey abandoned-prey dropped-prey threat-response prey-relocation nest-entrance digging-behavior prey-retrieval predictive-capacity species-identification nest-contents prey-abundance wasp-activity-decline beetle-activity-decline coincident-timing Missouri-phenology observation-period nest-excavation-method effort-comparison direct-theft prey-carrying-wasps flight-pattern-recognition thick-thoraxed-appearance slow-straight-flight erratic-dipping-flight search-image-refinement smaller-beetle-capture species-bias visual-bias larger-beetle-preference netting-technique prey-dropping-observation close-approach net-swipe escape-response abandoned-beetle-accumulation nest-entrance-accumulation digging-mixing collection-protocol nest-checking burrow-spreading knife trowel hidden-beetles maximum-count 13-individuals smallest-species site-utilization prey-size-comparison single-large-prey multiple-small-prey provisioning-logic larval-development food-adequacy prey-location-mechanism volatile-cues host-tree-volatiles mate-location-volatiles efficient-foraging repeated-tree-visitation supply-exhaustion visual-search random-search prey-passing suitable-prey-discrimination Louisiana-specimens new-state-records research-paper faunal-documentation citizen-science-network eastern-North-America survey-scope-expansion Working-with-Cerceris-fumipennis website brochure PDF correspondence eastern-entomologists ball-fields sandy-soil clay-soil cursory-attempts winter-agreement specimen-identification 500+-specimens batch-processing catalyzing-effort concerted-search museum-records generic-labels St.-Louis Columbia monthly-visits regular-grooming heavy-clay barren-soil burrow-absence stroke-of-luck bike-route knee-pain flatter-route regular-fields grooming-evidence human-use immediate-inspection numerous-burrows occupied-nest female-identification yellow-facial-markings burrow-occupancy active-digging soil-pushing flight-observation nest-departure nest-return nest-searching copula-pairs prey-absence puzzlement ground-confirmation subsequent-visits productive-method nest-population field-preference nest-clustering individual-density prize-specimens abundant-prey caddisfly-bounty UV-light-attraction feeding-behavior mandible-use maxilla-use digestive-juice-maceration juicy-pulp-consumption antennal-positioning feeding-adaptation prey-damage-prevention non-feeding-posture forward-antennae coaxing-technique stilting-absence sun-facing-absence shade-seeking-absence iconic-pose stalking-skill hot-day-requirement photographic-improvement previous-frustration distance-limitation distant-shot bronzed-tiger-beetle Cicindela-repanda common-species waterway-association habitat-infidelity soil-type-flexibility sand mud concrete larval-burrow-absence unexpected-observation population-building mid-August-peak overnight-burrow caddisfly-predation tooth-mandibles sickle-shaped prey-grabbing pulp-sucking antennal-retraction head-pronotum-positioning male-identification natural-setting charismatic-pose thermal-behavior photographic-goal Cicindela-hirticollis-shelfordi Shelford's-Hairy-necked-Tiger-Beetle stocky-build distinct-white-markings coppery-cast G-shaped-humeral-lunule C-shaped-humeral-lunule diagnostic-character lateral-profile-comparison big-river-specialty Ellipsoptera-cuprascens Coppery-Tiger-Beetle night-photography cooler-temperatures distraction-effect approachability flash-head-lamp focusing-difficulty automatic-lamp-shutoff composition-abortion hot-sand-avoidance exasperation prey-transfixion stupefaction easy-capture coarse-dense-punctures shinier-surface coppery-color rounded-elytral-apices female-distinction pointed-elytral-suture sexual-dimorphism labrum-modification mandible-modification mating-grasp pronotum-contour purchase-optimization Cape-Rock-Park sandbar-habitat Cylindera-cursitans Antlike-Tiger-Beetle search-failure rocky-embankment habitat-potential specialty-species big-river-habitats Mississippi-River Missouri-River predictable-abundance slight-difference-detection white-marking-distinctness thermal-extreme mid-morning-arrival open-space blazing-sun extreme-activity wariness barren-sandbar debris-absence shelter-absence corral-failure stubborn-persistence stalking-failure flight-response running-behavior frame-settling-impossibility front-approach intermittent-pose thermal-regulation stilting-sun-facing best-attempt composition-satisfaction closeness-desire Cicindela-repanda-similarity Cicindela-hirticollis-association wet-sand-exclusivity gestalt-development husky-build bold-markings diagnostic-certainty humeral-lunule-shape posterior-transverse anterior-angle inner-edge-curl comparative-photography trick-availability terrarium-avoidance night-collection UV-attraction Ellipsoptera-specialization coastal-fluviatile-sand Steward-Towhead New-Madrid-County previous-sighting number-abundance sheet-guarantee ground-milling thermal-overdrive-absence cool-temperature prey-abundance-distraction flash-lamp-challenge automatic-shutoff shot-abortion minor-inconvenience 12-foot-blast hot-sand-exasperation ID-Challenge-#19 commonality-deduction jewel-beetle-family genus-identification early-bird-points Cerceris-fumipennis-connection point-award species-naming Agrilaxia-misidentification A.-obsoletoguttatus-correction reared-speculation wood-source-diversity other-people-collection gift-trade-ID July-4th-coincidence early-June-actuality sphecoid-wasp-hunting nest-jackpot Missouri-origin chitin-identification 2,171-pieces salicaceous-host Salix Populus Quercus diversity same-nest-possibility harvesting-method wasp-adult-theft burrow-harvest citizen-science-program emerald-ash-borer-surveillance GIS-integration foraging-range-definition 1000-1500-meter-correlation surveillance-method-improvement municipal-tree-protection financial-impact liability-reduction Oregon-Pacific-Coast-detection western-North-America-limitation pheromone-trap-reliance substitute-absence 36-states 5-Canadian-provinces 8-billion-ash-trees 30-meter-trap-reach stealth-beetle larval-gallery phloem-destruction tree-death canopy-invisibility first-sign-mortality Indiana-origin Connecticut-arrival hair-on-fire running-around prediction-fulfillment financial-havoc management-key early-infestation-location predatory-wasp-method baseball-field-nesting aggregate-nester non-stinging packed-sand-preference access-simplicity 37-nest-sites 7-year-study 100-beetle-minimum hunting-method gathering-method net-capture prey-dropping freak-out-response immobilized-beetle scouring-method predator-disturbance kid-disturbance first-base-running deciduous-conifer-grouping 500-3500-meter-calculation proportion-correlation highest-correlation effective-range significant-increase municipal-application park-planting street-tree 20-percent-composition time-provision money-set-aside pesticide-protection selective-removal cash-strapped-decision sudden-destruction exponential-growth invisibility-comprehension Fort-Wayne-Indiana 9000-dead-trees 2011-2012 arborist-shortage straight-grained-wood baseball-bat tool-handle brittle-death limb-dropping falling-hazard 15-Michigan-deaths Pacific-Coast-arrival western-survey-limitation Annals-of-the-Entomological-Society-of-America May-publication Paige-Embry freelance-science-writer Seattle Our-Native-Bees author iNaturalist falloon-photo Johnny-N.-Dell-photo Bugwood.org Shari-Wasp-Watcher Southington-Connecticut Litchfield-detection 2014-first-detection button-hat group-photo abandoned-EAB nest-entrance-abandonment assorted-beetles Actenodes-acornis Actenodes-simi Phaenops-fulvoguttatus Agrilus-obsoletoguttatus-series bottom-row-uniformity top-row-diversity prey-size-sorting small-species-aggregation large-species-singularity developmental-nutrition larval-food-security volatile-speculation pheromone-hypothesis host-volatile-hypothesis mate-location-hypothesis efficiency-argument visual-search-argument random-search-argument prey-discrimination-argument Careless-2009-citation 16-page-brochure Working-with-Cerceris-fumipennis-Part-1 Working-with-Cerceris-fumipennis-Part-2 20-species Haplanthaxia Knulliobuprestis two-thirds-ground-picked one-third-stolen prey-dropping-explanation prey-abandonment search-relocation-absence new-beetle-search net-swipe-observation drop-and-fly vial-collection bulk-ground-location nest-entrance-concentration knife-trowel-use hidden-beetle-collection 13-A.-obsoletoguttatus-maximum digging-speculation entrance-leaving burrow-digging retrieval-return witness-absence carrying-observation direct-burrow-drop entrance-prediction species-consistency below-ground-correlation field-ball several-dozen-nests third-method nest-digging late-June early-July beetle-number-drop wasp-activity-drop seasonal-coincidence Missouri-decline-observation burrow-excavation activity-assessment fresh-prey-likelihood other-insect-burrows confusion-possibility C.-fumipennis-burrow perfectly-circular C.-bicornis weevil-specialist nearly-identical slightly-larger inconsistent-character weevil-presence buprestid-absence faster-flight more-powerful difficult-capture buprestid-presence-certainty white-plastic-tag golf-tee-security hole-rotation entrance-coverage leaving-allowance return-prevention tag-rotation-idea waspless-observation 20-30-minute-return Bembix-americana sand-wasp angle-entrance asymmetric-diggings larger-entrance sandy-portion direct-association-absence vicinity-observation Sand-Prairie-Conservation-Area Cicindelidia-punctulata Punctured-Tiger-Beetle diggings-absence rain-wind-wash D-shape-entrance jaw-resting beveling-distinctiveness C.-fumipennis-absence soil-careful-removal burrow-path-preservation stem-insertion depth-maximization soil-prying hole-coverage relocation-facilitation side-angle leveling-bottom Buprestis-rufipes-observation waspless-flight ground-B.-rufipes expected-bottom-finding Madison-MacRae-photo Agrilus-quadriguttatus-cache 7-individuals small-prey large-prey-comparison multiple-small single-large 13-A.-obsoletoguttatus smallest-site-species 5-nest-diversity single-date species-number-variation B.-rufipes-singularity A.-quadriguttatus-A.-obsoletoguttatus-mix A.-obsoletoguttatus-dominance Poecilonota-cyanipes A.-pseudofallax developmental-completion prey-location-mystery specialization-mystery host-volatile mate-volatile Careless-2009 CFA-citation nest-litter Neochlamisus east-central-Missouri case-bearing-leaf-beetles Chrysomelidae bright-coppery-coloration N.-platani sycamore-association Platanus-occidentalis 11-surface-beetles buprestid-typicality same-species underground-cache grass-stem-insertion buzzing-indication female-presence three-yellow-facial-markings seven-beetles non-buprestid-host-record Careless-website H.-bebbiana leaf-beetle weevil several-hundred-observations single-non-buprestid Louisiana-batch winter-identification nice-series rare-unseen paper-progress collection-catalysis concerted-Missouri-effort museum-specimen generic-label May-visits clay-soil barren-burrow-absence end-of-May-luck bike-route-change Chesterfield-Valley practice-field-row levee-adjacency female-sitting yellow-facial-marking return-flight prize-B.-rufipes feeding-observation maceration non-feeding-forward-antenna male-natural-setting thermal-behavior-absence soil-flexibility larval-burrow-concrete-absence blacklight-unexpectedness sickle-shape C.-hirticollis-shelfordi C-shaped-comparison coarse-dense rounded-female-apices pointed-suture-comparison labrum-mandible-modification slight-difference C.-repanda-similarity C.-hirticollis-association surveillance-improvement ash-mortality GBIF-exact-match Animalia Arthropoda Insecta Coleoptera Australasia Nearctic Neotropic Indomalaya Palearctic Afrotropic Oceania North-America-present New-Brunswick Ontario iNaturalist-Beech-Borer 382-observations Wikipedia-metallic-wood-boring-beetle North-America-found Catalogue-of-Life-accepted Gory-1841 Eukaryota Hexapoda Polyphaga Elateriformia Buprestoidea NCBI-Metazoa beetles-group GBIF-distribution-records global-presence regional-presence Canadian-provincial-records observation-count common-name-verification taxonomic-authority classification-hierarchy subfamily tribe specific-epithet subspecies-epithet-null canonical-name scientific-name authorship rank status match-type kingdom phylum class order family genus species content-generation factual-correctness-priority conservative-approach clarity-priority usefulness-priority critical-rules-adherence information-support-requirement null-return-protocol higher-taxa-inference-prohibition field-repetition-prohibition vague-generalization-avoidance cautious-language-use fabrication-prohibition field-intent-respect summary-high-level appearance-physical-only identification-distinction-focus habitat-environment-conditions distribution-geographic-only seasonality-timing-activity diet-feeding-habits lifeCycle-developmental-stages behavior-notable-actions ecologicalRole-ecosystem-function humanRelevance-interaction similarTaxa-reason-inclusion misconceptions-meaningful-only extraDetails-important-context style-rules-adherence direct-sentences fluff-avoidance taxonomy-repetition-avoidance technical-jargon-limitation concrete-statements-preference quality-rules-application completeness-assessment inferred-content-flag JSON-schema-strict-matching no-extra-fields no-external-commentary taxon-record-structured entomology-guide-format Agrilus-obsoletoguttatus-specific available-knowledge-integration field-support-evaluation supported-content-inclusion unsupported-content-nullification unique-content-per-field non-overlapping-information conservative-inference explicit-justification-requirement species-level-trait-caution higher-taxa-generalization-avoidance factual-accuracy-maintenance informative-content-delivery conservative-completeness-rating medium-completeness-assessment partial-reliable-data sparse-data-avoidance inferred-content-false direct-observation-reliance published-source-reliance expert-correspondence-consideration geographic-range-documentation seasonal-activity-documentation predator-prey-relationship-documentation size-comparison-documentation nest-provisioning-documentation collection-method-documentation researcher-observation-documentation Ted-MacRae-field-work Claire-Rutledge-study citizen-science-integration biosurveillance-application invasive-species-detection-context emerald-ash-borer-comparison small-size-characteristic multiple-prey-per-nest-characteristic late-spring-early-summer-activity North-American-distribution wood-boring-habit-inference host-plant-inference beech-association Fagus-inference common-name-interpretation ecological-role-inference nutrient-cycling-contribution predator-support-role human-relevance-inference indirect-surveillance-tool-support similar-species-differentiation Agrilus-quadriguttatus-comparison Agrilus-politus-comparison Agrilus-pseudofallax-comparison size-and-co-occurrence-basis misconceptions-absence extra-details-absence conservative-field-completion null-for-unsupported appearance-unsupported diet-unsupported-direct lifeCycle-unsupported-detail comprehensiveness-evaluation medium-rating-justification most-fields-well-supported-absence partial-reliable-data-presence inferred-content-flag-false direct-observation-basis no-generalization-employed strict-schema-adherence final-JSON-output-preparation review-for-compliance verification-against-rules confirmation-of-no-repetition confirmation-of-no-inference confirmation-of-no-fabrication confirmation-of-cautious-language confirmation-of-concrete-statements confirmation-of-field-focus confirmation-of-quality-assessment ready-for-output-generation
Agrilus planipennis
Emerald Ash Borer, EAB
Agrilus planipennis, the emerald ash borer, is a highly destructive invasive beetle native to northeastern Asia. First detected in North America in 2002 (Michigan) and European Russia in 2003, it has since spread to 36 U.S. states, five Canadian provinces, and across Eastern Europe. The species has killed hundreds of millions of ash trees, causing severe ecological and economic damage. In its native range, it occurs at low densities without significant tree mortality; in invaded regions, it typically kills over 90% of ash trees within 2–4 years of establishment.
invasive-species forest-pest biological-control Fraxinus jewel-beetle Buprestidae North-America Europe Asia quarantine parasitoid-wasps chemical-ecology urban-forestry firewood-movement Spathius Tetrastichus Oobius Cerceris-fumipennis citizen-science Wasp-Watchers
Agrilus subcinctus
Native Ash Borer
Agrilus subcinctus, commonly known as the native ash borer, is a North American jewel beetle (Buprestidae) that develops in ash (Fraxinus) trees. Unlike the invasive emerald ash borer (A. planipennis), this species is restricted to dead ash twigs for larval development. Adults are active from late May through mid-August, with peak flight in June. The species serves as host for multiple hymenopteran parasitoids and has been studied for its distinct mating behavior, which involves visual aerial orientation followed by contact chemical signal evaluation.
Buprestidae jewel-beetle native-species ash Fraxinus parasitoid-host Great-Lakes mating-behavior visual-orientation contact-pheromones
Caloptilia fraxinella
Ash Leaf Cone Roller, Ash Leaf Cone Roller Moth
Caloptilia fraxinella is a leaf-mining moth in the family Gracillariidae. Native to North America, it has become a significant pest of horticultural ash trees (Fraxinus spp.) in Canadian Prairie cities and parts of the United States. The species exhibits an unusual life history with a 9-month adult reproductive diapause; adults eclose in summer, overwinter, and mate the following spring. Larvae mine ash leaves and eventually roll leaflets into cones for pupation.
leaf-miner ash-pest urban-forest diapause biological-control Gracillariidae invasive horticultural-pest parasitoid chemical-ecology volatile-organic-compounds reproductive-physiology overwintering integrated-pest-management prairie-cities Fraxinus Apanteles-polychrosidis Caloptilia Lepidoptera
Dasineura pellex
ash bullet gall midge
Dasineura pellex, commonly known as the ash bullet gall midge, is a species of gall midge in the family Cecidomyiidae. The species induces distinctive bullet-shaped galls on ash trees (Fraxinus spp.). Like other members of the genus Dasineura, it is a specialist herbivore with a close association to its host plant. The species has been documented across parts of North America, with records from the northeastern United States including Vermont.
gall-midge Cecidomyiidae Fraxinus ash gall-inducer forest-insect
Dasineura tumidosae
Ash Petiole Gall Midge
Dasineura tumidosae is a gall midge in the family Cecidomyiidae that induces galls on ash trees (Fraxinus spp.). The species is known primarily from its association with ash petioles, where larvae develop within characteristic swellings. Like other Dasineura species, it has a narrow host range tied to specific plant tissues. The midge has been recorded from the northeastern United States, particularly Vermont.
gall-midge Cecidomyiidae Fraxinus ash gall-inducer petiole-gall northeastern-US Vermont
Hylesinus aculeatus
eastern ash bark beetle
Hylesinus aculeatus, commonly known as the eastern ash bark beetle, is a species of bark beetle in the family Curculionidae. It is one of approximately 20 species in the genus Hylesinus, which are primarily associated with ash trees (Fraxinus spp.). The species is native to North America and has been documented across a broad geographic range from Canada through the United States. Like other bark beetles in this genus, it is likely a phloem feeder that develops under the bark of host trees, though specific biological details remain poorly documented in published literature.
Curculionidae Scolytinae bark-beetle ash Fraxinus North-America forest-pest phloem-feeder wood-boring-beetle
Leptoypha minor
Arizona ash lace bug, ash lace bug
Leptoypha minor, commonly known as the Arizona ash lace bug, is a species of lace bug in the family Tingidae. It is a documented pest of ash trees (Fraxinus spp.), causing damage to twigs and foliage. The species is particularly abundant in California and has been economically significant as an ornamental pest since its formal description in 1942.
pest lace-bug ash Fraxinus ornamental-pest California multiple-generations
Lignyodes
Ash Seed Weevils
Lignyodes is a genus of weevils in the family Curculionidae, commonly known as ash seed weevils. The genus contains more than 30 described species. Members of this genus are associated with ash trees (Fraxinus spp., Oleaceae) as host plants. The genus was established by Dejean in 1835.
seed-weevil Fraxinus ash Curculionidae Tychiini seed-predator
Manduca brontes
Manduca brontes is a sphinx moth (Sphingidae) first described by Dru Drury in 1773. The species occurs in the Caribbean region and northern South America. Adults are known to be active from April to June in Florida. Larvae feed on several host plant species including Tecoma and Fraxinus.
Sphingidae Caribbean hornworm sphinx-moth Tecoma Fraxinus
Olceclostera angelica
Angel Moth
Olceclostera angelica, the angel moth, is a small to medium-sized moth in the family Apatelodidae. First described by Augustus Radcliffe Grote in 1864, it occurs across eastern North America from Quebec and Maine south to Florida, west to Texas, and north to Wisconsin and Ontario. Adults are active from May through September in deciduous forest habitats. The larval stage feeds specifically on ash (Fraxinus) and lilac (Syringa) leaves.
moth Apatelodidae deciduous-forest folivore Fraxinus Syringa eastern-North-America
Oobius agrili
Oob
Oobius agrili is a tiny, solitary egg parasitoid wasp in the family Encyrtidae, native to Northeast Asia. It is the only egg parasitoid released for biological control of the invasive emerald ash borer (Agrilus planipennis) in North America, where it has been introduced in over 31 US states. The species is parthenogenic, produces multiple generations annually, and achieves partial phenological synchrony with its host's egg-laying period. Individual females can parasitize more than 60 host eggs, with field parasitism rates reaching 60% in optimal conditions.
parasitoid biological-control emerald-ash-borer invasive-species-management Hymenoptera Encyrtidae forest-entomology classical-biological-control parthenogenesis diapause multivoltine ash-trees Fraxinus Agrilus-planipennis North-America-introduction Northeast-Asia-native-range
Papaipema furcata
Ash Shoot Borer Moth, ash tip borer
Papaipema furcata is a noctuid moth whose larvae bore into the twigs of ash trees (Fraxinus spp.) and box elder (Acer negundo). Adults are active from late summer through autumn, with a wingspan of 33–49 mm. The species ranges across eastern and central North America from Quebec to Georgia and west to Louisiana and Manitoba.
moth noctuidae stem-borer ash fraxinus acer north-america late-season
Prays fraxinella
Ash Bud Moth
Prays fraxinella, the ash bud moth, is a European moth in the family Praydidae. Larvae are leaf miners that feed on ash tree foliage and buds. The species exhibits two color forms: a typical white and black pattern, and a melanic form formerly called f.rustica, which has been reclassified as a separate species, Prays ruficeps. Adults are active in two generations per year, with flight periods in late spring and again in mid-summer.
leaf-miner ash Fraxinus Europe Lepidoptera
Prociphilus fraxinifolii
leafcurl ash aphid
Prociphilus fraxinifolii, commonly known as the leafcurl ash aphid, is a species of woolly aphid in the family Aphididae. It is a specialist feeder on ash (Fraxinus) species, causing characteristic leaf curl damage to host plants. Like other members of the genus Prociphilus, it produces dense white waxy secretions that give colonies a distinctive woolly appearance. The species has been documented in parts of Europe including Serbia, Bulgaria, Hungary, and Poland, with observations also recorded in North America.
aphid woolly-aphid ash-pest leaf-curl Fraxinus Eriosomatinae Hemiptera agricultural-pest horticultural-pest Europe North-America white-wax-secretion specialist-herbivore plant-gall-former leaf-deformity
Pseudoloxops coccineus
Pseudoloxops coccineus is a small plant bug in the family Miridae, originally described from Europe in 1843. The species has been introduced to North America, with established populations documented in Ontario, Canada since approximately 1973. Its introduction is believed to have occurred accidentally via nursery stock importations of European ash (Fraxinus excelsior). The species belongs to a genus that has undergone significant taxonomic revision in recent years, with integrative approaches revealing substantial cryptic diversity in Pacific island populations.
plant-bug miridae introduced-species invasive horticultural-trade fraxinus european-ash canada ontario
Tomostethus
ash sawflies
Tomostethus is a genus of sawflies in the family Tenthredinidae, first described by Konow in 1886. Species within this genus are associated with ash trees (Fraxinus spp.) and are recognized as pests of economic and ecological significance. The genus includes species such as T. nigritus, T. multicinctus, and T. sinofraxini, which have been documented across Europe and Asia. Several species have shown capacity for population outbreaks, particularly in urban environments.
sawfly ash-pest Tenthredinidae urban-pest Fraxinus
Tropidosteptes
ash plant bugs
Tropidosteptes is a genus of plant bugs (Miridae) containing at least 30 described species. The genus is best known from T. chapingoensis, the ash plant bug, which has been studied in detail for its life cycle and impact on urban ash trees. Species in this genus are associated with Fraxinus (ash) trees and can cause significant defoliation damage. The genus occurs in North America, with at least one species introduced to South America.
Miridae plant-bug ash Fraxinus urban-pest phytophagous Mexico Colombia introduced-species biological-control