Woodlice
Guides
Armadillidae
Tropical Pill Woodlice, Pill Bugs
Armadillidae is the largest family of terrestrial isopods (woodlice), comprising approximately 80 genera and 700 species. Members are characterized by their ability to enroll into a complete sphere (conglobation), a defensive behavior that distinguishes them from most other woodlice families. The family exhibits a strongly convex body form adapted for this rolling behavior, with antennae fully enclosed within the sphere when enrolled—a key difference from the superficially similar family Armadillidiidae. Armadillids occupy diverse habitats across tropical and subtropical regions worldwide, with some species adapted to arid environments and others to cave systems.
Cixius yufengi
Cixius yufengi is a species of planthopper in the family Cixiidae, described by Tsaur in 1993. The species belongs to a genus of small, often inconspicuous insects that feed on plant sap. Very little specific information is available about this particular species.
planthopperCixiidaeCalifornia-endemicTsaur-1993AuchenorrhynchaFulgoromorphasap-feederHemipterainsectarthropodHexapodaAnimaliaEukaryotaDelphacoideaCixiinaeCixiiniCixiusspeciesacceptedGBIFCatalogue-of-LifetaxonomydistributionCaliforniaUSAUnited-StatesNorth-Americaendemicinvertebratehemipterantrue-bugbuginsectaanimalhexapodeukaryotearthropodacixius-yufengiyufengiTsaur1993scientific-namecanonical-nameauthorshiprankstatusgenusspecific-epithetclassificationtaxonomy-matchexactkingdomphylumclassorderfamilydistribution-recordsBuglifeendemic-speciesBritish-endemicsIvell's-Sea-AnemoneEdwardsia-ivelliWidewater-LagoonSussexextinctlikely-extinctnot-seen-in-over-forty-years19731983dance-flyPoecilobothrus-majesticusEssex1907Caledonian-PlanthopperCixius-caledonicusnot-seen-for-70-yearsManx-Shearwater-FleaCeratophyllus-fionnus1960sTurk's-Earth-CentipedeNothogeophilus-turkiIsles-of-ScillyIsle-of-Wight1988never-seen-againconservationJames-Harding-MorrisbookBritish-endemic-invertebratesCraig-MacadamCeltic-WoodlouseMetatrichoniscoides-celticusWaleswestern-fringes-of-England1980sChater's-BristletailDilta-chateriiridescentjumping-powers1990sLundy-Cabbage-Flea-BeetlePsylliodes-luridipennisLundy-IslandDevonco-endemismLundy-Cabbageendemic-plantLundy-Cabbage-WeevilCeutorhynchus-contractus-pallipestaxonomic-uncertaintyHorrid-Ground-weaverNothophantes-horridusPlymouthdevelopmentNorthern-February-Red-StoneflyBrachyptera-putataScotlanddrummingabdomen-tappingBritish-Cave-ShrimpNiphargus-glennieiblindghostly-palecavesdamp-rock-fissurestemporary-puddleshumid-cavesrediscoveredprotectedsurvivalconservation-prioritiesglobal-responsibilityevolutionary-twistsecological-intriguehopenatural-heritageBack-from-the-BrinkRSPBBig-Garden-BirdwatchBSBINew-Year-Plant-Huntplantswildlifenaturecommunicationspublic-engagementcampaignsrare-speciesobscure-speciesoverlooked-speciesirreplaceable-specieslocal-wondersglobal-stakesisolationthousands-of-yearsmillions-of-yearsevolutionlandscapesRed-SquirrelHedgehogEuropeshared-speciesnowhere-elseno-backupno-second-chancescelebrationprotectioncherishrecogniseawarenesshabitat-protectionresearchforgotten-creaturesspotlightslipping-through-the-cracksunknown-to-publicrarely-surveyedbarely-hanging-onalready-goneuncomfortable-truthimportant-speciesleast-knownstrangedeeply-unsettlingsole-global-responsibilitylose-them-everywhereorganisationsfighting-to-changeextraordinary-workrarestmost-threatenedentirely-overlookedforgottenevolvedstep-with-Britain's-landscapesfamiliar-speciesshare-with-Europepopulation-overseasreintroducelose-themwrittenjourneyoverlookedirreplaceablefound-nowhere-else-on-Earthcall-to-recogniseprotectuniquely-oursavailable-nowbooksellersspecies-found-nowhere-else-on-Earthpassionate-nature-enthusiastlifelong-loveexploringnatural-worldtrekkingmountainsrare-flowersscouringfenselusive-mothsinvestigatingexotic-invertebrateshothousesfascinationunwaveringprofessional-lifeconservation-sectorhigh-impact-campaignsinspiredEngland's-rarestmost-obscure-speciesmissionBritain-and-Irelandfall-in-love-with-plantsSHAREFacebookLinkedInguest-blogauthorhow-many-speciesfound-only-in-Britainsimple-questioncomprehensive-listresearchingwriting2022referenceburied-in-booksscattered-across-internettucked-awayminds-of-species-expertsresultover-700-speciesat-least-another-100-subspeciesoccur-nowhere-else-on-Earthtotal-global-responsibilityvery-few-peoplename-even-a-single-onestruckmost-irreplaceable-specieslive-or-diedecisions-made-within-our-borderstop-of-conservation-prioritiescelebratedunderstoodset-outtell-their-storiesunique-invertebratesincredibly-fortunateBuglife's-Conservation-Directorfirst-timecompiling-report20-speciesfive-species-of-flyfour-species-of-beetletwo-stonefliesone-eachwoodlousecentipedemillipedefleabristletailspidershrimpsea-anemonetell-storiesgo-out-and-find-thempicked-fivetrack-downbumped-intocouple-moresearched-under-coastal-rocksexquisitepearly-translucenttiny-speciesbarely-2.5mm-longfirst-discoveredknown-only-from-Walesnearbysearched-dampferny-woodlandsalien-lookingastonishing-jumping-powersnamed-new-to-sciencetravelledDevon's-Lundy-Islandtry-and-seeparticularly-rare-pairingonly-known-exampleendemic-beetlepossibly-endemicspend-their-liveswintry-visitsearchelusive-and-threatenedfound-in-just-a-few-siteswithin-the-cityperpetually-under-pressureowes-its-survivaltireless-effortsguided-tourprime-Northern-February-Red-Stoneflyhabitatblew-my-mindmusical-prowessstoneflies-'drum'tapping-their-abdomensslithering-through-tightmuddyunderground-tunnelspersonal-favouriteutterly-gorgeousspends-its-lifechance-searchDevon-coastfirst-sightingalmost-thirty-yearsones-we've-lostimmediately-drawnonly-ever-knownunfortunatelyCraig's-reporthasn't-been-seenover-forty-yearsfirst-collectedlast-seenwithin-a-decadeknowing-this-species-existedgone-foreversadlynot-uncommon-themeendemic-invertebratesdiscoveredhasn't-been-foundover-a-century70-yearsdon't-think-anyonespottedsince-the-1960snot-long-afterfirst-describedfirst-foundstrange-and-uncomfortable-truthsome-of-the-most-important-speciesalso-some-of-the-least-knownby-definitionBritain's-sole-global-responsibilitylose-them-heredespite-that-significancethankfullyorganisations-fightingraising-awarenessrarest-and-most-threatenedotherwise-remainleading-edge-researchevolved-in-stepBritain's-landscapesthousandsunlike-more-familiar-speciesno-population-overseasnowhere-to-reintroducewhy-I-wrote-Endemicutterly-uniquemosseswoodlicebeetlesbuttercupsstories-full-ofright-attentionactionstill-be-savedheld-onpossiblewithin-our-reachall-good-booksellersoverlooked-and-irreplaceableBacks-Goldilocks-ButtercupHeather-StuckeyAbout-the-Authortrekking-up-mountainsscouring-fensinvestigating-exotic-invertebratesfascination-with-wildlifeRSPB's-Big-Garden-BirdwatchBSBI's-New-Year-Plant-HuntBack-from-the-Brink-projectcare-deeplycurrentlyensure-everyoneopportunitywork-with-BSBISHARE-ONCordioniscus
Cordioniscus is a genus of small terrestrial isopods (woodlice) in the family Styloniscidae. The genus contains approximately 18 described species. Members are classified within the suborder Oniscidea, the group containing all fully terrestrial isopods.
Detonella
Detonella is a genus of terrestrial isopods (woodlice) in the family Detonidae, established by Lohmander in 1927. The genus contains at least two described species: Detonella papillicornis from North America and Detonella sachalina from the Russian Far East. These woodlice belong to the suborder Oniscidea, the group containing all fully terrestrial isopods.
Detonidae
woodlice
Detonidae is a family of terrestrial isopods (woodlice) in the order Isopoda, containing at least 3 genera and more than 30 described species. Members of this family are primarily coastal, inhabiting upper intertidal and supratidal zones of rocky shores. The genus Deto includes species with complex taxonomic histories and documented cases of cryptic diversity, such as Deto echinata in southern Africa.
Elumoides
Elumoides is a genus of terrestrial isopods in the family Eubelidae, established by Taiti and Ferrara in 1983. These woodlice belong to the suborder Oniscidea, representing a group of fully terrestrial crustaceans. The genus is part of the diverse isopod fauna found in tropical and subtropical regions.
Eubelidae
Eubelid Pill Woodlice
Eubelidae is a family of woodlice (terrestrial isopods) comprising approximately 50 genera. Members are primarily associated with tropical forest habitats, with the highest diversity concentrated in African tropical forests. The family extends to parts of South-East Asia and the Arabian Peninsula, reflecting a disjunct tropical distribution pattern.
Gabunillo
Gabunillo is a genus of small, eyeless, pigmentless woodlice in the family Armadillidae. Described in 1983 from Gabon, it now includes four species found in Gabon, Brazil, and São Tomé. Members of this genus are characterized by their ability to fully conglobate (roll into a ball), complete absence of eyes, and lack of pseudotracheae. The genus is distinguished from the similar Synarmadillo by cephalon structure, telson shape, and the absence of ventral teeth on thoracic segments.
Haplophthalmus
Haplophthalmus is a genus of small terrestrial isopods (woodlice) in the family Trichoniscidae. The genus contains 47 described species, with several showing restricted geographic ranges that have led to conservation concerns. Two species, H. abbreviatus and H. rhinoceros, are listed as vulnerable on the IUCN Red List due to their limited distributions in the Balkans.
Ligidium
Rock Slaters
Ligidium is a genus of small, forest-dwelling woodlice (terrestrial isopods) in the family Ligiidae, containing approximately 46–68 species with ongoing taxonomic revision. The genus exhibits exceptional genetic differentiation and cryptic diversity, with populations often showing high isolation even over short geographic distances. Species are distributed across temperate regions of the Northern Hemisphere, with notable radiations in Japan, Greece, Turkey, the Caucasus, Central Asia, China, Taiwan, and North America. Ligidium has been proposed as a paleogeological marker organism due to its strong correlation between phylogenetic patterns and regional geological history.
Littorophiloscia
Littorophiloscia is a genus of terrestrial isopods (woodlice) established by Hatch in 1947, containing at least 20 described species. The genus belongs to the family Halophilosciidae (sometimes referenced as Philosciidae in older sources). Species in this genus have been documented as hosts for trichomycete fungi in their hindguts. The genus shows a tropical distribution pattern, with documented occurrences in the Caribbean region.
Oniscidae
sow bugs, woodlice
Oniscidae is a family of terrestrial isopods (woodlice) within the suborder Oniscidea. The family includes the common woodlouse Oniscus asellus and at least six confirmed genera: Oniscus, Oroniscus, Phalloniscus, Rabdoniscus, Rodoniscus, and Sardoniscus. Members are detritivores that contribute to decomposition and nutrient cycling in terrestrial ecosystems. Some species exhibit behavioral sensitivity to environmental contaminants, suggesting potential use as bioindicators.
Philosciidae
Philosciid Woodlice
Philosciidae is a family of terrestrial isopod crustaceans commonly known as woodlice. The family exhibits a cosmopolitan distribution with strong representation in tropical and subtropical regions, particularly in the Americas, Africa, and Oceania. Species diversity is highest in the Neotropics, with numerous genera and species described from Brazil alone. Several species have adapted to cave environments (troglobitic), including notable examples in the genera Benthana and Benthanoides from iron-ore caves in the Amazon biome. The family is less diverse in the Holarctic realm, where only a few species occur.
Porcellio
woodlice
Porcellio is a large genus of terrestrial isopods (woodlice) in the family Porcellionidae, containing approximately 191 described species with a near-cosmopolitan distribution. The genus includes the well-studied common rough woodlouse (Porcellio scaber), which has served as a model organism for research on behavioral ecology, personality traits in invertebrates, and physiological processes such as ecdysis. Most species were described by 1950, though new species continue to be discovered. Members of this genus cannot conglobate (roll into a ball), distinguishing them from pill bugs in related genera.
Rhyscotidae
Rhyscotid woodlice
Rhyscotidae is a family of terrestrial isopods (woodlice) within the suborder Oniscidea. The family comprises two genera, Rhyscotus and Rhyscotoides, and is characterized by small to medium-sized woodlice with typical isopod body plans. Members are found in tropical and subtropical regions, particularly in the Americas. The family was established by Budde-Lund in 1904 and represents a distinct lineage within the Crinocheta clade of terrestrial isopods.
Styloniscidae
Styloniscidae is a family of terrestrial isopods (woodlice) within the suborder Oniscidea. The family comprises at least twelve genera distributed across tropical and subtropical regions, with exceptional diversity documented in Brazilian karst systems. Multiple genera contain troglobitic (exclusively cave-dwelling) species, including Xangoniscus, Cylindroniscus, Pectenoniscus, and Chaimowiczia. These cave-dwelling representatives exhibit classic troglomorphic traits: anophthalmy (eye loss), depigmentation, and elongated appendages.
Trachelipodidae
Trachelipodid Woodlice
Trachelipodidae is a family of terrestrial isopods (woodlice) within the suborder Oniscidea. The family was established by Strouhal in 1953 and is classified within the infraorder Orthogonopoda and parvorder Crinocheta. Members are found across Europe and North America, with documented occurrences in Denmark, Norway, Sweden, the United States (including Vermont), and Romania. The genus Trachelipus is the best-known within the family, with species such as T. trilobatus having been subject to recent distributional studies.
Trichoniscus
pygmy woodlice
Trichoniscus is a genus of small terrestrial isopods (woodlice) containing over 100 species. The genus is notable for including species with both sexual and parthenogenetic reproductive modes, most prominently exemplified by Trichoniscus pusillus, which exists as diploid sexual and triploid asexual forms. These isopods are among the smallest woodlice, typically only a few millimeters in length, and are significant study organisms for research on the evolutionary maintenance of sexual reproduction.
Tylidae
Calloused Beach Pillbugs and Heller's Isopod
Tylidae is a family of terrestrial isopods (woodlice) comprising approximately 27 species, with all but one species placed in the genus Tylos; the sole exception is Helleria brevicornis. Together with Ligiidae, Tylidae represents an early-diverging lineage among woodlouse families. Members are primarily associated with coastal sandy beach habitats in the supralittoral zone.