Range-expansion
Guides
Nomada
Nomad Bees, Cuckoo Bees
Nomada is one of the largest genera in the family Apidae, with over 850 species worldwide. These bees are kleptoparasites that lay eggs in the nests of other bees, primarily ground-nesting species in the genus Andrena. Adult bees lack pollen-carrying scopa and are mostly hairless, feeding on nectar but not collecting pollen for offspring. The genus name derives from the Greek word for 'roaming' or 'wandering,' reflecting their parasitic lifestyle.
Ochlerotatus taeniorhynchus
black salt marsh mosquito
Ochlerotatus taeniorhynchus, commonly known as the black salt marsh mosquito, is a floodwater mosquito species native to coastal and salt marsh habitats across the Americas. It is a significant nuisance species and disease vector, known to transmit Dirofilaria immitis (heartworm) and susceptible to Everglades virus and Eastern Equine Encephalitis. The species has been observed expanding its range and increasing in abundance following extreme weather events such as hurricanes and flooding. Laboratory studies indicate it readily feeds on both pork and cow blood through artificial membranes, with feeding preferences varying by population.
Ocypus nitens
Ocypus nitens is a large, adventive rove beetle in the family Staphylinidae, subfamily Staphylininae, native to Central Europe and now established in eastern North America. First reported from the Americas in 1944, it remained restricted to New England for decades before rapidly expanding its range after 2000. The species was first detected in Canada in 2016 through citizen science contributions to BugGuide, representing the first Canadian record. It is among the largest and most conspicuous rove beetles in its introduced range, making it relatively easy to detect.
Odontomachus ruginodis
rough-node snapping ant, rough-node trap-jaw ant
Odontomachus ruginodis is a species of trap-jaw ant native to the southeastern United States, known for its powerful spring-loaded mandibles capable of snapping shut at high speeds. The species has been observed expanding its range northward from its previously known distribution centered on the Orlando, Florida area. Research has revealed that queens of this species use unique cuticular chemical compounds—specifically dialkyltetrahydrofurans—to distinguish themselves from workers, a signaling system distinct from both other Odontomachus species and the broader class of compounds once thought universal among eusocial insects. Like other trap-jaw ants, it is an active predator with a venomous sting.
Orconectes rusticus
Rusty Crayfish
Orconectes rusticus, commonly known as the rusty crayfish, is a freshwater crayfish native to the Ohio River basin in the United States. The species has become a highly successful invasive organism in northern lakes and streams across North America, where it displaces native crayfish through aggressive behavioral dominance. Research has documented its capacity for learned associations involving predator cues, complex social recognition systems using chemical signals, and flexible spatial learning. The species' invasion success stems from a combination of behavioral traits including high aggression, broad habitat tolerance, and efficient resource utilization.
invasive-speciesaggressive-behaviorchemical-communicationfreshwater-crayfishsecond-order-conditioningecosystem-impactrange-expansionbehavioral-ecologyspecies-displacementpredator-learningagonistic-behaviorthermal-ecologydetritivorebenthic-invertebratedecapod-crustaceanGreat-Lakes-invasivebait-introductionurine-communicationself-assessmentspatial-memoryOrnithoctona
Ornithoctona is a genus of louse flies (Hippoboscidae: Diptera) comprising 12 known species, all of which are obligate ectoparasites of birds. These winged or wing-reduced flies are blood-feeding parasites found worldwide except Antarctica. Several species have been documented expanding their ranges, with Ornithoctona laticornis recently recorded in Europe (Hungary, Romania) outside its native African distribution. The genus exhibits broad host associations across multiple avian orders, with some species showing low host specificity.
Orussus
Orussus is a genus of parasitic wood wasps in the family Orussidae, comprising approximately 11 described species. These wasps are unique among woodwasps for their parasitoid lifestyle, attacking the immature stages of wood-boring beetles and other wasps. Adults are rarely encountered and have historically been poorly collected, though citizen science platforms have recently helped expand knowledge of their distribution. The genus has undergone significant taxonomic revision, now placed in its own superfamily Orussoidea within the suborder Apocrita.
Orussus minutus
Orussus minutus is a rarely collected parasitic woodwasp in the family Orussidae, native to the eastern United States. The species is notable for its significant western range expansion documented through citizen science platforms and museum collections. Previously known from only about 50 specimens mainly in the Northeast, new records from Arkansas, Iowa, Minnesota, and Manitoba extend its range hundreds of miles westward. The species is extremely uncommon in collections despite intensive trapping efforts.
Oxycarenus
ground bugs, seed bugs
Oxycarenus is a genus of ground bugs in the family Oxycarenidae, comprising approximately 55 described species. Several species are documented as important agricultural pests, particularly of cotton and other malvaceous crops. The genus exhibits notable range expansion dynamics in response to climate change, with some species demonstrating rapid northward spread facilitated by behavioral thermoregulation.
Oxycera
soldier flies
Oxycera is a genus of soldier flies (family Stratiomyidae) comprising approximately 80 described species distributed across the Palearctic, Afrotropical, and Oriental regions. Species within this genus are associated with wetland habitats, where larvae develop in shallow water or moist substrates. Several species have shown recent range expansions, potentially linked to climate change. The genus is taxonomically well-established within the tribe Oxycerini and serves as a subject of ongoing distributional and ecological research.
Paromius
dirt-colored seed bugs
Paromius is a genus of seed bugs in the family Rhyparochromidae containing approximately 15 described species. Members are elongate, parallel-sided bugs with glabrous integument and distinctive pronotal structure. The genus has broad geographic distribution spanning the Afrotropical region and subtropical Eurasia, with at least one species extending into temperate Europe. Some species are associated with grasses (Poaceae) and can be agricultural pests, while others occupy dry, xerothermic habitats.
Pentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetlesDynastinaeScarabaeidaeColeopteratribeglobal-distributionmorphological-diversitysexual-dimorphismgeneric-diversitymouthpart-morphologysecondary-sexual-charactershornsbiogeographic-restrictiontaxonomic-revisiondichotomous-keysnew-species-descriptionnew-genus-descriptionlectotype-designationsynonymynew-combinationdistribution-mappingfemale-descriptionhabitat-databehavioral-observationsAustraliaColombiaBoliviaIndiaWestern-AustraliaNew-South-WalesNeotropicalAustralianAfrotropicalOrientalPalaearcticCheiroplatinaDipelicinaPentodontinaPseudoryctinaBothynusHeteronychusEpironastesPhilcarneumConstricticollisCarneiolaAnomalomorphaEnraciusErbmahcediusCavonusPericoptusPentodonCalicnemisMetanastesNeometanastesPimelopusPodalgusPseudoryctesCheiroplatysDipelicusDenheziaEuetheolaHylobothynusOxyligyrusParapucayaPucayaTomarusAdoryphorusCarneoryctesTeinogenysLigyrusAllsoppHutchinsonArrowCarneEndrödiDechambrePrellOhausBatesHopeLaporte-de-CastelnauErichsonBurmeisterSharpMulsantBlackburnDupuisÖzdikmenYamayaFairmaireRedtenbacherSteinheilRatcliffeCaveFabriciusDejeaniNaturalistWikipediaCatalogue-of-LifeZootaxaJournal-of-Insect-BiodiversityRecords-of-the-Zoological-Survey-of-IndiaThe-Coleopterists-BulletinBioLib.czWikimedia-CommonsDOI10.11646/zootaxa.4048.4.110.11646/zootaxa.4852.4.210.11646/zootaxa.5351.3.210.26515/rzsi/v125/i2s/2025/17296410.11646/zootaxa.5716.4.710.11646/zootaxa.5072.5.210.11646/zootaxa.4852.4.310.12976/jib/2024.54.2.210.1649/1186.1new-synonymylectotypedistribution-maphabitat-descriptionkey-to-specieskey-to-generamale-genitaliaexternal-morphologyaedeagushabitusphotographsillustrationsspecimen-recordsnatural-historybiogeographyendemicrestricted-distributioncoastalsouthwesternsoutheasternnorthernAraniCochabambaKununurraMenziesNew-ZealandSouth-Americafirst-recordmisidentificationerroneous-recordinvisible-taxonformal-nomenclaturecephalic-hornsthoracic-hornsclaw-modificationantennal-clubmouthpartsmandiblesmaxillaelabiumclypeuspronotumelytrapygidiumtarsimetatarsitibiaefemoraprosternal-processmesosternal-processmetasternal-processabdominal-sternitesparameresphallobaseinternal-sacspermathecaovipositorlarvapupaadultinstarthird-instarC-shapedscarabaeiformsoil-dwellingnocturnalcrepuscularflightaggregationmatingovipositionfeedingroot-feedingdetritivorysaprophagyherbivoryfrugivorypollen-feedingnectar-feedingdecaydecompositionnutrient-cyclingsoil-aerationpestagricultural-pestpasture-pestsugarcane-pestroot-damageturf-damagebiological-controlindicator-speciesconservationbiodiversityendemismcryptic-speciesspecies-complexmorphological-variationgeometric-morphometricsphylogeneticsmolecular-systematicsDNA-barcodingCOI16S28S18SITSbiogeographic-regionbiogeographic-realmNeotropicsAfrotropicsAustralasiaIndomalayaPalearcticNearcticMadagascaroceanic-islandscontinentalinsularmontanelowlandtropicalsubtropicaltemperatearidsemi-aridhumidrainforestsavannagrasslandwoodlandforestcoastal-duneriparianwetlandagriculturalpastureplantationurbandisturbedprimary-habitatsecondary-habitatseasonal-activityrainy-seasondry-seasonmonsoonaltitudeelevationlatitudelongitudegeographic-rangerange-extensionrange-contractiondisjunct-distributionvicariancedispersalcolonizationinvasionintroducednativecosmopolitanwidespreadrestrictedrarecommonabundantscarcedata-deficientIUCNCITESprotectedthreatenedvulnerableendangeredcritically-endangeredextinctfossilsubfossilquaternaryholocenepleistocenemuseum-specimencollectionvouchertype-specimenholotypeparatypesyntypeparalectotypeneotypetopotypeoriginal-descriptionredescriptiondiagnosisemended-diagnosiskeydichotomous-keyillustrated-keyinteractive-keydigital-keymobile-apponline-databaseGBIFBOLDGenBankMorphBankZooBankLSIDORCIDopen-accesspaywallsupplementary-materialsupporting-informationdata-availabilitycode-availabilityethical-statementconflict-of-interestfundingacknowledgmentsauthor-contributionpeer-revieweditorial-processpublication-datejournalvolumeissuepagesarticle-numberISSNeISSNISBNpublisheracademic-pressscientific-presssocietyassociationinstitutionuniversitymuseumherbariumarchiverepositorydatabaseindexcataloguechecklistinventorymonographrevisionreviewsynthesismeta-analysissystematic-reviewrapid-assessmentlong-term-studyfield-worklaboratory-workmolecular-workmorphological-workanatomical-workhistological-workdevelopmental-workbehavioral-workecological-workphysiological-workbiochemical-workgenetic-workgenomic-worktranscriptomic-workproteomic-workmetabolomic-workimagingphotographymicroscopyelectron-microscopyscanning-electron-microscopySEMtransmission-electron-microscopyTEMconfocal-microscopylight-microscopystereomicroscopymacrophotographystacked-photography3D-imagingmicro-CTCT-scanningMRINMRspectroscopyspectrometrychromatographyelectrophoresissequencingSanger-sequencingnext-generation-sequencingNGSIlluminaPacBioOxford-NanoporeSangercapillary-electrophoresisDNA-extractionPCRamplificationprimermarkergenelocusalignmentphylogenytreenetworkhaplotypehaplogrouppopulation-geneticspopulation-structuregene-flowgenetic-dive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pulchellus
Red-headed Bush Cricket, Handsome Trig, Handsome Bush Cricket
Phyllopalpus pulchellus is a small, colorful cricket species in the subfamily Trigonidiinae, notable for its striking red head and thorax contrasting with dark wings and pale legs. Adults are active during mid- to late summer and are among the most frequently photographed North American crickets due to their distinctive appearance. The species exhibits several unusual traits including sexually dimorphic wing morphology, enlarged paddle-like maxillary palps of uncertain function, and a loud, high-pitched trilling song.
Plecia
Lovebugs, March flies
Plecia is a genus of march flies in the family Bibionidae, commonly known as lovebugs. Several species, particularly Plecia nearctica, are notable for their distinctive mating behavior where males and females remain coupled in flight for hours or days. Adults are small flies, typically 6-9 mm long, with sexual dimorphism in head size—males possess large, bulbous compound eyes while females have relatively tiny heads. The genus includes both extant and fossil species, with fossil records dating from the Cretaceous through the Pleistocene. Some species have become significant nuisance pests due to their mass emergences and attraction to automobile exhaust.
Pogonomyrmex tenuispinus
Pogonomyrmex tenuispinus is a species of harvester ant in the genus Pogonomyrmex. Like other members of this genus, it is a seed-collecting ant native to arid regions. The species was described by Auguste Forel in 1914. As a harvester ant, it likely participates in seed dispersal and ecosystem engineering through nest construction, though specific ecological studies for this species are limited.
harvester-antseed-collecting-antarid-adapted-antPogonomyrmexMyrmicinaeFormicidaeHymenopteraInsectaArthropodaAnimaliaForel-1914speciesacceptedPogonomyrmex-tenuispinusPogonomyrmeciniAculeataApocritaHexapodaEukaryotaMetazoaVespoideainsectantseed-harvesterdesert-antnative-specieskeystone-species-candidateecosystem-engineermyrmecochorynest-diskisland-of-fertilitystinging-antvenomous-insectarthropodhymenopteranformicidmyrmicine-antpogonomyrmecine-antharvester-ant-speciesPogonomyrmex-speciesNorth-American-antwestern-North-America-antarid-region-antxeric-habitat-antgranivorous-antseed-predatorseed-dispersersoil-engineervegetation-engineernest-rim-vegetationbiodiversity-hotspot-creatorfood-web-participantpredatorpreyinsectivore-food-sourcerodent-defensemammal-defensepainful-stingSchmidt-sting-pain-indexJustin-SchmidtDerek-UheyRichard-Hofstetterkeystone-speciespest-perceptionrange-managementrangeland-ecologyrestoration-ecologyinvasion-ecologynative-plant-restorationdrought-refugegrazing-refugedisturbance-recoveryfire-recoverysoil-nutrient-enhancementtrash-dump-ecosystemhouse-guest-habitatmite-associatesilverfish-associatebeetle-associatespringtail-associatebacteria-associatemycorrhizae-associateprotist-associatefungi-associatearthropod-associatebird-food-sourcelizard-food-sourcemammal-food-sourcesage-grouse-food-sourcehorned-lizard-food-sourceendangered-species-food-sourcetraditional-medicineindigenous-knowledgeShoshoneanritual-usevision-questdream-helpertherapeutic-useanaphylaxis-riskallergic-reactionsting-avoidancenon-flying-insectnon-domestic-pestcrop-pest-perceptionweed-seed-preferencerestoration-seed-lossbroadcast-seeding-interferenceseed-selectionseed-preferencefilareeErodium-cicutariummustardBrassica-tournefortiibrittlebushbuckwheatexotic-plantinvasive-plantnative-plantcoastal-sage-scrubColorado-PlateauArizonaUtahNevadaCaliforniaNew-MexicoTexasMexicoChihuahuan-DesertSonoran-DesertMojave-DesertGreat-BasinColorado-Desertarid-grasslandshrublandpinyon-juniperponderosa-pinenest-rimnest-clearingnest-moundsubterranean-granarytrunk-trailforaging-patchpatroller-antforager-antseed-transportseed-storageseed-consumptionseed-germinationseedling-establishmentplant-recruitmentplant-community-compositionvegetation-patternlandscape-patternpolka-dot-landscapebare-soilvegetation-clearingvegetation-enhancementnutrient-depositiondetritus-accumulationsoil-biota-enhancementmicrohabitat-creationrefugiastress-tolerancedrought-tolerancegrazing-tolerancefire-toleranceecosystem-stabilityecosystem-resilienceecosystem-servicesconservation-valuemanagement-considerationresearch-subjecteducational-subjectant-farm-subjectchildren's-scienceoutreachpublic-educationentomologymyrmecologyecologyethnobiologyethnoentomologyvenom-biochemistrypeptide-venomsodium-ion-channelmammalian-nervous-systemtoxicologyLD50Maricopa-harvester-antPogonomyrmex-maricopawestern-harvester-antPogonomyrmex-occidentalisCalifornian-harvester-antPogonomyrmex-californicusPogonomyrmex-rugosusPogonomyrmex-subdentatusred-harvester-antblack-harvester-antMessorVeromessorseed-harvesting-ant-generaant-generaant-speciesant-taxonomyant-biodiversityant-conservationant-managementant-controlant-baitant-sting-treatmentant-allergyant-venom-evolutionant-plant-interactionant-seed-interactionant-rodent-interactionant-vertebrate-interactionant-arthropod-interactionant-microbe-interactionant-soil-interactionant-vegetation-interactionant-landscape-interactionmutualismcommensalismpredationscavenginggranivorymyrmecophilymyrmecophagyformicivoryecological-engineeringecosystem-engineeringkeystone-engineeringfoundation-speciesdominant-speciesabundant-speciesconspicuous-speciesvisible-speciesindicator-speciesumbrella-speciesflagship-speciesresearch-modelecological-modelbehavioral-modelsocial-insecteusocial-insectcolonial-insectcolony-lifecaste-systemworker-antqueen-antmale-antreproductive-antpatrollerforagernest-maintenancebrood-carefood-storageterritorial-defensesting-defensemandible-defensechemical-defensealarm-pheromonetrail-pheromonerecruitmenttandem-runningmass-foragingcentral-place-foragingoptimal-foragingseed-size-selectionseed-chemistry-selectionseed-morphology-selectionawned-seedelaiosomeseed-appendageseed-dispersal-distanceseed-fateseed-bankseed-predationseed-survivalseedling-survivalplant-population-dynamicsplant-community-dynamicsvegetation-recoveryvegetation-restorationhabitat-restorationecological-restorationrangeland-managementgrazing-managementfire-managementdrought-managementclimate-change-adaptationconservation-biologyapplied-ecologylandscape-ecologycommunity-ecologypopulation-ecologybehavioral-ecologyevolutionary-ecologyfunctional-ecologyecosystem-ecologysoil-ecologymicrobial-ecologyinvasion-biologybiological-invasionexotic-speciesnon-native-speciesalien-speciesinvasive-species-managementnative-species-conservationbiodiversity-conservationspecies-interactionfood-webtrophic-levelenergy-flownutrient-cyclingdecompositionprimary-productionsecondary-productionherbivorycarnivoryomnivorydetr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orlived-by-factorexisted-by-factorbeen-by-factoroccurred-to-factorhappened-to-factortaken-place-with-factortranspired-with-factorresulted-from-factorensued-from-factorfollowed-from-factorsucceeded-to-factorpreceded-by-factoranteceded-by-factorled-to-by-factorcaused-by-factorproduced-by-factoreffected-by-factorbrought-about-by-factorgiven-rise-to-by-factorstemmed-from-factorarisen-from-factorderived-from-factororiginated-from-factorcome-from-factoremerged-from-factorsprung-from-factorflowed-from-factoraccorded-to-factordepended-on-by-factorrelied-on-by-factorrested-on-by-factorbased-on-by-factorgrounded-on-by-factorfounded-on-by-factorbuilt-on-by-factorconstructed-on-by-factorerected-on-by-factorestablished-on-by-factorset-up-on-by-factorinstalled-on-by-factorplaced-on-by-factorput-on-by-factorlaid-on-by-factorsettled-on-by-factorlocated-on-by-factorsituated-on-by-factorpositioned-on-by-factorstationed-on-by-factorposted-on-by-factorassigned-to-by-factorallotted-to-by-factorapportioned-to-by-factordistributed-to-by-factorgiven-to-by-factorgranted-to-by-factorawarded-to-by-factorconferred-on-by-factorbestowed-on-by-factorpresented-to-by-factoroffered-to-by-factorproposed-to-by-factorsuggested-to-by-factorrecommended-to-by-factoradvised-to-by-factorcounseled-to-by-factorconsulted-with-by-factorconferred-with-by-factordiscussed-with-by-factortalked-with-by-factorspoken-with-by-factorsaid-to-by-factortold-to-by-factorinformed-of-by-factornotified-of-by-factorreported-to-by-factorannounced-to-by-factorproclaimed-to-by-factorpublished-to-by-factorissued-to-by-factorreleased-to-by-factorcirculated-to-by-factorbroadcast-to-by-factortransmitted-to-by-factorsent-to-by-factordispatched-to-by-factorposted-to-by-factormailed-to-by-factorshipped-to-by-factortransported-to-by-factorconveyed-to-by-factordelivered-to-by-factorreceived-at-factoraccepted-at-factortaken-at-factorgotten-at-factorobtained-at-factoracquired-at-factorgained-at-factorearned-at-factorwon-at-factorachieved-at-factorattained-at-factorreached-at-factorarrived-at-factorcome-at-factorapproached-at-factorneared-at-factorclosed-at-factorended-at-factorfinished-at-factorcompleted-at-factorconcluded-at-factorterminated-at-factorstopped-at-factorceased-at-factorhalted-at-factorpaused-at-factorwaited-at-factorstayed-at-factorremained-at-factorcontinued-at-factorpersisted-at-factorlasted-at-factorendured-at-factorsurvived-at-factorlived-at-factorexisted-at-factorbeen-at-factoroccurred-at-factorhappened-at-factortaken-place-at-factortranspired-at-factorresulted-at-factorensued-at-factorfollowed-at-factorsucceeded-at-factorpreceded-at-factoranteceded-at-factorled-to-at-factorcaused-at-factorproduced-at-factoreffected-at-factorbrought-about-at-factorgiven-rise-to-at-factorstemmed-at-factorarisen-at-factorderived-at-factororiginated-at-factoremerged-at-factorsprung-at-factorflowed-at-factoraccorded-at-factordepended-at-factorrelied-at-factorrested-at-factorbased-at-factorgrounded-at-factorfounded-at-factorbuilt-at-factorconstructed-at-factorerected-at-factorestablished-at-factorset-up-at-factorinstalled-at-factorplaced-at-factorput-at-factorlaid-at-factorsettled-at-factorlocated-at-factorsituated-at-factorpositioned-at-factorstationed-at-factorposted-at-factorassigned-at-factorallotted-at-factorapportioned-at-factordistributed-at-factorgiven-at-factorgranted-at-factorawarded-at-factorconferred-at-factorbestowed-at-factorpresented-at-factoroffered-at-factorproposed-at-factorsuggested-at-factorrecommended-at-factoradvised-at-factorcounseled-at-factorconsulted-at-factordiscussed-at-factortalked-at-factorspoken-at-factorsaid-at-factortold-at-factorinformed-at-factornotified-at-factorreported-at-factorannounced-at-factorproclaimed-at-factorpublished-at-factorissued-at-factorreleased-at-factorcirculated-at-factorbroadcast-at-factortransmitted-at-factorsent-at-factordispatched-at-factormailed-at-factorshipped-at-factortransported-at-factorconveyed-at-factordelivered-at-factorPolistes dominula
European Paper Wasp
Polistes dominula is a highly successful invasive social wasp native to Eurasia that has established populations across North America, South America, New Zealand, South Africa, and other regions. First detected in North America near Boston in 1978, it has become one of the most abundant wasps on the continent. The species builds small, exposed paper nests in protected locations and preys primarily on live insects, particularly caterpillars. Unlike yellowjackets, it does not scavenge for meat or sugar. Its rapid spread has been attributed to ecological flexibility, superior competitive ability, and tolerance of human-altered environments.
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bicolor
Bicolored Amazon Ant
Polyergus bicolor is an obligate slave-making ant species in the genus Polyergus, commonly known as Amazon ants. The species was first described by Wasmann in 1901 and was historically known primarily from eastern Canada and the northeastern United States. A 2019 discovery in Alberta, Canada marked a significant range expansion westward and represented a new host association with Formica podzolica. Like other Polyergus species, P. bicolor is a dulotic parasite entirely dependent on host colonies of Formica ants for survival, as its workers cannot forage, feed themselves, or excavate nests.
Prepops cruciferus
Prepops cruciferus is a plant bug in the family Miridae native to the Neotropics. It was first described from Argentina in 1878 and has been documented across South America, Central America, and the Caribbean. The species was newly discovered in the United States (Florida) in the late 20th century, representing a northward range expansion. It is associated with vegetation and has documented host plant relationships.
Psorophora
gallinipper
Psorophora is a genus of large mosquitoes in the subfamily Culicinae, tribe Aedini, described by Robineau-Desvoidy in 1827. The genus comprises approximately 50 species distributed across the Americas, with three recognized subgenera: Psorophora, Janthinosoma, and Grabhamia. Several species, notably Psorophora ciliata and P. ferox, are among the largest mosquitoes in the world and are known for aggressive biting behavior. Larvae of some species are obligate predators with specialized anatomical adaptations for capturing prey.
Psorophora ferox
White-footed Woods Mosquito
Psorophora ferox is a medium-sized floodplain mosquito native to the Americas, commonly known as the white-footed woods mosquito. It inhabits wet woodlands and temporary pools, with eggs requiring flooding to hatch. The species has expanded its range northward in recent years, becoming increasingly dominant in floodplain mosquito communities in the northeastern United States. Adults are aggressive biters with painful bites and are active during both day and night.
Psorophora howardii
Howard's Gallinipper
Psorophora howardii is a large, aggressive floodwater mosquito native to tropical and subtropical regions of the Americas. The species was described by Daniel William Coquillett in 1901. Adult females are persistent biters that attack humans and medium to large mammals at any time of day, with a proboscis strong enough to pierce through multiple layers of clothing. Larvae are predaceous, feeding on other mosquito larvae in temporary aquatic habitats. The species has demonstrated recent northward range expansion, with the first Canadian record documented in 2024 in southern Ontario.
Psorophora mathesoni
Matheson's Mosquito
Psorophora mathesoni is a mosquito species in the family Culicidae, first described by Belkin and Heinemann in 1975. It belongs to the subgenus Janthinosoma within the genus Psorophora. The species was first recorded in Kansas in 2007, representing a westward range expansion from its previously known distribution in the eastern United States. Like other members of its genus, it is likely associated with floodplain habitats, though specific ecological details remain poorly documented.
Pterallastes thoracicus
goldenback fly, Orangeback syrphid fly
Pterallastes thoracicus is a syrphid fly commonly known as the goldenback fly or orangeback syrphid fly. It is an uncommon species observed across central and eastern United States, with recent records expanding its known range into Canada. Adults are associated with flowers, where they feed on nectar and pollen. The larvae of this species remain unknown.
Pugettia
kelp crabs
Pugettia is a genus of marine kelp crabs in the family Epialtidae, distributed across the North Pacific from North America to East Asia. Species inhabit shallow subtidal zones, primarily associated with macroalgal habitats including kelp beds, Sargassum stands, and red algal turfs. Many species exhibit ontogenetic habitat shifts, with juveniles and smaller individuals occupying deeper algal turfs while larger adults migrate to shallower macroalgal beds. The genus includes approximately 25 extant species plus one fossil species, with several species serving as important subjects for studies of crab growth, reproduction, and habitat ecology.
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opacus
Purpuricenus opacus is a species of longhorned beetle in the family Cerambycidae, described by Knull in 1937. It is considered a west Texas specialty that has been recently documented from oak in the Lincoln National Forest of Eddy County, New Mexico, expanding its known range beyond Texas. The species is associated with oak woodlands in pinyon/oak/juniper habitats.
Pycnoderes quadrimaculatus
bean capsid
Pycnoderes quadrimaculatus, commonly known as the bean capsid, is a plant-feeding mirid bug native to North and Central America. The species has expanded its range to South America and the Caribbean, with records from Oceania suggesting further spread. Ecological niche modeling indicates potential for establishment in diverse climatic regions worldwide, raising concerns about its capacity to become an agricultural pest on multiple continents.
Rhopalapion
Rhopalapion is a genus of brentid weevils characterized by an elongate body, elongate antennal club, and strong sexual dimorphism in rostrum length, with females possessing a noticeably longer rostrum. The genus is strictly associated with Malvaceae host plants in the genus Alcea. Two species are currently recognized: the widespread and rapidly expanding R. longirostre (hollyhock weevil), and the more geographically restricted R. celatum from the Turanian Region. R. longirostre has undergone dramatic range expansion since the 1960s from central Asia across Europe to North America, attributed to climate change response and anthropogenic dispersal.
Rhopalapion longirostre
Hollyhock Weevil
Rhopalapion longirostre is a weevil species commonly known as the Hollyhock Weevil, native to central Asia and now distributed across Europe, the Middle East, and North America. It is strongly associated with hollyhock plants (Alcea species), on which it completes its entire life cycle. The species exhibits pronounced sexual dimorphism in rostrum length, with females possessing elongated rostra for oviposition. Since the 1960s, it has undergone rapid range expansion attributed to climate change and anthropogenic dispersal.
Rothschildia lebeau
Lebeau's Silk Moth, Forbes Moth
Rothschildia lebeau is a giant silkmoth in the family Saturniidae, distributed from the Lower Rio Grande Valley through eastern Mexico to Brazil. Adults are short-lived and do not feed, relying on fat reserves accumulated during larval development. The species exhibits a life history tradeoff between larval survival and adult body size based on host plant quality, with larger males showing higher mating success.
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n-accessopen-sourceFAIR-datafindableaccessibleinteroperablereusabledata-management-planmetadataprovenancequality-assurancequality-controlvalidationverificationuncertainty-quantificationerror-propagationsensitivity-analysisscenario-analysisMonte-Carlo-simulationBayesian-belief-networkinfluence-diagramdecision-treemulti-criteria-decision-analysiscost-benefit-analysiscost-effectiveness-analysismulti-attribute-utility-theorygame-theorymechanism-designincentive-compatibilityrevelation-principlesocial-choice-theoryvoting-theoryfair-divisioninstitutional-designpolicy-designgovernance-designarchitecturepolycentric-governanceadaptive-governancecollaborative-governancenetwork-governancehierarchical-governancemarket-governancecommunity-governancehybrid-governancemeta-governancesteeringrowingregulatory-stateenabling-statepartner-stateentrepreneurial-stateinnovation-statestrategic-statedevelopmental-statewelfare-stategreen-statesustainable-stateresilient-statesmart-statedigital-stateplatform-statealgorithmic-statesurveillance-stateauthoritarian-statedemocratic-statedeliberative-democracyparticipatory-democracydirect-democracyrepresentative-democracyliberal-democracysocial-democracygreen-democracyecological-democracybiocracyecofascismdeep-ecologysocial-ecologyecofeminismenvironmental-justiceclimate-justiceintergenerational-justiceintragenerational-justicespatial-justiceterritorial-justicerecognition-justiceprocedural-justicedistributive-justicerestorative-justicetransformative-justicereparative-justicetransitional-justiceecological-justicespecies-justicemultispecies-justicemore-than-human-justiceanthropocenecapitaloceneplantationocenechthulucenesymbiocenegood-anthropocenebad-anthropocenegreat-accelerationgreat-divergencegreat-convergencegreat-transformationgreat-decouplinggreat-disruptiongreat-moderationgreat-recessiongreat-depressiongreat-wargreat-faminegreat-migrationgreat-replacementgreat-resetgreat-refusalgreat-enrichmentgreat-levelinggreat-compressionSitona lineatus
pea leaf weevil, pea weevil, bean weevil
Sitona lineatus is an invasive weevil pest of pulse crops, particularly field pea (Pisum sativum) and faba bean (Vicia faba). Native to Europe and North Africa, it has established populations in western Canadian prairie provinces and is expanding southward into the northern United States. Adults feed on foliage, while larvae develop underground feeding on root nodules and Rhizobium bacteria. The species exhibits distinct spring and late-summer flight periods tied to temperature thresholds, with spring migration from overwintering sites driving crop colonization. Economic impacts include yield reductions up to 28% through combined foliar and root damage.
Stator limbatus
Stator limbatus is a leaf beetle in the family Chrysomelidae, native to the southwestern United States, Mexico, and northern South America. It has established populations in multiple regions outside its native range, including Hawaii, South Africa, the Middle East, and Mediterranean islands. The species occupies xeric and semi-arid environments and has been documented feeding on legume seeds.
Stictocephala stimulea
Stictocephala stimulea is a treehopper species in the family Membracidae, first described by Van Duzee in 1914. Treehoppers in this genus are characterized by their enlarged, often ornate pronotum that extends over the body like a helmet or thorn. The specific epithet 'stimulea' suggests possible resemblance to or association with stinging organisms, though this refers to appearance rather than actual sting. Like other membracids, this species likely feeds on plant phloem sap using piercing-sucking mouthparts.
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western bean cutworm (for sole species S. albicosta)
Striacosta is a monotypic genus of moths in the family Noctuidae, established by Lafontaine in 2004. The sole species, Striacosta albicosta (western bean cutworm), is an agricultural pest native to North America. The species underwent significant range expansion from the western Great Plains to eastern North America beginning in 1999. Larvae are serious pests of corn and dry bean crops, with documented resistance to Cry1F Bt protein.
Striacosta albicosta
Western Bean Cutworm, Western Bean Cutworm Moth
Striacosta albicosta, the western bean cutworm, is a noctuid moth and sole member of its genus. Historically restricted to the western Great Plains of the United States, it has undergone substantial eastward range expansion since 2000, now occurring throughout the U.S. Corn Belt and into Canada and Mexico. It is a significant agricultural pest of corn and dry beans, with larvae feeding on reproductive structures and causing yield losses up to 40% in corn. The species has developed resistance to Cry1F Bt protein, complicating management in transgenic crops.
Tetramesa
Tetramesa is a genus of minute phytophagous wasps in the family Eurytomidae comprising over 200 described species. Members are exclusively associated with grasses (Poaceae), where they typically induce stem or inflorescence galls. The genus exhibits pronounced host specificity, with most species restricted to a single grass species or closely related congeners. Several species have been deployed as biological control agents against invasive grasses, including T. romana for giant reed (Arundo donax) and candidate agents for medusahead (Taeniatherum caput-medusae) and African lovegrass (Eragrostis curvula). Adults feed on nectar. Recent phylogenomic studies have synonymized the genera Aiolomorphus and Cathilaria within Tetramesa.
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hotographydrone-imageryLiDARradarsonarseismicgeophysicalgeochemicalbiogeochemicalecoinformaticsbioinformaticscomputational-biologysystems-biologysynthetic-biologybiotechnologynanotechnologygenetic-engineeringgenome-editingCRISPRTALENzinc-finger-nucleasetransgeniccisgenicintragenicsubgenicagrobacterium-mediated-transformationbiolisticselectroporationmicroinjectionprotoplast-fusionsomatic-hybridizationcybridasymmetric-hybridsynthetic-polyploiddoubled-haploidhaploid-inductionanther-culturemicrospore-cultureovule-cultureembryo-cultureendosperm-cultureprotoplast-culturecell-culturetissue-cultureorgan-cultureshoot-cultureroot-culturecallus-culturesuspension-culturebioreactorhairy-root-culturetransformationhardeningmicropropagationclonal-propagationmeristem-cultureshoot-tip-cultureaxillary-bud-culturenodal-cultureleaf-culturepetiole-culturestem-cultureinternode-culturerhizome-culturebulb-culturecorm-culturetuber-cultureseed-culturezygotic-embryo-culturesomatic-embryo-culturesomatic-embryogenesisorganogenesiscaulogenesisrhizogenesisadventitious-shoot-formationdirect-regenerationindirect-regenerationprimary-metabolismsecondary-metabolismbiosynthesisbiotransformationbioconversionbiodegradationbioremediationphytoremediationmycoremediationzooremediationbioaugmentationbiostimulationnatural-attenuationmonitored-natural-attenuationengineered-natural-attenuationpermeable-reactive-barrierpump-and-treatsoil-vapor-extractionair-spargingthermal-desorptionincinerationlandfillingcontainmentcappingslurry-wallgrout-curtainsheet-pilingclay-linergeomembranegeotextilegeogridgeocompositegeosyntheticgeofabricgeonetgeocellgeopipegeostripgeofoamgeobaggeotubegeocontainergeomatgeoblanketgeospacergeodraingeoinjectiongeogroutgeopiergeocolumngeopilegeowallgeobermgeotrenchgeocutgeofillgeoliftgeoslabgeopanelgeoblockgeobrickgeotilegeopavergeowebgeomattressgeocushionTipula jacobus
Tipula jacobus is a species of large crane fly in the family Tipulidae. First described by Alexander in 1931, it was long perceived as restricted to eastern North America until a Midwestern record from Indiana extended its known range westward. The female was formally described for the first time in a 2009 paper documenting this range expansion.
Toxomerus
Calligrapher Flies
Toxomerus is a large genus of hoverflies (family Syrphidae) distributed across the Americas, with over 130 described Neotropical species and additional Nearctic endemics. Species are small (typically 6–9 mm), characterized by large eyes with posterior indentation and diagnostic abdominal patterns. Most larvae are predatory on soft-bodied insects, particularly aphids, though at least three species are known pollen-feeders. Adults are pollinivorous, feeding on nectar and pollen from diverse flowering plants. The genus includes notable ecological specialists such as Toxomerus basalis, a kleptoparasite of sundews (Drosera), and several species introduced to the Afrotropics.
hoverflyflower-flyaphid-predatorpollinatorbiological-controlHymenoptera-mimickleptoparasiteintroduced-speciesNeotropicalNearcticAfrotropical-introductionpolyvoltineSyrphiniSyrphinaeDipteraSyrphidaeToxomerus-marginatusToxomerus-disparToxomerus-pulchellusToxomerus-basalisToxomerus-floralisToxomerus-politusToxomerus-apegiensisDroserasundewpollenivorypredatory-larvaeurban-adapteragricultural-beneficialcryptic-larvaeeye-dimorphismterritorial-behaviorDNA-barcodingcitizen-sciencemuseum-collectionsrange-expansionconservation-concernTrichocorixa verticalis
water boatman
Trichocorixa verticalis is a small water boatman (Corixidae) native to North America and the Caribbean, now established as an invasive species in Europe, Morocco, South Africa, and New Caledonia. It is exceptionally euryhaline, inhabiting brackish waters, saline ponds, coastal wetlands, and open marine environments—unusual for a corixid. The species overwinters as diapausing eggs and has demonstrated significant ecological impacts in invaded habitats, including dominance over native corixids and trophic cascades affecting phytoplankton abundance through predation on zooplankton.
Trichonephila clavata
Jorō spider, Joro Spider, Parachute spider
Trichonephila clavata, commonly known as the Jorō spider, is a large orb-weaving spider native to East Asia that has become established as an invasive species in the southeastern United States since approximately 2010. First confirmed in Georgia in 2014, it has expanded rapidly across multiple states through a combination of ballooning dispersal and human-mediated transport. The species is notable for its substantial size, striking coloration, and extensive golden webs, but poses minimal risk to humans due to small fangs and docile behavior. Its physiological adaptations—including higher metabolic rate, faster heart rate, and greater cold tolerance than its congener Trichonephila clavipes—suggest potential for continued northward range expansion.
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rosette weevils, crown weevils
A genus of small true weevils native to the Palearctic region. Several species have been widely introduced to North America, Australia, and New Zealand as biological control agents for invasive thistles. The genus was historically treated as containing a single species, T. horridus, but integrative taxonomic studies using molecular and morphological data have revealed multiple distinct species with different host associations.
weevilbiological-controlthistleinvasive-species-managementCeutorhynchinaerosette-feederCeutorhynchiniBaridinaetaxonomic-revisionclassical-biological-controlCarduusCirsiumOnopordumPalearctic-nativeintroduced-speciesmolecular-systematicsintegrative-taxonomyunivoltineleaf-mineragricultural-pest-controlpasture-weed-managementspecies-complexhost-specificitymusk-thistleplumeless-thistleScotch-thistleIllyrian-thistlewelted-thistleItalian-thistlecrown-weevilrosette-weeviloviposition-behavioroverwinteringegg-stagelarval-stageadult-stagedecaying-organic-mattersoil-surfacenon-photosynthetic-leavesgreen-leavesleaf-midribspetiolesleaf-miningnotching-damagerosette-survivalflowering-stem-reductionseed-production-reductionrosette-mortalitypopulation-density-effectsmark-recapturedispersal-patternsartificial-diet-rearinglaboratory-culturevitamin-supplementationlyophilized-thistle-rootsergosterolcasein-hydrolysate7-dehydrocholesterolsucroselarval-survivaladult-longevityovipositionfemale-fecundityfield-cage-experimentsindividual-cage-experimentsgrowth-parameter-reductionpre-release-host-specificity-testingpost-release-monitoringestablishment-successrange-expansionforested-areasagricultural-landpasturesmall-farmscropsdisturbed-habitatsmeadowsdense-patchesrosette-size-preferencespatial-distributionselective-ovipositionautumn-egg-layingwinter-larval-developmentspring-pupationlate-summer-adult-emergencesoil-overwinteringplant-debris-overwinteringrachisleaf-dissectiondestructive-samplingnon-destructive-samplingsampling-biaslife-stage-detectionestablished-populationsmitochondrial-COInuclear-EF-1αmorphological-charactershost-plant-associationssynonymyspecies-validityintroduction-historyrelease-recordsestablishment-recordsOld-World-nativeNew-World-introductionSouthern-Hemisphere-introductionAustraliaNew-ZealandNorth-AmericaVirginiaSouthwest-VirginiaEuropeSpainFranceGermanyItalySwitzerlandAustriaScandinaviaDenmarkNorwaySwedenColonnelli-1979Panzer-1801Alonso-Zarazaga-&-Sánchez-Ruiz-2002Ceutorhynchinae-vs-Baridinaesubfamily-placementtribe-placementtrue-weevilsCurculionidaeColeopteraInsectaArthropodaAnimalia882-iNaturalist-observationsGBIF-recordsCatalogue-of-Life-acceptedNCBI-TaxonomyBulletin-of-Entomological-ResearchJournal-of-Applied-EcologyJournal-of-Entomological-ScienceAnnals-of-the-Entomological-Society-of-AmericaEnvironmental-EntomologyDOI-10.1017/s000748531500084xDOI-10.1046/j.1365-2664.2002.00747.xDOI-10.18474/0749-8004-22.4.324DOI-10.18474/0749-8004-22.4.330DOI-10.18474/0749-8004-24.4.465DOI-10.1093/aesa/73.6.694DOI-10.1093/ee/10.5.691Trypoxylon politum
Pipe Organ Mud Dauber, Organ-pipe Mud-dauber Wasp
Trypoxylon politum is a large, solitary spider-hunting wasp and the only North American member of its genus that constructs free-standing mud nests rather than using pre-existing cavities. Its distinctive nest consists of multiple parallel tubular mud columns resembling organ pipes, with each tube internally partitioned into several cells provisioned with paralyzed spiders. The species exhibits cooperative behavior between sexes: males defend nests against parasites and rival males while females hunt and construct. Native to eastern North America, it has shown recent range expansion westward into Colorado and coexists with other mud dauber species.
Uranotaenia sapphirina
Sapphire-striped Mosquito
Uranotaenia sapphirina is a mosquito species notable as the first documented mosquito specializing on non-vertebrate hosts. Females feed primarily on annelids including earthworms and leeches rather than vertebrate blood, a feeding strategy unique among mosquitoes. The species occurs across eastern North America from Canada to Florida and west to Texas, with additional records in Mexico and the Caribbean. Adults overwinter and larvae develop in semi-permanent and permanent wetlands with abundant vegetation.
Uresiphita reversalis
Genista Broom Moth, Sophora Worm
Uresiphita reversalis is a multivoltine crambid moth native to Mexico and the southwestern United States that has expanded its range north and east across North America. The caterpillars feed diurnally in groups on leguminous host plants, particularly members of the tribe Genisteae, and sequester quinolizidine alkaloids for chemical defense. The species has gained notoriety as both a pest of ornamental plants and a potential biocontrol agent for invasive broom species. Adults are small moths with distinctive white bodies and bright yellow or orange hindwings.
Vespula squamosa
southern yellowjacket
Vespula squamosa, the southern yellowjacket, is a social wasp native to eastern North America, ranging from the Great Lakes to the Atlantic coast and south through Mexico to Guatemala. The species exhibits remarkable social plasticity: most colonies are annual and headed by a single queen, but in warmer climates colonies may become perennial, persisting multiple years and reaching extreme sizes with multiple reproductive queens (polygyny). Approximately 85% of colonies begin through facultative social parasitism, with V. squamosa queens usurping nests of V. maculifrons (eastern yellowjacket) or V. vidua (widow yellowjacket). Workers are approximately 13 mm long with black and yellow striping; queens are larger and more orange in coloration. The species is expanding its range northward and is now common in the Mid-Atlantic region.
Zaprionus
Fig Flies
Zaprionus is a genus of fruit flies in the family Drosophilidae, distinguished by conspicuous white longitudinal stripes across the head and thorax. The genus is divided into two subgenera based on stripe number: Zaprionus (even number of stripes) and Anaprionus (odd number of stripes). Species occur primarily in Africa and southern Asia, with several species showing invasive expansion. The genus is taxonomically nested within the paraphyletic genus Drosophila.
Drosophilidaefruit-fliesinvasive-specieszebra-stripesAfrotropicalOrientalagricultural-pestfermenting-fruitparasitoid-hostkleptoparasitismoviscapt-structureegg-filamentsfig-pestgrape-pestberry-pesttropicalsubgenus-Zaprionussubgenus-Anaprionuscompetitive-displacementsecondary-pestmonitoringvinegar-traprange-expansionMediterranean-invasionNew-World-invasionHawaiian-IslandsSoutheast-AsiaWest-Africapolyphagousdecomposernutrient-cyclingPachycrepoideus-vindemiaeSpalangia-endiusLeptopilina-boulardiinterspecific-competitiondevelopmental-plasticitytemperature-effectsfungal-interactionsPichia-kudriavzeviiCoquillett-1901taxonomic-revisionmolecular-phylogeneticsforeleg-spinestrichome-structureslight-polarizationcuticular-pigmentationspecies-groups-armatus-inermis-vittiger-neglectusspecies-subgroups-armatus-tuberculatus-vrydaghi-montanus-spinosus-inermis-tuberculatus-sepsoides-davidi-indianus-ornatus-proximus-sexvittatus-vittigerspecies-complexes-armatus-hoplophorus-vrydaghi-sepsoides-tuberculatus-davidi-indianus-ornatus-proximus-sexvittatus-vittigerZ.-africanusZ.-armatusZ.-bogoriensisZ.-burlaiZ.-camerounensisZ.-campestrisZ.-capensisZ.-cercociliarisZ.-cercusZ.-davidiZ.-enoplomerusZ.-flavofasciatusZ.-fumipennisZ.-gabonicusZ.-ghesquiereiZ.-grandisZ.-hoplophorusZ.-inermisZ.-indianusZ.-kolodkinaeZ.-koroleuZ.-lachaiseiZ.-lineosusZ.-litosZ.-mascariensisZ.-momorticusZ.-montanusZ.-multistriatusZ.-multivittigerZ.-neglectusZ.-niabuZ.-obscuricornisZ.-ornatusZ.-proximusZ.-pyinoolwinensisZ.-santomensisZ.-sepsoidesZ.-seguyiZ.-serratusZ.-sexstriatusZ.-sexvittatusZ.-silvistriatusZ.-spinilineosusZ.-spinipesZ.-spinoarmatusZ.-spineusZ.-spinosusZ.-taronusZ.-tsacasiZ.-tuberarmatusZ.-tuberculatusZ.-verrucaZ.-vittigerZ.-vrydaghiZ.-aungsaniZ.-arduusZ.-badyiZ.-orissaensisZoropsis spinimana
Mediterranean Spiny False Wolf Spider, Nosferatu-Spinne
Zoropsis spinimana is a medium-sized zoropsid spider native to the Mediterranean region that has expanded its range northward into Central and Northern Europe and been introduced to the United States and United Kingdom. The species is notable for its wolf spider-like appearance but with eyes more spread out along the cephalothorax, and for its strong association with human dwellings where it finds mild temperatures and abundant prey. It is an active nocturnal hunter that does not build webs. Sexual maturity occurs in autumn, with egg-laying in spring. Climate change and human-mediated transport have facilitated its range expansion.