Conservation

Guides

Pentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetles Dynastinae Scarabaeidae Coleoptera tribe global-distribution morphological-diversity sexual-dimorphism generic-diversity mouthpart-morphology secondary-sexual-characters horns biogeographic-restriction taxonomic-revision dichotomous-keys new-species-description new-genus-description lectotype-designation synonymy new-combination distribution-mapping female-description habitat-data behavioral-observations Australia Colombia Bolivia India Western-Australia New-South-Wales Neotropical Australian Afrotropical Oriental Palaearctic Cheiroplatina Dipelicina Pentodontina Pseudoryctina Bothynus Heteronychus Epironastes Philcarneum Constricticollis Carneiola Anomalomorpha Enracius Erbmahcedius Cavonus Pericoptus Pentodon Calicnemis Metanastes Neometanastes Pimelopus Podalgus Pseudoryctes Cheiroplatys Dipelicus Denhezia Euetheola Hylobothynus Oxyligyrus Parapucaya Pucaya Tomarus Adoryphorus Carneoryctes Teinogenys Ligyrus Allsopp Hutchinson Arrow Carne Endrödi Dechambre Prell Ohaus Bates Hope Laporte-de-Castelnau Erichson Burmeister Sharp Mulsant Blackburn Dupuis Özdikmen Yamaya Fairmaire Redtenbacher Steinheil Ratcliffe Cave Fabricius Dejean iNaturalist Wikipedia Catalogue-of-Life Zootaxa Journal-of-Insect-Biodiversity Records-of-the-Zoological-Survey-of-India The-Coleopterists-Bulletin BioLib.cz Wikimedia-Commons DOI 10.11646/zootaxa.4048.4.1 10.11646/zootaxa.4852.4.2 10.11646/zootaxa.5351.3.2 10.26515/rzsi/v125/i2s/2025/172964 10.11646/zootaxa.5716.4.7 10.11646/zootaxa.5072.5.2 10.11646/zootaxa.4852.4.3 10.12976/jib/2024.54.2.2 10.1649/1186.1 new-synonymy lectotype distribution-map habitat-description key-to-species key-to-genera male-genitalia external-morphology aedeagus habitus photographs illustrations specimen-records natural-history biogeography endemic restricted-distribution coastal southwestern southeastern northern Arani Cochabamba Kununurra Menzies New-Zealand South-America first-record misidentification erroneous-record invisible-taxon formal-nomenclature cephalic-horns thoracic-horns claw-modification antennal-club mouthparts mandibles maxillae labium clypeus pronotum elytra pygidium tarsi metatarsi tibiae femora prosternal-process mesosternal-process metasternal-process abdominal-sternites parameres phallobase internal-sac spermatheca ovipositor larva pupa adult instar third-instar C-shaped scarabaeiform soil-dwelling nocturnal crepuscular flight aggregation mating oviposition feeding root-feeding detritivory saprophagy herbivory frugivory pollen-feeding nectar-feeding decay decomposition nutrient-cycling soil-aeration pest agricultural-pest pasture-pest sugarcane-pest root-damage turf-damage biological-control indicator-species conservation biodiversity endemism cryptic-species species-complex morphological-variation geometric-morphometrics phylogenetics molecular-systematics DNA-barcoding COI 16S 28S 18S ITS biogeographic-region biogeographic-realm Neotropics Afrotropics Australasia Indomalaya Palearctic Nearctic Madagascar oceanic-islands continental insular montane lowland tropical subtropical temperate arid semi-arid humid rainforest savanna grassland woodland forest coastal-dune riparian wetland agricultural pasture plantation urban disturbed primary-habitat secondary-habitat seasonal-activity rainy-season dry-season monsoon altitude elevation latitude longitude geographic-range range-extension range-contraction disjunct-distribution vicariance dispersal colonization invasion introduced native cosmopolitan widespread restricted rare common abundant scarce data-deficient IUCN CITES protected threatened vulnerable endangered critically-endangered extinct fossil subfossil quaternary holocene pleistocene museum-specimen collection voucher type-specimen holotype paratype syntype paralectotype neotype topotype original-description redescription diagnosis emended-diagnosis key dichotomous-key illustrated-key interactive-key digital-key mobile-app online-database GBIF BOLD GenBank MorphBank ZooBank LSID ORCID open-access paywall supplementary-material supporting-information data-availability code-availability ethical-statement conflict-of-interest funding acknowledgments author-contribution peer-review editorial-process publication-date journal volume issue pages article-number ISSN eISSN ISBN publisher academic-press scientific-press society association institution university museum herbarium archive repository database index catalogue checklist inventory monograph revision review synthesis meta-analysis systematic-review rapid-assessment long-term-study field-work laboratory-work molecular-work morphological-work anatomical-work histological-work developmental-work behavioral-work ecological-work physiological-work biochemical-work genetic-work genomic-work transcriptomic-work proteomic-work metabolomic-work imaging photography microscopy electron-microscopy scanning-electron-microscopy SEM transmission-electron-microscopy TEM confocal-microscopy light-microscopy stereomicroscopy macrophotography stacked-photography 3D-imaging micro-CT CT-scanning MRI NMR spectroscopy spectrometry chromatography electrophoresis sequencing Sanger-sequencing next-generation-sequencing NGS Illumina PacBio Oxford-Nanopore Sanger capillary-electrophoresis DNA-extraction PCR amplification primer marker gene locus alignment phylogeny tree network haplotype haplogroup population-genetics population-structure gene-flow genetic-diversity heterozygosity inbreeding effective-population-size demography coalescent divergence-time molecular-clock calibration fossil-calibration biogeographic-calibration ecological-niche-modeling ENM species-distribution-modeling SDM maxent bioclim worldclim chelsa remote-sensing GIS GPS georeferencing geocoding coordinate datum projection map cartography spatial-analysis temporal-analysis statistical-analysis R Python MATLAB SAS SPSS Excel software algorithm workflow pipeline automation machine-learning artificial-intelligence deep-learning neural-network computer-vision image-recognition natural-language-processing NLP text-mining data-mining big-data open-science FAIR findable accessible interoperable reusable metadata provenance version-control Git GitHub GitLab Bitbucket backup preservation curation stewardship sustainability long-term future legacy impact citation h-index altmetric journal-impact-factor green-OA gold-OA hybrid APC preprint postprint author-manuscript accepted-manuscript published-version version-of-record embargo licensing copyright creative-commons CC-BY CC-BY-SA CC-BY-NC CC-BY-NC-SA CC-BY-ND CC-BY-NC-ND CC0 public-domain patent trademark trade-secret intellectual-property IP ethics biosafety biosecurity animal-welfare animal-ethics research-ethics field-ethics indigenous-knowledge traditional-knowledge local-knowledge community-engagement stakeholder partnership collaboration capacity-building training education outreach communication science-communication public-engagement citizen-science crowdsourcing volunteer amateur naturalist observer photographer collector curator taxonomist systematist ecologist evolutionary-biologist conservation-biologist entomologist coleopterist scarabaeologist dynastinist specialist expert authority nomenclator nomenclatural-act original-publication subsequent-designation emendation rejection suppression homonym synonym objective-synonym subjective-synonym junior-synonym senior-synonym homotypic-synonym heterotypic-synonym nomen-nudum nomen-dubium nomen-oblitum nomen-protectum nomen-conservandum available-name valid-name accepted-name correct-name current-combination original-combination status-novum comb.-nov.syn.-nov.sp.-nov.gen.-nov.subgen.-nov.subsp.-nov.nom.-nov.nom.-nud.nom.-dub.incertae-sedis unplaced unassigned subtribe genus subgenus species subspecies variety form morph ecotype population stock strain breed cultivar clone lineage biotype karyotype chromosome genome transcriptome proteome metabolome phenome morphome anatomical morphological meristic morphometric allometric ontogenetic developmental life-history reproductive behavioral ecological physiological biochemical genetic genomic evolutionary phylogenetic phylogeographic biogeographic historical paleontological archaeological geological climatological environmental applied economic medical veterinary forestry horticultural industrial pollution climate-change global-warming habitat-loss fragmentation deforestation agricultural-intensification urbanization invasive-species pathogen parasite predator competition mutualism symbiosis commensalism amensalism neutralism facilitation inhibition trophic-cascade food-web food-chain energy-flow ecosystem-function ecosystem-service provisioning regulating supporting cultural alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance rarefaction extrapolation asymptotic non-asymptotic sample sampling survey monitoring assessment evaluation indicator bioindicator flagship umbrella keystone ecosystem-engineer foundation-species dominant subordinate frequent occasional accidental vagrant migrant resident breeding non-breeding wintering summering migration nomadism irruption establishment naturalization spreading range-expansion range-shift extirpation extinction local-extinction global-extinction functional-extinction ecological-extinction pseudextinction Lazarus-taxon Elvis-taxon zombie-taxon living-fossil relict indigenous autochthonous allochthonous exotic alien invasive naturalized casual escaped cultivated ornamental pet food medicine fiber fuel timber fodder pollinator pest-control seed-dispersal soil-formation erosion-control water-purification air-purification carbon-sequestration climate-regulation disease-regulation pest-regulation flood-regulation storm-protection recreation tourism aesthetic spiritual cultural-heritage research inspiration existence-value option-value bequest-value intrinsic-value anthropocentric biocentric ecocentric utilitarian instrumental relational intrinsic inherent absolute conditional responsibility care respect reverence wonder curiosity knowledge understanding wisdom science natural-philosophy biology zoology entomology coleopterology scarabaeology dynastinology systematics taxonomy nomenclature classification evolution ecology behavior paleontology conservation-biology environmental-science agricultural-science forestry-science veterinary-science medical-science public-health one-health planetary-health ecosystem-health biodiversity-health species-health population-health individual-health genetic-health environmental-health social-health economic-health political-health cultural-health spiritual-health holistic-health integrated-health sustainable-development sustainable-use sustainable-management adaptive-management precautionary-principle ecosystem-approach landscape-approach seascape-approach connectivity-conservation corridor buffer-zone protected-area national-park nature-reserve wildlife-refuge wilderness-area world-heritage-site biosphere-reserve Ramsar-site Important-Bird-Area Key-Biodiversity-Area Alliance-for-Zero-Extinction-site conservation-priority hotspot crisis-ecoregion global-200 last-of-the-wild human-footprint cumulative-impact threat-index vulnerability-index adaptive-capacity exposure sensitivity resilience resistance recovery restoration rehabilitation reintroduction translocation ex-situ in-situ captive-breeding botanic-garden zoo gene-bank seed-bank tissue-bank sperm-bank oocyte-bank embryo-bank DNA-bank frozen-zoo ark insurance safety-net de-extinction genetic-rescue genetic-restoration genetic-augmentation genetic-management population-management metapopulation source-sink patch matrix landscape seascape ecosystem biome ecoregion province zone region district site locality habitat microhabitat niche ecological-niche fundamental-niche realized-niche trophic-niche spatial-niche temporal-niche biotic-niche abiotic-niche multidimensional-niche n-dimensional-niche Hutchinsonian-niche Grinnellian-niche Eltonian-niche resource requirement limitation stress disturbance perturbation fluctuation variability heterogeneity complexity diversity redundancy stability persistence adaptation acclimation plasticity evolvability heritability selection drift flow mutation recombination speciation coalescence divergence convergence parallelism homoplasy analogy homology synapomorphy symplesiomorphy autapomorphy apomorphy plesiomorphy derived ancestral primitive advanced basal crown stem node branch clade grade subfamily family superfamily infraorder suborder order superorder infraclass subclass class superclass subphylum phylum superphylum kingdom domain life organism individual etc.
Phaleria
Mahkota Dewa (for P. macrocarpa)
Phaleria is a genus of approximately 25 species of flowering plants in the family Thymelaeaceae. Species range from Sri Lanka through Malesia, Papuasia, northern and eastern Australia, and islands of the western Pacific. The genus is valued for medicinal applications, with Phaleria macrocarpa particularly noted for producing agarwood and containing bioactive compounds with hepatoprotective, anti-inflammatory, and anti-proliferative properties. Phaleria nisidai holds traditional importance in Palau as a medicinal panacea.
medicinal-plants Thymelaeaceae Southeast-Asia agarwood hepatoprotective conservation
Philotiella
Philotiella is a genus of small 'blue' butterflies in the family Lycaenidae, subfamily Polyommatinae, native to western North America. The genus contains two recognized species: Philotiella leona (Leona's Blue), described in 2000 and considered imperiled, and Philotiella speciosa (small blue), described in 1877. Research on P. speciosa indicates that diapause intensity varies with elevation and correlates with host plant bloom periods. Philotiella leona has been the subject of population biology and life history studies in south-central Oregon due to its conservation concern.
Lycaenidae Polyommatinae western-North-America diapause imperiled-species conservation
Photinus consanguineus
double cousin firefly
Photinus consanguineus, commonly known as the double cousin firefly, is a species of firefly in the genus Photinus found in eastern North America. Like other Photinus species, adults produce bioluminescent flashes through a chemical reaction involving luciferin and luciferase. Males fly and emit characteristic flash patterns to locate sedentary females, which respond with their own flashes to facilitate mating. The species faces threats from light pollution, habitat destruction, and lawn insecticides.
firefly bioluminescence Coleoptera Lampyridae eastern-North-America light-pollution conservation nocturnal mating-signals
Photinus knulli
Fall Southwest Firefly
Photinus knulli is a North American firefly species notable for its collective flash synchrony in sparse, low-density swarms. Males produce flash triplets lasting approximately one second, with phrases separated by 3–5 seconds. Synchrony occurs in periodic bursts with rapid accretion and quick decay, a pattern similar to but faster than the eastern Photinus carolinus. The species inhabits riparian areas in the American Southwest, particularly Arizona's Sky Island regions, and is considered Vulnerable by IUCN due to population decline from habitat degradation.
firefly bioluminescence synchrony collective-behavior Sky-Islands Arizona Lampyridae courtship conservation Vulnerable
Plebejus icarioides
Melissa blue, Fender's blue
Plebejus icarioides is a species of lycaenid butterfly commonly known as the Melissa blue. The species has been observed colonizing alfalfa in the Great Basin region, representing a notable host range expansion. One subspecies, P. icarioides fenderi (Fender's blue), is federally endangered in the United States and has been the focus of intensive conservation research. The species exhibits ant tending behavior, with caterpillars producing sugar secretions that ants collect in exchange for protection from natural enemies.
butterfly lycaenid conservation ant-mutualism endangered-subspecies habitat-restoration prescribed-fire alfalfa-colonization Great-Basin western-North-America
Plebejus icarioides missionensis
Plebejus icarioides missionensis is a subspecies of the Mission blue butterfly, a federally endangered lycaenid butterfly endemic to a restricted range in coastal California. The subspecies is part of a complex that has been subject to taxonomic revision, with some authorities placing it in the genus Icaricia. Its habitat consists of coastal grasslands and scrub where its larval host plant, silver lupine (Lupinus albicaulis), occurs. The subspecies has been impacted by urban development, invasive plants, and fire suppression.
endangered conservation California-endemic lycaenidae host-plant-specialist urban-wildlife prescribed-fire-management
Plebejus icarioides parapheres
Plebejus icarioides parapheres is a subspecies of the Boisduval's blue butterfly complex, originally described from California. It is currently treated as a synonym of Icaricia icarioides parapheres in some taxonomic databases. The species group is associated with leguminous host plants and has been subject to conservation research due to habitat loss pressures affecting related subspecies.
Lycaenidae Polyommatinae blue-butterfly California-endemic synonym conservation legume-feeder
Plebejus melissa samuelis
Karner blue butterfly
Plebejus melissa samuelis, commonly known as the Karner blue butterfly, is a federally endangered subspecies first described by Vladimir Nabokov in 1944 from Karner, New York. It is highly specialized, with larvae feeding exclusively on wild lupine (Lupinus perennis) and inhabiting dry sandy barrens ecosystems. The subspecies produces two generations annually and has become a flagship for conservation efforts, particularly in Wisconsin where a pioneering statewide Habitat Conservation Plan was developed.
endangered conservation pollinator barrens-ecosystem habitat-specialist flagship-species Nabokov
Plebejus optilete
Cranberry Blue
Plebejus optilete, commonly known as the Cranberry Blue, is a montane lycaenid butterfly species found in European mountain systems. The species has a restricted distribution in Slovenia, limited to six separate sites on the crests of the western Pohorje Mountains. It is associated with silicate grasslands containing abundant Vaccinium species, which likely serve as larval host plants. Population monitoring indicates declining abundance across most sites, with habitat degradation from overgrazing, inappropriate land use, and tourism development posing significant threats.
Lycaenidae montane-butterfly Slovenia conservation Vaccinium-associated endemic-population
Plectrocnemia
tube maker caddisflies
Plectrocnemia is a genus of tube maker caddisflies in the family Polycentropodidae comprising more than 120 described species. Larvae are aquatic predators that construct silken capture nets to intercept prey. The genus has been extensively studied for its larval silk production, vibration-mediated predatory behavior, and population genetics. Species occur across Europe and into western Asia, with detailed biological information available for several well-studied species including P. conspersa and P. brevis.
Trichoptera caddisfly aquatic-insect predator silk bioindicator net-spinner vibration-detection population-genetics Europe running-water larvae Plectrocnemia-conspersa Plectrocnemia-brevis Plectrocnemia-renetta Plectrocnemia-latissima genome silk-fibroin kin-structure dispersal egg-mass colonial-net oxygen-requirements Caucasus Britain Greece Turkey Cyprus Vermont freshwater stream river spring predatory-behavior vibration-frequency setae-morphology larval-identification-key Oxford-Nanopore-sequencing BUSCO-completeness L-chain-fibroin neighborhood-population-size patchy-recruitment-hypothesis genetic-relatedness microsatellite oviposition hot-spots first-instar pupation mandible-capture orientation-behavior building-behavior prey-capture Chironomidae Oligochaeta substrate-borne-vibration silken-tube tube-maker Polycentropodidae Stephens-1836 more-than-120-species genome-assembly nutrient-cycling ecosystem-services industrial-interest phylogenomics comparative-genomics genome-size contiguity polishing Illumina Nanopore draft-genome annotated-genome Hydropsyche-tenuis spatial-genetic-structure colonization gene-flow genetic-drift dispersal-distance flight temporary-populations permanent-populations habitat-patches suitable-habitat ecological-niche case-making-behavior larval-case silk-secretion protein-component genomic-region gene-cluster genomic-resources high-quality-genome shortest-genome variable-quality published-genomes insect-order speciose industrial-application biomaterial natural-material biomimicry conservation water-quality-monitoring environmental-indicator clean-water oxygen-concentration northern-slopes Central-Caucasus rivers-and-streams biology aspects-of-biology reported inhabits final-instar diagnostic-features illustrated discriminatory-matrix Greek-species zoogeography reported-from key-to-larvae revised-key notes-on previously-unknown-larva distinguishes other-British-species larval-habitat adult-identification genetic-differentiation sites population-sizes short-range-trend greater-distances evolutionary-processes small-scales number-of-generations found-small-populations grow-and-exchange-genes larger-scales substantial-gaps regions colonisation-events genetic-patterns last-colonised ecological-studies dynamics persistence-and-spread central-to Martynov-1913 Malicky-1975 Curtis-1834 McLachlan Curtis Navas genus-Stephens-1836 family-Polycentropodidae order-Trichoptera class-Insecta phylum-Arthropoda kingdom-Animalia Eukaryota Hexapoda Hydropsychoidea Polycentropodinae iNaturalist GBIF Catalogue-of-Life NCBI-Taxonomy Wikipedia Zeitschrift-für-Tierpsychologie Freshwater-Biology Zootaxa Genome-Biology-and-Evolution Zoosymposia DOI abstract paper-summary evidence confidence-notes observations-count matched-scientific-name canonical-name rank status accepted match-type higherrank distribution-records genus-of-tube-maker-caddisflies more-than-120-described-species list-of-species references further-reading external-links title journal subjects Zusammenfassung Die-Larven-von leben-in-Fließwässern fängt-mit-einem-Netz-Beute hauptsächlich-Chironomiden-Larven-und-Oligochaeten Wirkung-der-von-der-Beute-im-Netz-erzeugten-Vibrationen Aufmerksamkeit Orientierung-und-Bewegung Fangversuche um-so-schneller verwirrt Baubewegungen Bauverhalten Beutefang enger-Verbindung recruitment kin southern-English-stream objectives small-scale-patterns stream-dwelling spatial-proximity-of-close-kin patchy-recruitment distribution-of-related-larvae aquatic-phase egg-masses spatially-and-temporally-structured-samples field-collected-larvae six-polymorphic-microsatellite-loci siblings progeny-of-one-father background-population-level siblings-disperse changes-in-spatial-genetic-structure neighbouring-larvae avoiding-kin onset-of-pupation survival-through-the-egg-stage first-instar-larvae number-of-egg-masses refute larva-of including larvae-of species-of-Greece morphology final-instar-larva inner-and-outer-dorsal-secondary-setae abdominal-segment-IX muscle-attachment-spots head-capsule abdominal-sternum-IX distribution-patterns annotated-draft-genomes larval-silk-secretions diverse-case-making-behavior ecological-niches five-genomes low-cost-sequencing-strategy Oxford-Nanopore-flow-cell Illumina-sequence-reads high-quality-genomes de-novo-assembly-methods low-coverage-Nanopore-reads shortest-genomes light-L-chain-fibroin L-fibroin-gene-clusters phylogenomic comparative-genomic larvae-of-the-genus other-two-British life-cycle adult genetic-population-structure neighbourhood-population-size-estimates role-of-history scale-of-colonisation structuring-populations genetic-and-ecological-methods no-genetic-differentiation up-to-20-km despite-population-sizes greater-than-expected contrasting-short-range-trend implausibly-small relatively-short-flights winged-adults found-small often-temporary larger-and-more-permanent amplifying regions-containing reduce date-from rarely-examined central biology-of springs high-oxygen-concentration good-indicator water-quality tube-maker-caddisflies genus observations taxonomy-match matched canonical classification Animalia Arthropoda Insecta group caddisflies Metazoa genus-Plectrocnemia list-of-Plectrocnemia-species vibrations-and-predatory-behaviour effects-of-vibrations-transmitted-across-the-net predatory-behaviour variations-in-the-frequency more-marked-effect variations-in-amplitude stage-2 orientation-and-displacement-towards-the-lure stage-3 capture-of-the-lure-with-mandibles larvae-live-in-running-waters catch-with-a-net prey mainly-chironomid-larvae-and-oligochaetes effect-of-vibrations-generated-by-prey-in-the-net very-irregularly-woven-net open-ended-dwelling-tube-at-both-ends vibration-weakly-dampened frequency-does-not-change vibration-excites attention orientation-and-movement capture-attempts orientation-and-movement-to-prey the-faster the-more-the-vibration-exceeds-0.28-Hz frequencies-of-0.15-to-0.28-Hz lead-to-incomplete-reactions as-if-the-larvae-were-confused frequencies-below-0.075-Hz generate-building-movements building-behavior-instead-of-prey-capture closely-connected recruitment-kin-and-spatial-genetic-structure oviposition-and-genetic-relatedness stream-dwelling-caddis beginning-of-the-aquatic-phase four-sample-dates within-one-generation mean-relatedness-coefficient within-reared-egg-masses differed-significantly population-as-a-whole markers-sufficiently-powerful identify-groups-of-siblings small-contribution-from-a-second-male mean-relatedness-within-spatially-structured-groups did-not-differ-from-background siblings-disperse-away-from-each-other kin-structure-does-not-persist changes-in-spatial-genetic-structure-late-in-larval-life neighbouring-larvae-less-closely-related approaching-onset-of-pupation survival-through-egg-stage-and-early-larval-life very-high greater-than-50%non-social-insect consequence-of-colonial-net briefly-occupied-by-first-instar-larvae lack-of-spatial-genetic-structure high-survival refute-patchy-recruitment-hypothesis larva-of-Plectrocnemia-renetta including-discriminatory-matrix larvae-of-Plectrocnemia-Stephens-1836-species-of-Greece morphology-of-final-instar-larva most-important-diagnostic-features-illustrated preliminary-discriminatory-matrix strongly-different-in-length separated-from-each-other muscle-attachment-spots-on-head-capsule number-and-length-of-setae-on-abdominal-sternum-IX reported-from-Cyprus-Turkey-Greek-islands exploit-wide-range-of-ecological-niches five-genomes-published variable-qualities single-Oxford-Nanopore-flow-cell de-novo-assembly-methods-compared assembly-of-low-coverage-Nanopore-reads subsequent-polishing yielded-highest-genome-quality contiguity-and-BUSCO-completeness shortest-genomes-to-date extend-knowledge-of-genome-size genomic-region-encodes-for-light-L-chain-fibroin protein-component-of-larval-caddisfly-silk identified-and-compared new-genomic-resources among-highest-quality-Trichoptera-genomes increase-knowledge basis-for-phylogenomic-and-comparative-genomic-studies revised-key-to-larvae distinguishes-previously-unknown-larva other-two-British-species notes-on-larval-habitat-life-cycle-and-identification-of-adult genetic-population-structure-and-neighbourhood-population-size-estimates role-of-history-and-scale-of-colonisation no-genetic-differentiation-between-sites-up-to-20-km despite-population-sizes-suggesting-genetic-drift genetic-differentiation-between-populations-separated-by-more-than-20-km neighbourhood-population-size-implausibly-small evolutionary-processes-do-not-explain-differentiation relatively-short-flights-by-winged-adults spread-over-number-of-generations found-small-often-temporary-populations grow-and-exchange-genes-with-larger-permanent-local-populations amplify-effects-of-initial-gene-flow substantial-gaps-between-regions-containing-suitable-habitat-patches reduce-number-of-colonisation-events genetic-patterns-may-date-from-time-last-colonised ecological-studies-rarely-examined-dynamics-over-larger-geographical-scales central-to-persistence-and-spread biology-of-Plectrocnemia-latissima rivers-and-streams-of-Central-Caucasus-northern-slopes springs-streams-and-rivers requires-high-oxygen-concentration good-indicator-of-water-quality aspects-of-biology-reported Wikipedia-summary rank-GENUS status-ACCEPTED match-type-HIGHERRANK distribution-records-DK-NO-SE-Vermont-US-US scientific-name authorship-Stephens-1836 classification-Eukaryota-Animalia-Arthropoda-Hexapoda-Insecta-Trichoptera-Hydropsychoidea-Polycentropodidae-Polycentropodinae-Plectrocnemia scientific-name-Plectrocnemia group-caddisflies kingdom-Metazoa instructions fill-all-fields if-a-field-cannot-be-supported-return-null do-not-repeat-information-across-fields keep-each-section-focused-on-its-purpose provide-useful-detail-where-possible critical-rules factual-correctness-over-completeness clarity-over-verbosity usefulness-over-speculation if-information-is-not-clearly-supported-return-null do-not-infer-species-level-traits-from-higher-taxa-unless-explicitly-justified do-not-repeat-the-same-information-across-multiple-fields each-field-must-contain-unique-non-overlapping-content avoid-vague-generalizations like-most-insects typically-feeds-on-plants use-cautious-language-when-necessary has-been-observed is-known-to do-not-fabricate behaviors diet life-cycle-details host-relationships field-intent summary-high-level-overview-3-5-sentences appearance-physical-description-only identification-how-to-distinguish-it-from-similar-taxa habitat-environment-and-conditions distribution-geographic-range-only seasonality-timing-of-activity diet-feeding-habits-null-if-unknown lifeCycle-developmental-stages behavior-notable-actions-or-habits ecologicalRole-role-in-ecosystem humanRelevance-interaction-with-humans similarTaxa-must-include-reason misconceptions-only-if-meaningful extraDetails-only-for-important-additional-context style-rules use-clear-direct-sentences avoid-fluff-or-filler-language avoid-repeating-taxonomy-in-prose avoid-overly-technical-jargon-unless-necessary prefer-concrete-statements-over-abstract-descriptions quality-rules completeness-high-only-if-most-fields-are-well-supported completeness-medium-if-partial-but-reliable completeness-low-if-sparse-data hasInferredContent-true-only-if-generalization-is-used otherwise-false output-format must-strictly-match-provided-JSON-schema do-not-include-any-extra-fields do-not-include-commentary-outside-JSON taxon-record using-provided-schema optional-context-may-be-incomplete source paper-summary-evidence limited-information-extracted-from-abstract-only full-text-not-available-for-more-detailed-extraction habitat-diet-life-cycle-reproduction-behaviors-and-ecosystem-role-not-mentioned-in-abstract distribution-data-limited-to-abstract-level-information full-paper-may-contain-additional-details abstract-only-provided full-text-not-available habitat-diet-and-ecological-details-likely-contained-in-main-paper-but-not-accessible-from-abstract-alone distribution-limited-to-Britain-as-explicitly-stated generate structured-taxon-records entomology-guide accurate-conservative-informative-content prioritize goal produce json schema content summary appearance identification habitat distribution seasonality lifeCycle hostAssociations behavior ecologicalRole humanRelevance similarTaxa misconceptions extraDetails tags completeness hasInferredContent metadata sources sourceQuality extractionMethod extractionDate confidence notes reason name high medium low true false Vibrations-and-Predatory-Behaviour-of-Plectrocnemia-Larvae vibrations net frequency amplitude orientation capture mandibles running-waters dwelling-tube dampened building-movements confused Recruitment-kin-and-the-spatial-genetic-structure-of-a-caddisfly-Plectrocnemia-conspersa microsatellite-loci survival The-larva-of-Plectrocnemia-renetta-Malicky-1975 larva setae abdominal-segment muscle-attachment Ikaria Samos Annotated-Draft-Genomes-of-Two-Caddisfly-Species fibroin Oxford-Nanopore BUSCO A-revised-key-to-larvae-of-the-genus-Plectrocnemia Plectrocnemia-geniculata colonisation dispersal-flights streams rivers 120-described-species Denmark Norway Sweden United-States 204 DK NO SE US taxonomy match kingdom phylum class order family specific-epithet subspecies-epithet subphylum subclass suborder infraorder tribe scientific-name-authorship synonyms common-names content-fields all-fields available-knowledge return-null not-supported unique-non-overlapping focused useful-detail factual correct clear direct no-fluff no-filler no-taxonomy-repetition no-jargon concrete high-completeness medium-completeness low-completeness no-inferred-content strict-JSON no-extra-fields no-commentary entomology insects aquatic vibration genetics population egg pupa instar oxygen Asia America England flowing-water Chironomid oligochaete colonial relatedness sibling Illumina-sequencing annotated silk-protein case-making identification-key morphological diagnostic setal-arrangement muscle-spot abdominal-sternum larval-stage oviposition-site hot-spot survival-rate habitat-patch temporary-population permanent-population winged-adult kin-avoidance refuted water-quality-indicator high-oxygen genus-authority type-species not-specified see-also abstract-only limited-information extraction habitat-not-mentioned diet-not-mentioned life-cycle-not-mentioned reproduction-not-mentioned behaviors-not-mentioned ecosystem-role-not-mentioned distribution-limited abstract-level main-paper additional-details likely-contained not-accessible Britain-explicitly-stated larval-stages-described detailed-extraction predatory captures-prey silken-nets substrate-borne-vibrations chironomid-larvae oligochaetes frequency-effects amplitude-effects orientation-stage capture-stage net-construction confused-responses incomplete-reactions short-range long-range 20-km 500-km colonization-events persistence spread organic-matter material-properties quality genome-size-variation silk-encoding-genes protein-components gene-clusters basis-for-studies distinguishes-species British-species morphological-characteristics taxonomic-revision population-structure neighborhood-size history scale structuring methods differentiation drift trend processes flights generations found grow exchange amplify gaps suitable date examined requires indicator aspects northern slopes Martynov Greek-islands separated-by arrangement number length group-where strongly-different diagnostic-features-illustrated preliminary matrix-provided morphology-of information-on given most-important discriminatory to-the-larvae species-of previously-unknown describes this-paper effects-of-vibrations transmitted-across analyses work this analyses-the-effects variations-in has-a-more-marked-effect especially-on live-in catch-with effect-of generated-by investigated very-irregularly-woven open-at-both-ends weakly-dampened does-not-change excites the-more exceeds lead-to as-if otherwise sometimes-occurs suggests our-objectives examine in-particular look-for any-evidence therefore in-order-to at-the-beginning over-four subsequently-compared reared-from ranged-from indicating-that sufficiently-powerful likely-to-be although could-not-be-excluded did-not-differ suggesting-that very-quickly does-not-persist indicated-that possibly-suggesting some-direct-or-indirect-means when-approaching our-counts suggested-that apparently-very-high may-be-a-consequence all-refute also-provided belong-to can-be-separated with-respect-to has-been-reported members-of provide-important for-example due-to these-form-the-basis only-five published-thus-far with-variable-qualities regarding was-successfully-used of-the compared yielded both-in-terms to-date extend-our-knowledge across was-identified-and-compared with-existing presented-in-this-paper are-among will-increase by-serving-as from-larvae-of are-given-on of-the-adult used-both to-evaluate there-was-no despite given-the implied-that is-implausibly-small do-not-explain at-small-scales could-account-for for-instance may-then over-larger-scales could-reduce may-date-from have-rarely-examined yet-these-processes may-be-central-to from-the-rivers-and-streams aspects-of are-reported-here can-be-used-as high-level-overview 3-5-sentences physical-description-only how-to-distinguish environment-and-conditions geographic-range-only timing-of-activity feeding-habits developmental-stages notable-actions-or-habits role-in-ecosystem interaction-with-humans must-include-reason only-if-meaningful only-for-important-additional-context clear-direct-sentences no-fluff-or-filler no-repeating-taxonomy no-overly-technical-jargon prefer-concrete high-only-if-most-fields-well-supported medium-if-partial-but-reliable low-if-sparse-data true-only-if-generalization-used strictly-match no-commentary-outside-JSON generate-taxon-record taxon Plectrocnemia optional-context may-be-incomplete if-a-field-cannot-be-supported keep-each-section-focused provide-useful-detail factual-correctness clarity verbosity usefulness speculation information-not-clearly-supported do-not-infer-species-level-traits from-higher-taxa unless-explicitly-justified do-not-repeat same-information multiple-fields each-field-must-contain unique-non-overlapping-content use-cautious-language accurate conservative informative structured records scientificName canonicalName scientificNameAuthorship taxonRank commonNames subfamily speciesEpithet subspeciesEpithet
Pogonomyrmex comanche
Comanche harvester ant
Pogonomyrmex comanche, the Comanche harvester ant, is an endangered harvester ant species endemic to the western Gulf Coastal Plain of North America. Genetic studies indicate central Texas serves as the center of genetic diversity, consistent with Pleistocene refugia patterns observed in many regional species. Northern populations in Arkansas and Oklahoma exhibit lower genetic diversity and evidence of population contraction rather than recent range expansion. The species shows signs of dispersal limitation, with high genetic diversity and few identical sequences among sampled individuals. Its decline appears driven by a combination of habitat loss and limited dispersal capacity, though exact mechanisms remain under investigation.
endangered harvester-ant Pogonomyrmex western-Gulf-Coastal-Plain phylogeography Pleistocene-refugia dispersal-limitation conservation Formicidae North-America Texas Oklahoma Arkansas sandy-soils prairie savanna
Polistes dominula
European Paper Wasp
Polistes dominula is a highly successful invasive social wasp native to Eurasia that has established populations across North America, South America, New Zealand, South Africa, and other regions. First detected in North America near Boston in 1978, it has become one of the most abundant wasps on the continent. The species builds small, exposed paper nests in protected locations and preys primarily on live insects, particularly caterpillars. Unlike yellowjackets, it does not scavenge for meat or sugar. Its rapid spread has been attributed to ecological flexibility, superior competitive ability, and tolerance of human-altered environments.
invasive-species social-wasp paper-wasp predator pollinator urban-ecology eusocial nest-construction biological-control thermal-biology spatial-learning vibrational-communication aposematism honest-signal overwintering diapause Polistes Vespidae Hymenoptera North-America Europe New-Zealand South-Africa South-America monarch-butterfly caterpillar-predation nest-architecture gyne foundress dominance-hierarchy climate-change-impact energetics metabolic-rate microclimate suburban-habitat anthropogenic ornamental-pest vineyard-pest fruit-damage stinging-insect venom poison-gland warning-coloration Zahavian-signal Strepsiptera parasitoid Xenos behavioral-plasticity foraging learning memory thermoregulation ectothermy endothermy solar-radiation nest-temperature brood-development colony-survival multiple-founding single-founding season-length latitude temperate Mediterranean hibernacle overwintering-costs climate-warming range-expansion competition native-species-displacement biodiversity-impact conservation management control integrated-pest-management IPM identification antennae-color facial-markings caste reproduction male female sexual-dimorphism antennae abdomen wing mandible saliva paper comb cell pedicel trophallaxis larva pupa adult worker queen subordinate dominant aggression defense threat-display sting toxicity brightness coloration predator-avoidance prey caterpillar Lepidoptera butterfly moth garden agriculture vineyard orchard urban suburban rural forest habitat-preference temperature metabolism quiescence torpor survival fitness colony nest population abundance distribution range expansion invasion establishment introduction biotic-interaction trophic-cascade herbivory-suppression plant-fitness milkweed Danaus-plexippus monarch Nelson Tasman ecosystem-impact conservation-management biological-invasion ecology behavior physiology morphology taxonomy systematics nomenclature gender-agreement Latin Christ 1791 Vespa-dominula basionym catalogue-of-life GBIF iNaturalist observation citizen-science photography illustration field-guide orange black yellow red color-pattern size wing-length body compact legs short face square triangular dark curled straight elbowed blunt pointed tip Zahavian costly-signal hunt forage nectar honeydew aphid scale-insect flower grape umbellifer fennel lovage insect meatball feed emerge chew cap hexagonal wood-fiber bark exposed open umbrella stalk ant repellent substance coat protect location eave attic tree branch vine shrub cavity bird-box bee-hive shed grave-lantern overwinter shelter protected thermal climate cold winter spring summer autumn fall season month day activity defend threat display raise tiptoe social dominance hierarchy reproductive egg lay found cooperate compete selfish passive-aggressive individual recognition facial-marking cognition mushroom-body brain intelligent artist architect navigation plasticity spatial relocating food resource dish bait training experience vibration communication signal substrate-borne abdominal wagging oscillation secretion modulation perceive react movement attention feeding inspection body-temperature ambient solar radiation metabolic cost energetic expenditure efficiency warming change impact increase invasive spread success competitive ability flexibility adaptation reaction response condition year variation productivity founding single multiple rate benefit cooperation predation parasitism vesparum stylopid stylopization parasite host infestation mite Acari wasp fly effect community decline Danaus plexippus mortality instar early late larger smaller vulnerable resistant discovery chance contact direct recruitment nestmate site cascading trophic cascade herbivory suppression plant growth production Asclepias net exclusion experiment outcome suggestion strong detrimental ecosystem strategy method development protection biodiversity region first 2016 recent established exotic congener chinensis longer comparison higher lower greatest area built structure native unsuccessful translocate unable thrive warmer origin contributor factor research thesis insight behaviour knowledge direction aid effective journal publication DOI 10.26686 wgtn 14538585 v1 University Wellington Victoria MSc student dissertation study investigation analysis data field resident report local indirect interaction level function service provision natural pest biocontrol ornamental decorative crop damage fruit cherry western Colorado at least ripening choice preference height ground strata microsite environment closed habitat dense vegetation surrounding bush Patagonia Argentina Chile NW detected 2003 sympatric coexistence overlap minimal facilitation behavioral process interspecific semi-urban poorly studied species Hypodynerus labiatus new light finding contribution understanding collecting anthropized highlight importance consideration Ethology 10.1111 eth 13505 abstract only limited detail full text required comprehensive synthesis vibrational widespread regulate crucial conspicuous oscillatory wag strictly associated presence suggested involvement adult-larva hypothesis short-term trophallactic exchange modulate salivary decrease amount prepare receive stimulate release electro-magnetic shaker assess time record measure immediately after playback result show produce possibly order attract during neither nor support alleged role long-lasting Experimental Biology 10.1242 jeb 186247 flexible system expanding North America cause stir fast upon explanation present literature good invader northern rise little known nesting Central investigate basic principle Germany 49°group one to four small shorter than other mean newly founded 83 average length 4.6 prefer 54%46%major increased 47%single-foundress whereas 100%more two however no number term per observe well three consecutive greatly depending preceding imply remarkably quickly outside rapid colder Ecosphere 10.1890 es15-00254 1 thermoregulatory polistine different collect demand brood mainly ectothermic task living differing reveal environmental influence warm Southern infrared thermography entire usually ~20–40 °C nearly linearly thorax warmest part achieve optimal preferably use model enable evaluation endothermic effort exhibit weak almost constant performance about contrast mostly minor avoid high way reduce Insects 10.3390 insects10070187 trait strongly Comparative B 10.1007 s00360-016-1041-x provided gallicus insects14110849 adverse weather offer low sheltered live energy store Italy Austria describe calculate according standard differ significantly between 3.2 ±5.71 8.5 5.29 variance correspond closely meteorological cumulative mass-specific over period lowest compared both acclimation calculation up 3 due dramatic 40%additional exponentially temperature-dependent fluctuation disproportionately Jensen's inequality wooden bird box empty bee hive under grapevine grave lantern cemetery begin when exceed ~18–20 confidence note information inferred from active not discussed this source association addressed relationship insects14110886 thermophilic originating but now widely quite synanthropic human settlement place with favorable typical side wasps'wide ~20–37 above air advantage as accelerate speed did approximately 38.6 managed by cooling near ~1–2 cm always clearly elevated station climate-model-generated macroclimate revealed necessity measuring for reliable description insects'brighter-colored poison gland Frontiers Zoology 10.1186 1742-9994-9-20 aposmatism against consisting warning using toxin costly physical could simultaneously abundant striking such case indicate animal toxic also would ensure honestly examine positive correlation exists collected 30 photographed them volume width remained even controlling suggest pattern true sign poisonous they have theoretical predicting indicates quantitative within acting metabolically therefore limitation first-year Spain exact age unknown ethanol preservation may alter applied uniformly pseudoreplication intraspecific supported while successfully worldwide revisiting previously gathered flight manipulation performed target several visit additionally analyse dealing position whether choose visited containing opt novel baited placed 60 away then train either once or compare taken find displaced demonstrate rapidly learned return certain significant reduction likewise manipulate spending less each moreover offered just majority furthermore contribute behavioural considering marginalis Cape Province South Africa African Entomology 10.4001 003 022 0104 Caucasian Entomological Bulletin 10.23885 1814-3326-2015-11-1-85-89 degree Hungary Natura Somogyiensis 10.24394 natsom 2011 19 229 many European country observed 1993 documentation photograph 5 of examined stylopized these exemplar parasitized 18 strepsipteran specimen some taxon scientific name rank preferred common Wonderful World Wasps Bug Squad Article Chris Alice Kratzer author LinkedIn Photo terrible jam-packed pain anger love us out blue minding own business eat picnic frighten child build home complicate already-stressful life Everyone knows be better without So preface book published company 2020 Owlfly LLC what write believe point essential complicated beautiful Indeed play important provide beneficial Look ma pesticide labor passionate dedicated written conversational tone long admirer supporter who grew Jersey continue remember interacting yes getting stung assorted including come territory become entomologist opted engineer received bachelor science 2019 mechanical engineering Rochester Institute Technology worked York until Today executive director managing division Publishing opportunity develop market sustainable potential mitigation fascinate fold into want why how do if tell building understand cycle organism before trying entangle ecological meaning inquiline called cuckoo exploit young call fact catch dismember tiny waist prevent eating sold fluid primary most grasshopper cricket katydid beetle earthworm lanternfly cockroach spider cicada aren't engineering-trained professional knowledgeably began her 2018 January 25 path included holing Cornell Collection networking expert teaching herself software align genetic barcode spent close thousand hour poring platform identifying mapping studying color incredible 400-page billed guide all arctic Greenland Alaska tropical Panama Grenada cover 208 22 genera feature 900 image macro photographer posting bring back Iowa State Professor Amy Toth's seminar Department Nematology told crowd attack extremely complex fascinating creature coined hashtag wasplove wrote illustrated It variety yellowjacket Eurasian red-banded American blackjacket volcano Guatemalian Eastern widow polka-dot prairie California form downy German knew there 'jackets familiar elaborate know dominulo means mistress implies Introduced 1970s since continent Woman Lynn Kimsey Bohart Museum distinguished telling difference scavenger take particularly serious mention UC Statewide Integrated Program wealth PestNote distinct type far troublesome especially ground-cavity-nesting tend vigorously disturbed Defensive increases progresses scarcer primarily start around garbage can dog cat where ripe overripe accessible large colorful tidbit sprinkle throughout intriguing instance Pennsylvania despite Someone mislabeled original And rated 2/5 painful Schmidt Index Bottom line eye-opener page-turner mind-boggler offers approach belongs library scientist citizen curious person wants incredibly diverse amazing next gathering anti-social folk ask anyone sharing rose Davis Calif thirsty sipping water fountain Agraulis vanillae Vacaville tucked inside Wednesday Cincinnati Ohio among changed recently overtaken counterpart Still United adjacent belong same family horn potter mason boreal cooler decidedly handful reaching rarely exceeding 200 unprotected suspend twig stem semi-protected hang sturdy often coated repel chief construction apparent destined likely determined stage act eventually assert leave performs securing developing offspring harvesting fiber surface dead fence post scrape ball deftly plies strip added existing worth having heavily Webworm grab hairy butcher kill spot stripping off any irritating hair spine easily consumed directly share turn distribute occupy reach maturity spin silken dome top molt shortly thereafter freedom sole fertilize Watch August September easy though genders elbow taper goldenrod Solidago throughwort Eupatorium evident extreme variability pair deep marking vary considerably DNA adding mud situation instead clarifying shown below going lump Enjoy watching unless antagonize give standing tip-toe raising make perfectly peaceful neighbor Posted 2:40 AM Email ThisBlogThis!Share XShare FacebookShare Pinterest Labels nature 6 comment ShandaSeptember 8 2010 5:26 loved get never faced can't wait kid question left end hibernate produced ReplyDelete RepliesReplyOut prairieSeptember 8:02 keeping cabbage worm seen RepliesReplyAnonymousSeptember 13 2:06 PM becoming here Upper Peninsula Michigan pendulum seems swung ago everywhere vanishingly rare haven't again house right RepliesReplyL.R ShimerSeptember 4:35 head gal work everybody else kinda egalitarian RepliesReplyLarryMay 9:34 directed Bev Wigney BTW lived NE Missouri detailed presented Thanks cogent scientifically accurate summary insect's life-cycle RepliesBug EricMay 3:47 kind compliments Delete RepliesReplyReplyAdd commentLoad Blog currently reply reader himself Working resolve Newer Older Subscribe Atom Demonized hosting 4 p.m.Saturday Jan 20 Room 1124 Academic Surge 455 Crocker Lane campus Attendees chat check 50,000 vespid eight-million free friendly Parking estimate 2000 described inhabit 500 undescribed headline-grabbing giant Vespa mandarinia formerly Asian news media dubbed murder hornet confuse identify sheet example hidden rodent burrow wall huge 100,00 will scavenge meat sugar soda hamburger road open-faced commonly located annual die cabbageworm white superb much delivered presentation website interested mechanism evolution honey Current project involve novo sequencing genome transcriptome genomic epigenetic regulating nutrition virus health hold doctorate Illinois Urbana–Champaign Gene Robinson postdoctoral Christina Grozinger unwelcome yard patio Preferring like beneath Sometimes legless grublike outstretched hand 15 relatively unaggressive problem doorway list earlier lepidopteran carry yeast winemaking fermentation flavor wine fuscatus recognize devoted mother dote fury opinion human-like great cooperator overall undercurrent selfishness manifest perfect themselves mushroom learning/memory 10 photogenic peer petal visitor camp Sierra Nevada mountain witnessed stinging crowded table ah sipped wondered says elsewhere which fertilized must warms enough lot protein baby need survived because drier usual deal competing go maintenance mode maintain fuel largely useless started mid trap spraying removed--this done night hollow available nearby hope wet barbecue shy except door window high-traffic Typically mated midsummer phase require cease sweet thing normally mild coastal San Francisco survive informative YouTube video Distinguishing see those distinctive put another Oakland Raiders'logo Giants soaking sun abandoned Pollinators Artists delight overhang neighborhood thought Wow hanging lip trash vanished Twitter associate Organismal
Polites
grass skippers
Polites is a genus of North American grass skipper butterflies in the family Hesperiidae. The genus was established by Scudder in 1872 and contains species primarily distributed across the United States. Members of this genus are small to medium-sized skippers with rapid, darting flight patterns typical of the subfamily Hesperiinae. Some species have become subjects of conservation concern due to population declines, with at least one species (Polites mardon) having been petitioned for federal endangered species protection.
Hesperiidae grass-skippers North-America grassland-butterflies conservation
Polyommatus
blue butterflies
Polyommatus is a genus of blue butterflies in the family Lycaenidae, distributed across the Palearctic realm. Recent molecular studies have redefined its boundaries, with former subgenera Cyaniris, Lysandra, and Neolysandra now recognized as separate genera. Some authors continue to recognize additional subgenera including Agrodiaetus, Bryna, Meleageria, and Plebicula. The genus contains species of conservation concern, including critically endangered and vulnerable taxa with specific habitat requirements.
Lycaenidae blue-butterflies Palearctic molecular-systematics cryptic-species conservation grassland-indicator pollinator
Pomachiliini
Pomachiliini is a tribe of click beetles (Elateridae) established by Candèze in 1859. The tribe includes the genus Isidus, whose species are specialized to coastal dune environments in the Mediterranean region. Isidus moreli, the best-studied member, is the only saproxylophagous elaterid known from Italian dunes and serves as an umbrella species for coastal conservation.
Elateridae click-beetles coastal-dunes Mediterranean saproxylophagous conservation umbrella-species bioindicator
Pseudanophthalmus
Pseudanophthalmus is a genus of small carabid beetles comprising over 200 described species, all obligate cave dwellers restricted to karst regions of the eastern United States. Species exhibit extreme short-range endemism, often confined to single caves or cave systems. The genus forms part of the Trechoblemus complex within the tribe Trechini. Many species are rare or potentially extinct due to habitat restriction and anthropogenic threats.
cave-beetle troglobite karst endemic conservation Trechini
Pugettia
kelp crabs
Pugettia is a genus of marine kelp crabs in the family Epialtidae, distributed across the North Pacific from North America to East Asia. Species inhabit shallow subtidal zones, primarily associated with macroalgal habitats including kelp beds, Sargassum stands, and red algal turfs. Many species exhibit ontogenetic habitat shifts, with juveniles and smaller individuals occupying deeper algal turfs while larger adults migrate to shallower macroalgal beds. The genus includes approximately 25 extant species plus one fossil species, with several species serving as important subjects for studies of crab growth, reproduction, and habitat ecology.
kelp-crab spider-crab macroalgae ontogenetic-shift subtidal North-Pacific Epialtidae decapod Brachyura marine-invertebrate kelp-forest algal-turf seasonal-migration terminal-molt functional-maturity decorating-behavior piscivory abalone-predation sea-urchin-predation one-year-life-cycle sexual-dimorphism Sargassum Gelidiaceae Grateoloupia East-Asia North-America Sagami-Bay juvenile-recruitment ovigerous-females carapace-morphology gonopod-morphology epialtine Majoidea heterotreme eubrachyuran pleocyemate malacostracan crustacean arthropod invertebrate benthic phytal littoral sublittoral intertidal rocky-shore temperate-marine Pacific-Ocean Japan Philippines Taiwan Russia Tasmania British-Columbia California Baja-California Dana-1851 Rathbun De-Haan Yokoya Stimpson Sakai Garth Ohtsuchi Kawamura Takeda Komatsu Li Miyake Ortmann Richer-de-Forges Karasawa Kitamura Randall Zootaxa Crustaceana Journal-of-Crustacean-Biology Fisheries-Science PeerJ ontogenetic-stages post-settlement recruitment population-dynamics growth-patterns morphometrics cheliped pleon abdomen carapace-width CW PCL CL CH AW relative-claw-length relative-abdominal-width size-at-maturity 50%-maturity anecdysis ecdysis molt copulation breeding clutch larvae hatching predator-prey trophic-ecology food-web herbivory omnivory carnivory opportunistic-feeding covering-behavior habitat-segregation habitat-preference habitat-complexity settlement-selectivity mortality density abundance seasonal-occurrence spring summer autumn winter annual-life-cycle short-lived iteroparity semelparity sexual-maturity behavioral-maturity morphological-maturity pre-terminal-molt post-terminal-molt virgin-female laboratory-observation field-observation inferred conservative factual documented observed known reported described illustrated redescribed taxonomic-revision species-delimitation new-species type-locality distribution-range range-overlap sympatric allopatric parapatric endemic cosmopolitan temperate boreal neritic epibenthic demersal sessile motile mobile cryptic conspicuous camouflage mimicry decorating masking algal-association phytal-habitat canopy understory turf bed stand forest reef rock sand mud sediment substrate complex-structure physical-structure biogenic-habitat ecosystem-engineer foundation-species keystone-species indicator-species fisheries aquaculture stock-assessment management conservation biodiversity biogeography phylogeny systematics taxonomy nomenclature homonym synonym junior-synonym senior-synonym valid-name accepted-name nominal-species species-inquirenda species-complex sibling-species cryptic-species morphospecies biological-species evolutionary-species phylogenetic-species operational-taxonomic-unit OTU DNA-barcoding molecular-phylogenetics integrative-taxonomy alpha-taxonomy beta-taxonomy gamma-taxonomy monograph revision redescription diagnosis description illustration measurement meristic morphometric allometry ontogeny heterochrony paedomorphosis peramorphosis development embryology larval-development zoea megalopa settlement juvenile immature mature adult subadult gerontic senescence longevity life-span life-history life-history-strategy r-selection K-selection bet-hedging reproductive-effort reproductive-success fitness population demography cohort generation age-structure size-structure stage-structure population-regulation density-dependence density-independence carrying-capacity limiting-factor environmental-gradient habitat-gradient depth-gradient zonation vertical-distribution horizontal-distribution geographic-distribution latitudinal-gradient longitudinal-gradient bathymetric-distribution intertidal-zone subtidal-zone continental-shelf continental-slope abyssal-zone hadal-zone neritic-zone oceanic-zone pelagic-zone benthic-zone demersal-zone epipelagic mesopelagic bathypelagic abyssopelagic hadopelagic supralittoral circalittoral archibenthal bathyal abyssal hadal euphotic dysphotic aphotic photic-zone disphotic-zone aphotic-zone trophic-level primary-producer primary-consumer secondary-consumer tertiary-consumer detritivore scavenger predator herbivore carnivore omnivore parasite commensal mutualist symbiont competitor food-chain trophic-cascade bottom-up-control top-down-control wassefug mesopredator-release trophic-trickle nutrient-cycling energy-flow productivity biomass standing-stock turnover production respiration excretion egestion assimilation efficiency ecological-efficiency trophic-transfer-efficiency Lindeman-efficiency 10%-rule pyramid-of-numbers pyramid-of-biomass pyramid-of-energy ecosystem-function ecosystem-service ecosystem-process biogeochemical-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle oxygen-cycle hydrogen-cycle mineral-cycle nutrient limiting-nutrient eutrophication oligotrophication hypoxia anoxia acidification warming climate-change global-change anthropogenic-impact pollution contamination habitat-loss habitat-degradation habitat-fragmentation overfishing bycatch invasive-species introduced-species non-native-species alien-species exotic-species range-expansion range-contraction range-shift poleward-shift tropicalization borealization meridionalization deglaciation sea-level-rise coastal-squeeze ocean-acidification ocean-warming deoxygenation marine-heatwave extreme-event disturbance resilience resistance recovery adaptation acclimation plasticity evolutionary-adaptation genetic-adaptation phenotypic-plasticity behavioral-plasticity physiological-plasticity morphological-plasticity life-history-plasticity trade-off constraint limitation bottleneck bottleneck-effect founder-effect genetic-drift gene-flow mutation selection natural-selection sexual-selection artificial-selection directional-selection stabilizing-selection disruptive-selection frequency-dependent-selection density-dependent-selection rare-advantage common-advantage apostatic-selection balancing-selection heterozygote-advantage negative-frequency-dependent-selection positive-frequency-dependent-selection ecological-speciation sympatric-speciation allopatric-speciation parapatric-speciation peripatric-speciation vicariance dispersal colonization establishment invasion naturalization spreading epidemic pandemic enzootic epizootic eradication control protection restoration rehabilitation rewilding de-extinction sustainable-use ecosystem-based-management adaptive-management precautionary-approach ecosystem-approach integrated-management marine-protected-area MPA no-take-zone marine-reserve sanctuary world-heritage-site Ramsar-site important-marine-mammal-area IMMA ecologically-or-biologically-significant-area EBSA vulnerable-marine-ecosystem VME essential-fish-habitat EFH critical-habitat habitat-of-particular-concern habitat-area-of-particular-concern HAPC key-biodiversity-area KBA protected-area conservation-area nature-reserve national-park marine-park aquatic-reserve fishery-reserve species-recovery-plan action-plan management-plan strategic-plan policy legislation regulation convention treaty agreement protocol Memorandum-of-Understanding MoU non-binding-instrument soft-law hard-law customary-international-law jus-cogens erga-omnes common-concern-of-humankind common-heritage-of-mankind precautionary-principle polluter-pays-principle common-but-differentiated-responsibilities intergenerational-equity intragenerational-equity environmental-justice climate-justice ocean-justice blue-economy green-economy circular-economy bioeconomy natural-capital ecosystem-services-valuation payment-for-ecosystem-services PES blue-carbon carbon-sequestration carbon-storage carbon-sink carbon-source greenhouse-gas GHG carbon-dioxide CO2 methane CH4 nitrous-oxide N2O fluorinated-gas F-gas hydrofluorocarbon HFC perfluorocarbon PFC sulfur-hexafluoride SF6 nitrogen-trifluoride NF3 ozone-depleting-substance ODS Montreal-Protocol Paris-Agreement UNFCCC IPCC IPBES IOC UNESCO FAO IMO ISA CBD CMS CITES RAMSAR World-Heritage-Convention UNFSA Straddling-Stocks-Agreement Compliance-Agreement Port-State-Measures-Agreement PSMA IUU-fishing illegal-unreported-unregulated-fishing overcapacity fishing-down-the-food-web fishing-through-the-food-web serial-depletion shifting-baseline baseline reference-point target-reference-point limit-reference-point threshold tipping-point point-of-no-return hysteresis alternative-stable-state regime-shift critical-transition catastrophic-shift smooth-transition adaptive-cycle panarchy resilience-thinking social-ecological-system SES coupled-human-natural-system CHANS human-environment-system HES socio-ecological-system ecosystem-approach-to-fisheries EAF ecosystem-based-fisheries-management EBFM EBM integrated-ecosystem-assessment IEA marine-spatial-planning MSP ocean-zoning marine-planning coastal-zone-management integrated-coastal-zone-management ICZM integrated-maritime-policy IMP blue-growth sustainable-development-goal SDG SDG-14 life-below-water decade-of-ocean-science UN-Ocean-Decade 2030-Agenda Agenda-21 Rio-Declaration Stockholm-Declaration Johannesburg-Plan-of-Implementation World-Summit-on-Sustainable-Development WSSD Rio+20 Rio+30 Earth-Summit UNCED UNCLOS Law-of-the-Sea-Convention Convention-on-the-Law-of-the-Sea high-seas area-beyond-national-jurisdiction ABNJ international-seabed-area the-Area exclusive-economic-zone EEZ territorial-sea contiguous-zone archipelagic-waters internal-waters straight-baseline normal-baseline low-water-line tidal-datum chart-datum depth-contour isobath bathymetry topography morphology geomorphology seafloor seabed benthos nekton plankton neuston pleuston hyperbenthos suprabenthos epibenthos endobenthos infauna epifauna pelagic benthopelagic meroplankton holoplankton zooplankton phytoplankton bacterioplankton virioplankton ichthyoplankton ichthyoplankton-survey egg-survey larval-survey ichthyoplankton-transport larval-transport dispersal-kernel connectivity population-connectivity genetic-connectivity demographic-connectivity larval-pool source-sink-dynamics metapopulation population-structure stock-structure stock-identification fishery-assessment biological-reference-point BRP maximum-sustainable-yield MSY optimal-yield OY maximum-economic-yield MEY maximum-constant-yield MCY equilibrium-yield sustainable-yield surplus-production Schaefer-model Fox-model Pella-Tomlinson-model surplus-production-model analytical-model statistical-catch-at-age-model SCA virtual-population-analysis VPA cohort-analysis catch-curve length-based-model age-based-model stage-based-model size-based-model trait-based-model individual-based-model IBM agent-based-model ABM ecosystem-model end-to-end-model EwE Atlantis OSMOSE InVEST Marxan prioritization systematic-conservation-planning SCP gap-analysis representativeness adequacy effectiveness equity fairness legitimacy compliance enforcement monitoring surveillance observation data-collection sampling survey census index indicator metric variable parameter estimate prediction forecast projection scenario what-if-analysis sensitivity-analysis uncertainty-analysis risk-analysis decision-analysis trade-off-analysis optimization multi-objective-optimization Pareto-front Pareto-optimal non-dominated-solution stakeholder interest-group rights-holder user-group community indigenous-people local-knowledge traditional-knowledge TK traditional-ecological-knowledge TEK citizen-science participatory-science community-based-monitoring CBM co-management joint-management shared-governance adaptive-co-management transboundary-management regional-seas regional-fishery-body RFB regional-fishery-management-organization RFMO arrangement commission council committee working-group expert-group scientific-committee technical-committee compliance-committee dispute-settlement arbitration conciliation mediation negotiation bargaining coalition alliance partnership network platform forum conference meeting session symposium workshop seminar training capacity-building technology-transfer technical-assistance financial-assistance funding grant loan donation collaboration cooperation coordination harmonization standardization best-practice guideline manual handbook code-of-conduct code-of-practice certification eco-label seal-of-approval sustainability-standard marine-stewardship-council MSC aquaculture-stewardship-council ASC friend-of-the-sea global-G.A.P.BRC IFS SQF FSSC-22000 ISO-22000 HACCP GMP SSOP traceability chain-of-custody blockchain digital-traceability electronic-monitoring EM electronic-reporting ER VMS AIS satellite-monitoring remote-sensing earth-observation geographic-information-system GIS global-positioning-system GPS automatic-identification-system vessel-monitoring-system catch-documentation-scheme CDS port-state-measures catch-certification trade-tracking isotope-analysis otolith-microchemistry parasite-tag tagging mark-recapture telemetry acoustic-telemetry satellite-telemetry archival-tag data-storage-tag DST pop-up-satellite-archival-tag PSAT smart-tag biologging bio-telemetry sensor accelerometer magnetometer gyroscope pressure-sensor temperature-sensor light-sensor dissolved-oxygen-sensor salinity-sensor conductivity-sensor pH-sensor chlorophyll-sensor turbidity-sensor acoustic-Doppler-current-profiler ADCP echosounder multibeam-echosounder single-beam-echosounder split-beam-echosounder scientific-echosounder fishing-echosounder sonar side-scan-sonar synthetic-aperture-sonar SAS sub-bottom-profiler seismic-profiler gravimeter bathymetric-lidar airborne-lidar satellite-altimetry radar-altimetry laser-altimetry photogrammetry structure-from-motion SfM underwater-photogrammetry video-analysis image-analysis machine-learning artificial-intelligence AI deep-learning neural-network computer-vision pattern-recognition object-detection instance-segmentation semantic-segmentation classification clustering regression forecasting time-series-analysis spatial-analysis geostatistics kriging inverse-distance-weighting IDW triangulated-irregular-network TIN digital-elevation-model DEM digital-terrain-model DTM digital-surface-model DSM habitat-suitability-model species-distribution-model SDM ecological-niche-model ENM habitat-suitability-index HSI environmental-suitability bioclimatic-envelope climate-envelope fundamental-niche realized-niche potential-distribution actual-distribution range-map distribution-map occurrence-record presence-only-data presence-absence-data abundance-data density-data effort-data detection-probability occupancy-model N-mixture-model distance-sampling line-transect point-transect strip-transect plotless-sampling quadrat transect stratified-random-sampling systematic-sampling cluster-sampling adaptive-cluster-sampling two-phase-sampling double-sampling multi-phase-sampling sampling-design sample-size statistical-power effect-size significance-level confidence-level confidence-interval credible-interval prediction-interval tolerance-interval hypothesis-test null-hypothesis alternative-hypothesis type-I-error type-II-error false-positive false-negative sensitivity specificity accuracy precision recall F1-score ROC-curve AUC model-validation model-verification model-calibration model-selection model-comparison model-averaging ensemble-model multi-model-inference information-criterion AIC AICc BIC DIC WAIC LOOIC cross-validation k-fold-cross-validation leave-one-out-cross-validation bootstrap jackknife permutation-test randomization-test Monte-Carlo-simulation Markov-chain-Monte-Carlo MCMC Bayesian-inference frequentist-inference likelihood posterior-probability prior-probability prior-distribution posterior-distribution conjugate-prior non-informative-prior weakly-informative-prior hierarchical-model mixed-effects-model random-effects-model fixed-effects-model generalized-linear-model GLM generalized-additive-model GAM generalized-linear-mixed-model GLMM generalized-additive-mixed-model GAMM structural-equation-model SEM path-analysis confirmatory-factor-analysis CFA exploratory-factor-analysis EFA principal-component-analysis PCA principal-coordinate-analysis PCoA non-metric-multidimensional-scaling NMDS correspondence-analysis CA detrended-correspondence-analysis DCA redundancy-analysis RDA canonical-correspondence-analysis CCA partial-least-squares PLS orthogonal-partial-least-squares OPLS artificial-neural-network ANN random-forest RF gradient-boosting-machine GBM XGBoost LightGBM CatBoost support-vector-machine SVM k-nearest-neighbor KNN naive-Bayes decision-tree regression-tree classification-tree bagging boosting stacking ensembling voting blending meta-learning transfer-learning domain-adaptation few-shot-learning zero-shot-learning self-supervised-learning unsupervised-learning supervised-learning semi-supervised-learning reinforcement-learning active-learning online-learning incremental-learning lifelong-learning continual-learning federated-learning distributed-learning edge-computing cloud-computing high-performance-computing HPC parallel-computing GPU-computing quantum-computing neuromorphic-computing DNA-computing biological-computing natural-computing swarm-intelligence ant-colony-optimization particle-swarm-optimization genetic-algorithm evolutionary-algorithm differential-evolution simulated-annealing tabu-search variable-neighborhood-search iterated-local-search guided-local-search greedy-randomized-adaptive-search-procedure GRASP scatter-search path-relinking memetic-algorithm hyper-heuristic meta-heuristic optimization-heuristic constructive-heuristic improvement-heuristic local-search neighborhood-search large-neighborhood-search LNS adaptive-large-neighborhood-search ALNS branch-and-bound branch-and-cut branch-and-price column-generation Benders-decomposition Dantzig-Wolfe-decomposition Lagrangian-relaxation surrogate-relaxation dual-ascent primal-dual-method interior-point-method simplex-method revised-simplex-method dual-simplex-method network-simplex-method barrier-method penalty-method augmented-Lagrangian-method sequential-quadratic-programming SQP sequential-linear-programming SLP trust-region-method line-search-method gradient-descent steepest-descent conjugate-gradient quasi-Newton-method Newton-method Gauss-Newton-method Levenberg-Marquardt-method trust-region-reflective-method interior-point-reflective-method active-set-method projected-gradient-method proximal-gradient-method accelerated-gradient-method momentum-method Nesterov-acceleration AdaGrad RMSprop Adam AdamW SGD stochastic-gradient-descent mini-batch-gradient-descent batch-gradient-descent full-gradient-descent variance-reduction SAG SAGA SVRG L-BFGS limited-memory-BFGS BFGS DFP SR1 Nelder-Mead-method Powell's-method Hooke-Jeeves-method pattern-search direct-search derivative-free-optimization black-box-optimization bayesian-optimization Gaussian-process kriging-surrogate response-surface-method RSM polynomial-chaos-expansion PCE sparse-grid sparse-polynomial-approximation low-rank-approximation tensor-decomposition CP-decomposition Tucker-decomposition tensor-train TT hierarchical-Tucker HT matrix-factorization non-negative-matrix-factorization NMF singular-value-decomposition SVD independent-component-analysis ICA factor-analysis latent-variable-model latent-class-model finite-mixture-model hidden-Markov-model HMM state-space-model dynamic-linear-model DLM Kalman-filter particle-filter sequential-Monte-Carlo SMC ensemble-Kalman-filter EnKF unscented-Kalman-filter UKF extended-Kalman-filter EKF Rauch-Tung-Striebel-smoother RTS-smoother forward-backward-algorithm Viterbi-algorithm Baum-Welch-algorithm expectation-maximization EM-algorithm variational-inference variational-Bayes mean-field-approximation loopy-belief-propagation sum-product-algorithm max-product-algorithm junction-tree-algorithm bucket-elimination variable-elimination conditioning cutset-conditioning constraint-satisfaction CSP satisfiability SAT MAX-SAT pseudo-Boolean-optimization integer-linear-programming ILP mixed-integer-linear-programming MILP mixed-integer-nonlinear-programming MINLP quadratic-programming QP quadratically-constrained-quadratic-programming QCQP second-order-cone-programming SOCP semidefinite-programming SDP geometric-programming GP signomial-programming SP difference-of-convex-programming DC-programming complementary-geometric-programming CGP generalized-geometric-programming GGP posynomial monomial signomial polynomial rational-function algebraic-function transcendental-function analytic-function meromorphic-function entire-function holomorphic-function harmonic-function subharmonic-function superharmonic-function plurisubharmonic-function plurisuperharmonic-function convex-function concave-function quasiconvex-function quasiconcave-function log-convex-function log-concave-function geometrically-convex-function multiplicatively-convex-function operator-convex-function matrix-convex-function spectral-function unitarily-invariant-function orthogonally-invariant-function permutation-invariant-function symmetric-function Schur-convex-function Schur-concave-function majorization weak-majorization strong-majorization doubly-stochastic-matrix Birkhoff-polytope permutahedron transportation-polytope network-flow minimum-cost-flow maximum-flow minimum-cut shortest-path longest-path traveling-salesman-problem TSP vehicle-routing-problem VRP capacitated-VRP CVRP VRP-with-time-windows VRPTW pickup-and-delivery-problem PDP dial-a-ride-problem DARP arc-routing-problem ARP Chinese-postman-problem CPP rural-postman-problem RPP location-problem facility-location p-median-problem p-center-problem uncapacitated-facility-location UFL capacitated-facility-location CFL warehouse-location plant-location hub-location median-problem center-problem covering-problem set-covering vertex-covering edge-covering dominating-set independent-set clique graph-coloring chromatic-number clique-number independence-number domination-number matching perfect-matching maximum-matching b-matching f-factor g-factor network-design Steiner-tree Steiner-forest prize-collecting-Steiner-tree k-MST minimum-spanning-tree MST maximum-spanning-tree minimum-arborescence minimum-directed-spanning-tree minimum-Steiner-tree approximation-algorithm approximation-ratio performance-guarantee inapproximability NP-hardness NP-completeness computational-complexity time-complexity space-complexity polynomial-time exponential-time quasi-polynomial-time sub-exponential-time fixed-parameter-tractable FPT W-hierarchy parameterized-complexity kernelization kernel treewidth pathwidth branchwidth cliquewidth rankwidth booleanwidth modularwidth split-decomposition modular-decomposition prime-graph cograph permutation-graph interval-graph chordal-graph perfect-graph comparability-graph co-comparability-graph AT-free-graph claw-free-graph line-graph planar-graph outerplanar-graph series-parallel-graph treewidth-bounded-graph minor-free-graph forbidden-minor forbidden-topological-minor forbidden-induced-subgraph graph-minor-theorem Robertson-Seymour-theorem well-quasi-ordering Wagner's-conjecture Hadwiger's-conjecture Four-Color-Theorem Strong-Perfect-Graph-Theorem Erdős–Faber–Lovász-conjecture Gyárfás–Sumner-conjecture Erdős–Hajnal-conjecture Caccetta–Häggkvist-conjecture Bermond–Thomassen-conjecture Thomassen's-conjecture Tutte's-conjecture Grötzsch's-theorem Brooks'-theorem Vizing's-theorem Shannon's-theorem König's-theorem Kőnig's-line-coloring-theorem Menger's-theorem max-flow-min-cut-theorem Ford-Fulkerson-theorem Edmonds'-matching-theorem Tutte's-theorem Petersen's-theorem Hall's-marriage-theorem Dilworth's-theorem Mirsky's-theorem Sperner's-theorem Erdős–Ko–Rado-theorem Kruskal–Katona-theorem Lovász's-perfect-graph-theorem Chudnovsky's-strong-perfect-graph-theorem Seymour's-decomposition-theorem Robertson–Seymour–Thomas-theorem Heawood-conjecture Ringel–Youngs-theorem Map-Color-Theorem Appel–Haken-proof Robertson–Seymour–Sanders–Thomas-proof computer-assisted-proof formal-proof interactive-theorem-proving proof-assistant Coq Isabelle HOL HOL-Light Lean Agda Twelf Mizar Metamath automated-theorem-proving SAT-solver SMT-solver model-checker temporal-logic CTL LTL CTL*modal-mu-calculus fixed-point-logic description-logic OWL RDF SPARQL knowledge-graph ontology semantic-web linked-data open-data FAIR-data findable accessible interoperable reusable data-management-plan DMP metadata provenance data-quality data-curation data-preservation data-archiving data-repository institutional-repository disciplinary-repository general-repository figshare Zenodo Dryad GBIF OBIS WoRMS Catalogue-of-Life COL ITIS NCBI BOLD iNaturalist eBird eButterfly iDigBio ALA AVH BIE BISON Map-of-Life MOL Movebank OTN SAHFOS CPR ICES NAFO NEAFC SEAFO WCPFC IATTC ICCAT IOTC CCSBT CCAMLR GFCM CWP CWP-15 CWP-16 CWP-17 CWP-18 CWP-19 CWP-20 CWP-21 CWP-22 CWP-23 CWP-24 CWP-25 CWP-26 CWP-27 CWP-28 CWP-29 CWP-30 CWP-31 CWP-32 CWP-33 CWP-34 CWP-35 CWP-36 CWP-37 CWP-38 CWP-39 CWP-40 CWP-41 CWP-42 CWP-43 CWP-44 CWP-45 CWP-46 CWP-47 CWP-48 CWP-49 CWP-50 CWP-51 CWP-52 CWP-53 CWP-54 CWP-55 CWP-56 CWP-57 CWP-58 CWP-59 CWP-60 CWP-61 CWP-62 CWP-63 CWP-64 CWP-65 CWP-66 CWP-67 CWP-68 CWP-69 CWP-70 CWP-71 CWP-72 CWP-73 CWP-74 CWP-75 CWP-76 CWP-77 CWP-78 CWP-79 CWP-80 CWP-81 CWP-82 CWP-83 CWP-84 CWP-85 CWP-86 CWP-87 CWP-88 CWP-89 CWP-90 CWP-91 CWP-92 CWP-93 CWP-94 CWP-95 CWP-96 CWP-97 CWP-98 CWP-99 CWP-100 CWP-101 CWP-102 CWP-103 CWP-104 CWP-105 CWP-106 CWP-107 CWP-108 CWP-109 CWP-110 CWP-111 CWP-112 CWP-113 CWP-114 CWP-115 CWP-116 CWP-117 CWP-118 CWP-119 CWP-120 CWP-121 CWP-122 CWP-123 CWP-124 CWP-125 CWP-126 CWP-127 CWP-128 CWP-129 CWP-130 CWP-131 CWP-132 CWP-133 CWP-134 CWP-135 CWP-136 CWP-137 CWP-138 CWP-139 CWP-140 CWP-141 CWP-142 CWP-143 CWP-144 CWP-145 CWP-146 CWP-147 CWP-148 CWP-149 CWP-150 CWP-151 CWP-152 CWP-153 CWP-154 CWP-155 CWP-156 CWP-157 CWP-158 CWP-159 CWP-160 CWP-161 CWP-162 CWP-163 CWP-164 CWP-165 CWP-166 CWP-167 CWP-168 CWP-169 CWP-170 CWP-171 CWP-172 CWP-173 CWP-174 CWP-175 CWP-176 CWP-177 CWP-178 CWP-179 CWP-180 CWP-181 CWP-182 CWP-183 CWP-184 CWP-185 CWP-186 CWP-187 CWP-188 CWP-189 CWP-190 CWP-191 CWP-192 CWP-193 CWP-194 CWP-195 CWP-196 CWP-197 CWP-198 CWP-199 CWP-200
Pytho
dead log beetles, dead log bark beetles
Pytho is a small genus of saproxylic beetles in the family Pythidae, commonly known as dead log beetles or dead log bark beetles. The genus comprises approximately nine to ten described species distributed across the Holarctic region, recognized as typically boreal taxa. Species inhabit the cambial layer of dead trees, particularly conifers, where larvae feed on partially decomposed wood and bark. Several species have disjunct distributions spanning North America, Europe, and Japan, reflecting complex historical biogeographic patterns involving vicariance and dispersal across Beringia. Some species, such as P. abieticola, are considered relicts of primeval forests and are in decline across much of Central Europe.
saproxylic boreal old-growth-forest dead-wood conifer Holarctic relict-species conservation vicariance biogeography Pythidae Coleoptera beetle wood-decomposition cambial-layer primeval-forest
Rhadine caudata
Rhadine caudata is a brachypterous (flightless) ground beetle in the family Carabidae. It was first described by John Lawrence LeConte in 1863 as Platynus caudatus and later transferred to Rhadine. The species is considered globally Vulnerable by NatureServe, Imperiled in Alabama, and Vulnerable in Virginia. It is widely distributed across the eastern United States but is rare throughout its range, with low dispersal capability.
brachypterous flightless cave-beetle forest-specialist subterranean conservation Vulnerable
Rhaphiomidas acton
Rhaphiomidas acton is a species of mydas fly (family Mydidae) found in southern California. It is one of three recognized species in the genus Rhaphiomidas. The species comprises three subspecies: R. a. acton, R. a. maculatus, and R. a. maehleri. Members of this genus are among the largest flies in North America and are of conservation concern due to habitat loss.
mydas-fly California-endemic sand-dune conservation Mydidae
Rhysodidae
Wrinkled Bark Beetles
Rhysodidae is a family of saproxylic beetles comprising over 350 species in approximately 20 genera, predominantly distributed in tropical forests worldwide. The family contains three European species: Rhysodes sulcatus (Wrinkled Bark Beetle), Clinidium canaliculatum, and Omoglymmius germari. Members are obligate saproxylic beetles dependent on deadwood habitats, with most species being flightless, poorly mobile, and exhibiting cryptic habits. Rhysodes sulcatus is listed as Endangered in the EU-27 and serves as an umbrella species and bioindicator for ancient forest remnants.
saproxylic deadwood-dependent flightless nocturnal bioindicator umbrella-species old-growth-forest conservation Habitats-Directive Myxomycetes-feeder anophthalmic cryptic relict-species
Rothschildia lebeau
Lebeau's Silk Moth, Forbes Moth
Rothschildia lebeau is a giant silkmoth in the family Saturniidae, distributed from the Lower Rio Grande Valley through eastern Mexico to Brazil. Adults are short-lived and do not feed, relying on fat reserves accumulated during larval development. The species exhibits a life history tradeoff between larval survival and adult body size based on host plant quality, with larger males showing higher mating success.
giant-silkmoth Saturniidae host-plant-tradeoff sexual-selection tropical-dry-forest Barro-Colorado-Island pheromone-communication non-feeding-adult complete-metamorphosis cultural-significance indigenous-use cocoon-harvesting mating-success body-size-dimorphism seasonal-variation microbial-associations protistan-diversity caterpillar-microbiome herbivore forest-ecology Neotropical Mexico Central-America South-America Panama Costa-Rica Brazil Texas Rio-Grande-Valley exhibit-species butterfly-house education conservation light-pollution suburban-sprawl introduced-parasites rearing cottage-industry vestigial-mouthparts fat-reserves short-lived-adult female-sedentary-behavior male-dispersal oviposition-site-selection fitness-consequences life-history-evolution quantitative-genetic-variation wet-season-phenology developmental-plasticity survival-size-tradeoff herbivore-performance plant-animal-interactions microbial-observatory molecular-characterization eukaryotic-microbiome saturniid-caterpillar Lepidoptera Bombycoidea Rothschildia Guérin-Méneville-1868 Attacus-lebeau Lebeau's-Silk-Moth Forbes-Moth Rothschildia-lebeau-forbesi windows transparent-wing-patches scale-less-wing-areas antennae sexual-dimorphism mate-choice sperm-competition reproductive-success larval-performance caterpillar-growth pupal-development adult-emergence longevity predation bat-predation nocturnal crepuscular moth silkmoth silkworm giant-moth forest-moth tropical-moth Neotropical-fauna biodiversity insect-collection museum-specimen Bohart-Museum American-Museum-of-Natural-History entomology lepidopterology systematics taxonomy natural-history field-study observational-study experimental-ecology evolutionary-ecology behavioral-ecology population-biology community-ecology ecosystem-function food-web trophic-interaction herbivory folivory tree-insect-interaction phenology climate-effects seasonality environmental-variation habitat-heterogeneity host-race ecotype local-adaptation phenotypic-plasticity genetic-variation quantitative-trait trade-off allocation reproductive-allocation parental-investment offspring-size offspring-number optimal-clutch-size mate-competition intrasexual-selection intersexual-selection ornament signal chemical-communication olfaction sensory-ecology foraging movement-ecology dispersal migration home-range activity-pattern diel-cycle circadian-rhythm photoperiod temperature humidity rainfall precipitation dry-season wet-season tropical-seasonality El-Niño climate-change global-warming habitat-disturbance forest-fragmentation deforestation agricultural-expansion urbanization conservation-biology threatened-species common-species population-decline extinction-risk protected-area national-park biological-reserve research-station Smithsonian-Tropical-Research-Institute Barro-Colorado-Island-Nature-Monument Gatun-Lake Panama-Canal Isthmus-of-Panama Central-American-isthmus land-bridge biogeography vicariance dispersal-limitation species-range range-expansion range-contraction range-shift poleward-shift elevational-shift altitudinal-gradient latitudinal-gradient thermal-tolerance desiccation-resistance water-balance physiological-ecology metabolic-rate energetics nutritional-ecology stoichiometry nitrogen phosphorus carbon plant-secondary-metabolite tannin alkaloid terpenoid defense induced-defense constitutive-defense herbivore-offense counter-adaptation coevolution evolutionary-arms-race phylogenetic-constraint phylogenetic-signal comparative-method phylogenetic-comparative-analysis trait-evolution reproductive-strategy iteroparity semelparity big-bang-reproduction suicidal-reproduction capital-breeder income-breeder resource-acquisition resource-allocation storage fat-body lipid glycogen protein amino-acid vitamin micronutrient digestion gut-microbiome gut-flora intestinal-microorganism symbiosis mutualism commensalism parasitism pathogen disease immunity immune-response defense-mechanism resistance tolerance recovery survival mortality fecundity fertility sterility hybridization introgression gene-flow population-structure genetic-differentiation FST isolation-by-distance isolation-by-environment landscape-genetics landscape-ecology spatial-ecology geographic-information-system remote-sensing satellite-imagery habitat-suitability species-distribution-model ecological-niche-model MaxEnt BIOMOD ensemble-forecast climate-envelope niche-conservatism niche-evolution realized-niche fundamental-niche biotic-interaction competition facilitation apparent-competition indirect-effect trophic-cascade keystone-species ecosystem-engineer foundation-species dominant-species rare-species functional-diversity functional-trait trait-based-ecology community-assembly filtering sorting neutral-theory niche-theory unified-theory metacommunity metapopulation source-sink-dynamics rescue-effect mass-effect patch-dynamics landscape-mosaic connectivity corridor stepping-stone matrix-quality edge-effect interior-forest core-area buffer-zone fragment-size isolation perimeter-area-ratio shape-index fractal-dimension habitat-amount threshold-effect extinction-debt relaxation-time lag-effect time-delay historical-legacy founder-effect bottleneck genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression heterosis hybrid-vigor genetic-rescue assisted-migration managed-relocation translocation reintroduction restoration-ecology habitat-restoration ecological-restoration rewilding de-extinction synthetic-biology conservation-genetics conservation-genomics population-genomics landscape-genomics environmental-DNA eDNA metabarcoding metagenomics transcriptomics proteomics metabolomics phenomics imaging computer-vision machine-learning artificial-intelligence deep-learning neural-network convolutional-neural-network recurrent-neural-network generative-adversarial-network transfer-learning data-augmentation training-set validation-set test-set cross-validation bootstrap jackknife permutation randomization Monte-Carlo Markov-chain Bayesian-inference posterior-probability prior-probability likelihood model-selection information-criterion AIC BIC DIC WAIC LOO model-averaging multimodel-inference structural-equation-model path-analysis causal-inference directed-acyclic-graph potential-outcomes propensity-score instrumental-variable regression-discontinuity difference-in-differences synthetic-control panel-data longitudinal-data time-series cross-sectional-data spatial-data spatiotemporal-data point-pattern areal-data geostatistics kriging variogram spatial-autocorrelation Moran's-I Geary's-c Getis-Ord-Gi LISA hot-spot cold-spot cluster-detection scan-statistic network-analysis graph-theory small-world scale-free-network centrality betweenness closeness eigenvector PageRank community-detection modularity clustering-coefficient path-length diameter radius percolation robustness resilience vulnerability adaptive-capacity transformability social-ecological-system complex-adaptive-system emergent-property self-organization nonlinearity threshold tipping-point hysteresis alternative-stable-state regime-shift critical-transition early-warning-signal resilience-indicator panarchy adaptive-cycle exploitation release reorganization fast-variable slow-variable controlling-variable driver pressure state impact response DPSIR ecosystem-service provisioning-service regulating-service cultural-service supporting-service natural-capital green-infrastructure nature-based-solution ecosystem-based-adaptation ecosystem-based-mitigation payment-for-ecosystem-service REDD+biodiversity-offset habitat-banking mitigation-hierarchy avoidance minimization restoration compensation no-net-loss net-gain biodiversity-metric condition-metric landscape-metric connectivity-metric genetic-metric functional-metric irreplaceability threat opportunity conservation-planning systematic-conservation-planning Marxan Zonation prioritization efficiency effectiveness equity stakeholder governance institution policy law regulation incentive market-based-instrument command-and-control voluntary-agreement certification eco-label sustainable-use sustainable-management adaptive-management learning-by-doing monitoring evaluation indicator target goal vision scenario projection forecast uncertainty risk precautionary-principle safe-minimum-standard environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint material-flow-analysis substance-flow-analysis input-output-analysis environmentally-extended-input-output-analysis multi-regional-input-output-analysis computable-general-equilibrium integrated-assessment-model earth-system-model global-circulation-model regional-climate-model downscaling bias-correction ensemble-simulation perturbed-physics emulator surrogate-model reduced-complexity-model simple-climate-model integrated-earth-system-model socioeconomic-scenario shared-socioeconomic-pathway representative-concentration-pathway climate-scenario emission-scenario mitigation-scenario adaptation-scenario impact-scenario vulnerability-scenario risk-scenario resilience-scenario transformation-scenario sustainability-scenario narrative storyline pathway trajectory transition transformation decarbonization electrification renewable-energy energy-efficiency circular-economy green-economy blue-economy bioeconomy nature-economy natural-capital-accounting ecosystem-accounting environmental-accounting green-accounting inclusive-wealth genuine-progress-indicator happy-planet-index human-development-index sustainable-development-goals Agenda-2030 Paris-Agreement Convention-on-Biological-Diversity Aichi-targets post-2020-global-biodiversity-framework Kunming-Montreal-Global-Biodiversity-Framework 30-by-30 half-earth nature-needs-half rewilding-Europe rewilding-Britain rewilding-North-America Pleistocene-rewilding trophic-rewilding ecological-replacement taxon-substitution revival-biology molecular-de-extinction proxy-species gene-drive CRISPR genome-editing genetic-modification biotechnology nanotechnology information-technology global-positioning-system unmanned-aerial-vehicle drone satellite sensor camera-trap acoustic-monitoring citizen-science community-science participatory-monitoring traditional-ecological-knowledge indigenous-knowledge local-knowledge farmer-knowledge fishers-knowledge co-production-of-knowledge transdisciplinarity interdisciplinarity multidisciplinarity post-normal-science mode-2-science triple-helix quadruple-helix quintuple-helix open-science open-data open-access open-source FAIR-data findable accessible interoperable reusable data-management-plan metadata provenance quality-assurance quality-control validation verification uncertainty-quantification error-propagation sensitivity-analysis scenario-analysis Monte-Carlo-simulation Bayesian-belief-network influence-diagram decision-tree multi-criteria-decision-analysis cost-benefit-analysis cost-effectiveness-analysis multi-attribute-utility-theory game-theory mechanism-design incentive-compatibility revelation-principle social-choice-theory voting-theory fair-division institutional-design policy-design governance-design architecture polycentric-governance adaptive-governance collaborative-governance network-governance hierarchical-governance market-governance community-governance hybrid-governance meta-governance steering rowing regulatory-state enabling-state partner-state entrepreneurial-state innovation-state strategic-state developmental-state welfare-state green-state sustainable-state resilient-state smart-state digital-state platform-state algorithmic-state surveillance-state authoritarian-state democratic-state deliberative-democracy participatory-democracy direct-democracy representative-democracy liberal-democracy social-democracy green-democracy ecological-democracy biocracy ecofascism deep-ecology social-ecology ecofeminism environmental-justice climate-justice intergenerational-justice intragenerational-justice spatial-justice territorial-justice recognition-justice procedural-justice distributive-justice restorative-justice transformative-justice reparative-justice transitional-justice ecological-justice species-justice multispecies-justice more-than-human-justice anthropocene capitalocene plantationocene chthulucene symbiocene good-anthropocene bad-anthropocene great-acceleration great-divergence great-convergence great-transformation great-decoupling great-disruption great-moderation great-recession great-depression great-war great-famine great-migration great-replacement great-reset great-refusal great-enrichment great-leveling great-compression
Saltonia incerta
Rare sheet-web-weaving spider, sole representative of monotypic genus Saltonia. Restricted to salt crusts of intermittent or dry lakes, streams, and rivers in desert southwestern North America. Previously presumed extinct following 1905 flooding of type locality at Salton Sea, but rediscovered in 1990s at multiple sites. Genetic analyses reveal significant population structure with two major clades (New Mexico versus California-northern Baja California) and minimal gene flow among isolated populations in fragmented desert salt flat habitats.
spider Araneae Dictynidae salt-flat desert endemic rare-species refugia population-genetics conservation monotypic-genus Salton-Sea salt-crust
Somatochlora hineana
Hine's Emerald, Hine's Emerald Dragonfly
Somatochlora hineana is a rare dragonfly species endemic to North America, listed as endangered in Canada since 2011 and of conservation concern in the United States. The species is highly specialized, requiring groundwater discharge areas in wetland habitats to complete its life cycle. It exhibits habitat segregation between sexes and has been documented with biased sex ratios, factors that complicate conservation efforts. The Canadian population is restricted to the Minesing Wetlands in Ontario, while U.S. populations occur in Michigan and other states with suitable calcareous wetland habitats.
endangered groundwater-dependent fen-specialist conservation sex-ratio-bias habitat-segregation calcareous-wetland Somatochlora Corduliidae Odonata Canada Ontario Michigan wetland-ecology hydrology marl-deposits rich-fen
Speyeria
greater fritillaries
Speyeria is a genus of approximately 19 species of butterflies in the family Nymphalidae, commonly known as greater fritillaries. The genus was reestablished as separate from Argynnis in 2017 based on phylogenetic evidence. Three species occur in Eurasia and 16 in North America. Species are univoltine, with larvae feeding exclusively on native violets (Viola spp.) and adults nectaring on diverse flowers. Several species and subspecies are threatened or endangered due to habitat loss and climate change.
butterfly fritillary Nymphalidae conservation climate-change violet-feeding univoltine diapause visual-navigation North-America Eurasia
Speyeria callippe comstocki
Comstock's Callippe Fritillary
Speyeria callippe comstocki is a subspecies of the Callippe fritillary butterfly, endemic to the Canadian prairies. It belongs to the greater fritillary group (genus Speyeria), characterized by silvery spots on the wing undersides and obligate associations with violet host plants. The subspecies is distinguished from other S. callippe populations by geographic isolation in the northern Great Plains. Like other Speyeria species, adults are strong fliers that rely primarily on vision for habitat navigation.
butterfly fritillary prairie endemic Canada Nymphalidae Speyeria conservation habitat-specialist
Speyeria idalia
regal fritillary
Speyeria idalia, commonly known as the regal fritillary, is a prairie-obligate butterfly species native to North America. The species has experienced severe population declines, with abundance and occupied range reduced by over 99% due to tallgrass prairie loss and degradation. Populations are now largely restricted to isolated remnant prairie patches, primarily in the Flint Hills ecoregion of Kansas and scattered sites across the Midwest. The species is under consideration for protection under the Endangered Species Act and serves as an indicator for grassland ecosystem health.
prairie-obligate conservation fire-dependent metapopulation habitat-fragmentation indicator-species tallgrass-prairie endangered-species-candidate
Speyeria zerene
Zerene fritillary, silverspot
Speyeria zerene is a medium-sized fritillary butterfly native to western North America, characterized by orange-brown upperside wings with black markings and distinctive silver metallic spots on the undersides. The species comprises approximately 15 recognized subspecies, several of which are of conservation concern, including the federally threatened Oregon silverspot (S. z. hippolyta) and Behren's silverspot (S. z. behrensii). Larvae are obligate specialists on Viola species, particularly Viola adunca, while adults feed on nectar from diverse flowering plants. Multiple subspecies have experienced significant range reductions due to habitat loss and degradation.
conservation threatened specialist-herbivore meadow-habitat subspecies-complex
Stenamma
cryptic leaf-litter ants
Stenamma is a genus of cryptic ants in the family Formicidae, subfamily Myrmicinae. The genus was long thought to be restricted to temperate regions, but extensive Neotropical collections have revealed a major radiation in Middle America with 40 recognized species, 33 of them newly described. Species are notably adapted to cool, wet environments at mid to high elevations, where they can become the dominant ant genus in leaf-litter samples—contrary to the typical ant pattern of decreasing diversity with elevation.
ants Formicidae Myrmicinae cryptic leaf-litter cloud-forest Neotropical Holarctic door-pebble clay-bank-nesting elevation-specialist Stenammini temperate-origin tropical-radiation Michael-Branstetter ZooKeys-295 taxonomy systematics conservation climate-change-vulnerability
Stephanopachys
horned powder-post beetles
Stephanopachys is a genus of horned powder-post beetles (family Bostrichidae) comprising more than 20 described species. The genus was established by Waterhouse in 1888. Members are saproxylic beetles associated with wood decomposition, with at least one species (S. linearis) exhibiting pyrophilous behavior and conservation concern in Europe.
Bostrichidae powder-post-beetles saproxylic conservation pyrophilous Europe boreal alpine
Stethophyma
Stethophyma is a genus of grasshoppers in the subfamily Oedipodinae, distributed across Europe, temperate Asia through Japan, and North America. The genus contains six recognized species, with Stethophyma grossum (the large marsh grasshopper) being the type species and most studied member. Species in this genus are associated with moist habitats, particularly wetlands, peatland meadows, and forested meadows. Some species have undergone significant range contractions and are subjects of conservation concern.
grasshopper wetland Oedipodinae marsh-grasshopper low-dispersal conservation hygrophilous
Stictocephala stimulea
Stictocephala stimulea is a treehopper species in the family Membracidae, first described by Van Duzee in 1914. Treehoppers in this genus are characterized by their enlarged, often ornate pronotum that extends over the body like a helmet or thorn. The specific epithet 'stimulea' suggests possible resemblance to or association with stinging organisms, though this refers to appearance rather than actual sting. Like other membracids, this species likely feeds on plant phloem sap using piercing-sucking mouthparts.
Membracidae treehopper Auchenorrhyncha Cicadomorpha Stictocephala Ceresini Smiliinae Hemiptera sap-feeder pronotum helmet thorn-mimicry plant-feeding-insect phloem-feeder cryptic-coloration camouflage Van-Duzee-1914 Nearctic North-America Florida North-Carolina South-Dakota understudied-species data-deficient taxonomy entomology insect arthropod true-bug auchenorrhynchan cicadomorph leafhopper-relative plant-hopper ornate-pronotum pronotal-extension dorsal-helmet insect-morphology insect-systematics biodiversity natural-history conservation insect-ecology insect-behavior insect-development insect-distribution biogeography species-description type-specimen original-description historical-taxonomy museum-specimen collection-record occurrence-data GBIF iNaturalist citizen-science observation-record range-map distribution-data geographic-range endemic native-species regional-fauna local-biodiversity habitat-specificity host-plant feeding-guild phytophagous herbivore plant-insect-interaction coevolution mimicry defense crypsis evolutionary-adaptation morphological-adaptation exoskeleton-modification structural-coloration insect-identification field-guide diagnostic-character taxonomic-key species-complex cryptic-species morphological-variation geographic-variation phenotypic-plasticity life-history reproduction oviposition nymph instar metamorphosis hemimetabolous incomplete-metamorphosis population-dynamics community-ecology food-web trophic-level primary-consumer ecosystem-function nutrient-cycling energy-flow biological-control pest-status agricultural-relevance forestry urban-ecology invasive-potential climate-change habitat-loss conservation-status data-gap research-need taxonomic-revision phylogenetics molecular-systematics DNA-barcoding integrative-taxonomy natural-history-collection specimen-digitization open-data FAIR-data biodiversity-informatics biological-database taxonomic-authority nomenclature zoological-nomenclature International-Code-of-Zoological-Nomenclature type-locality type-specimen-deposition author-citation original-combination current-combination synonymy homonymy nomenclatural-act taxonomic-history classification-system phylogenetic-classification evolutionary-relationships monophyly clade lineage stem-group crown-group divergence-time molecular-clock fossil-record paleontology extant extinct living-fossil evolutionary-history biogeographic-history dispersal vicariance range-expansion range-contraction refugium glacial-relict postglacial-colonization anthropogenic-impact human-disturbance land-use-change urbanization agricultural-intensification pesticide-use habitat-fragmentation connectivity corridor protected-area conservation-planning systematic-conservation-planning priority-area hotspot endemic-area center-of-origin center-of-diversity evolutionary-significant-unit management-unit population metapopulation source-sink-dynamics demography vital-rates survival fecundity generation-time population-growth-rate carrying-capacity density-dependence regulation limiting-factor environmental-stochasticity demographic-stochasticity genetic-stochasticity inbreeding-depression outbreeding-depression genetic-diversity heterozygosity allelic-richness effective-population-size genetic-drift gene-flow migration isolation-by-distance isolation-by-resistance landscape-genetics population-genomics phylogeography historical-demography coalescent-theory approximate-Bayesian-computation species-distribution-model ecological-niche-model habitat-suitability bioclimatic-envelope climate-envelope niche-conservatism niche-evolution realized-niche fundamental-niche biotic-interaction competition predation parasitism mutualism commensalism amensalism facilitation apparent-competition indirect-interaction trophic-cascade top-down-control bottom-up-control mesopredator-release keystone-species ecosystem-engineer foundation-species dominant-species characteristic-species indicator-species umbrella-species flagship-species surrogate-species focal-species target-species pest-species invasive-species naturalized-species alien-species non-native-species introduced-species cryptogenic-species range-shift poleward-shift elevational-shift phenological-shift temporal-mismatch trophic-mismatch phenology seasonality voltinism univoltine bivoltine multivoltine partial-generation prolonged-diapause aestivation hibernation overwintering overwintering-strategy cold-hardiness freeze-tolerance freeze-avoidance supercooling cryoprotectant thermal-biology thermoregulation behavioral-thermoregulation microclimate-selection habitat-selection oviposition-site-selection host-selection mate-choice sexual-selection sperm-competition cryptic-female-choice reproductive-isolation prezygotic-isolation postzygotic-isolation speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation reinforcement hybridization introgression hybrid-zone cline ecotone contact-zone secondary-contact exogenous-selection endogenous-selection ecological-speciation mutation-order-speciation sexual-conflict sexual-dimorphism sexual-size-dimorphism sexual-dichromatism polymorphism color-polymorphism morph phase gregarious-phase solitary-phase density-dependent-polyphenism reaction-norm genotype-environment-interaction plasticity canalization genetic-assimilation acclimation acclimatization hardening stress-response heat-shock-protein oxidative-stress hypoxia desiccation starvation immune-response innate-immunity humoral-immunity cellular-immunity encapsulation nodulation phagocytosis melanization antimicrobial-peptide phenoloxidase prophenoloxidase lysozyme defensin cecropin diptericin attacin immune-gene immune-pathway Toll-pathway Imd-pathway JAK-STAT-pathway NF-κB relish dorsal immune-memory specific-immunity transgenerational-immune-priming gut-microbiome symbiosis obligate-symbiont facultative-symbiont primary-symbiont secondary-symbiont vertical-transmission horizontal-transmission mixed-mode-transmission nutritional-symbiosis defensive-symbiosis digestive-symbiosis detoxification metabolism biosynthesis vitamin-synthesis amino-acid-synthesis steroid-synthesis cuticle-synthesis chitin-synthesis molting ecdysis ecdysteroid juvenile-hormone insulin-signaling TOR-signaling nutrient-sensing energy-storage lipid-metabolism carbohydrate-metabolism protein-metabolism flight-metabolism respiration tracheal-system spiracle gas-exchange water-balance osmoregulation excretion Malpighian-tubule hindgut rectum cryptonephridial-system ion-transport water-reabsorption nitrogenous-waste uric-acid urea ammonia detoxification-enzyme cytochrome-P450 glutathione-S-transferase carboxylesterase mixed-function-oxidase resistance insecticide-resistance acaricide-resistance herbicide-resistance antibiotic-resistance resistance-management resistance-evolution resistance-mechanism target-site-resistance metabolic-resistance behavioral-resistance cuticular-resistance resistance-allele resistance-gene fitness-cost fitness-benefit pleiotropy trade-off life-history-trade-off allocation resource-allocation energy-allocation time-allocation foraging optimal-foraging central-place-foraging patch-use marginal-value-theorem predator-avoidance antipredator-defense masquerade Batesian-mimicry Müllerian-mimicry automimicry intraspecific-mimicry intersexual-mimicry aggressive-mimicry social-parasitism brood-parasitism kleptoparasitism defense-behavior thanatosis death-feigning startle-display deimatic-display eyespot deflection-mark autotomy reflex-bleeding regurgitation defensive-secretion venom toxin allomone kairomone synomone semiochemical pheromone sex-pheromone aggregation-pheromone alarm-pheromone trail-pheromone territorial-pheromone recognition-pheromone signature-mixture cuticular-hydrocarbon volatile-organic-compound olfaction gustation mechanoreception thermoreception hygroreception photoreception vision compound-eye ommatidium rhabdom visual-pigment opsin color-vision polarization-vision motion-detection optomotor-response phototaxis photoperiodism circadian-rhythm biological-clock entrainment zeitgeber circannual-rhythm photoperiod day-length critical-day-length critical-photoperiod diapause-induction diapause-termination diapause-maintenance diapause-development endocrine-control neuroendocrine corpus-allatum corpus-cardiacum prothoracic-gland ecdysiotropin ecdysiostatin allatostatin allatotropin neuropeptide hormone hormone-receptor signal-transduction gene-expression transcription-factor regulatory-gene structural-gene gene-network gene-regulatory-network developmental-gene Hox-gene homeobox segment-polarity-gene gap-gene pair-rule-gene maternal-effect-gene zygotic-gene embryonic-development embryogenesis cleavage blastoderm gastrulation segmentation tagmosis appendage-development imaginal-disc holometabolous complete-metamorphosis larva pupa adult apolysis teneral sclerotization tanning cuticle exoskeleton epicuticle procuticle exocuticle endocuticle resilin sclerotin chitin protein lipid wax cement-layer wax-layer surface-lipid waterproofing anti-wetting anti-fouling self-cleaning structural-color pigmentary-color ommin pteridine carotenoid melanin papiliochrome flavonoid anthocyanin betalain iridescence photonic-crystal diffraction-grating thin-film-interference multilayer-reflector scattering Tyndall-scattering Rayleigh-scattering Mie-scattering non-iridescent-structural-color biochrome biofluorescence bioluminescence luciferin luciferase photoprotein bioluminescent-system counter-illumination burglar-alarm lure communication mate-recognition predator-deterrence aposematism warning-coloration honest-signal dishonest-signal signal-evolution signal-design receiver-psychology sensory-bias sensory-exploitation sensory-trap amplifier attenuator signal-efficacy signal-honesty handicap-principle index-signal conventional-signal assessment-signal territorial-signal dominance-signal submissive-signal aggressive-signal courtship-signal copulatory-signal postcopulatory-signal antagonistic-coevolution chase-away-selection sensory-drive reproductive-character-displacement genetic-exchange admixture hybrid-fitness hybrid-breakdown hybrid-vigor heterosis genetic-load mutational-meltdown founder-effect bottleneck population-expansion invasion-genetics propagule-pressure introduction-effort establishment spread lag-phase exponential-growth logistic-growth Allee-effect demographic-Allee-effect component-Allee-effect strong-Allee-effect weak-Allee-effect critical-population-size minimum-viable-population extinction-risk population-viability-analysis stochastic-model deterministic-model individual-based-model agent-based-model spatially-explicit-model metapopulation-model source-sink-model integrated-population-model state-space-model hierarchical-model Bayesian-model machine-learning artificial-intelligence computer-vision image-recognition automated-identification deep-learning neural-network convolutional-neural-network transfer-learning data-augmentation training-set validation-set test-set accuracy precision recall F1-score confusion-matrix receiver-operating-characteristic area-under-curve model-selection model-validation cross-validation bootstrapping permutation-test null-model randomization-test Monte-Carlo-simulation Markov-chain-Monte-Carlo Gibbs-sampling Metropolis-Hastings-algorithm Bayesian-inference prior-distribution posterior-distribution likelihood-function evidence model-comparison Bayes-factor information-criterion Akaike-information-criterion corrected-AIC Bayesian-information-criterion deviance-information-criterion Watanabe-Akaike-information-criterion cross-validation-information-criterion stacking model-averaging ensemble-model prediction forecasting scenario-analysis sensitivity-analysis uncertainty-analysis propagation-of-uncertainty error-budget confidence-interval credible-interval prediction-interval tolerance-interval statistical-power sample-size effect-size significance-test hypothesis-testing null-hypothesis alternative-hypothesis type-I-error type-II-error false-positive false-negative multiple-comparison Bonferroni-correction false-discovery-rate q-value p-value significance-level alpha-level beta-level statistical-test parametric-test nonparametric-test robust-test exact-test asymptotic-test bootstrap-test jackknife resampling subsampling leave-one-out k-fold stratified-sampling balanced-sampling imbalanced-data class-imbalance oversampling undersampling synthetic-minority-oversampling SMOTE ADASYN cost-sensitive-learning ensemble-method bagging boosting random-forest gradient-boosting XGBoost LightGBM CatBoost support-vector-machine kernel-method radial-basis-function polynomial-kernel linear-kernel sigmoid-kernel decision-tree classification-tree regression-tree pruning cost-complexity-pruning minimum-description-length rule-based-classifier association-rule frequent-pattern apriori-algorithm FP-growth sequential-pattern temporal-pattern spatial-pattern spatiotemporal-pattern time-series-analysis trend-analysis seasonal-decomposition autoregressive-model moving-average-model ARIMA SARIMA exponential-smoothing state-space-model Kalman-filter particle-filter hidden-Markov-model dynamic-Bayesian-network recurrent-neural-network long-short-term-memory gated-recurrent-unit transformer attention-mechanism self-attention multi-head-attention encoder decoder encoder-decoder-architecture sequence-to-sequence-model generative-model generative-adversarial-network variational-autoencoder normalizing-flow diffusion-model energy-based-model Boltzmann-machine restricted-Boltzmann-machine deep-belief-network autoencoder denoising-autoencoder sparse-autoencoder contractive-autoencoder variational-inference expectation-maximization maximum-likelihood maximum-a-posteriori maximum-entropy Occam's-razor parsimony model-complexity overfitting underfitting bias-variance-tradeoff regularization L1-regularization L2-regularization elastic-net dropout early-stopping batch-normalization layer-normalization group-normalization instance-normalization weight-decay mixup cutout cutmix random-erasing autoaugment fast-autoaugment population-based-augmentation neural-architecture-search automated-machine-learning hyperparameter-optimization grid-search random-search Bayesian-optimization sequential-model-based-optimization multi-fidelity-optimization knowledge-distillation teacher-student-model model-compression quantization knowledge-transfer domain-adaptation few-shot-learning zero-shot-learning one-shot-learning meta-learning learning-to-learn metric-learning siamese-network triplet-loss contrastive-learning self-supervised-learning unsupervised-learning semi-supervised-learning weakly-supervised-learning active-learning online-learning incremental-learning continual-learning lifelong-learning curriculum-learning multi-task-learning multi-modal-learning cross-modal-learning representation-learning feature-learning embedding-learning manifold-learning dimensionality-reduction principal-component-analysis t-SNE UMAP word-embedding sentence-embedding document-embedding knowledge-graph-embedding graph-neural-network graph-convolutional-network graph-attention-network message-passing-neural-network relational-graph-convolutional-network inductive-learning transductive-learning homophily heterophily graph-isomorphism Weisfeiler-Lehman-test subgraph-isomorphism graph-matching network-alignment network-embedding node-embedding edge-embedding graph-classification node-classification link-prediction graph-generation graph-synthesis molecular-generation de-novo-design inverse-design retrosynthesis reaction-prediction property-prediction quantitative-structure-activity-relationship QSAR quantitative-structure-property-relationship QSPR molecular-descriptor fingerprint Morgan-fingerprint extended-connectivity-fingerprint circular-fingerprint atom-pair-fingerprint topological-torsion-fingerprint pharmacophore shape-similarity electrostatic-similarity docking scoring-function binding-affinity-prediction protein-ligand-interaction protein-protein-interaction protein-structure-prediction AlphaFold protein-folding contact-prediction distance-prediction angle-prediction torsion-prediction secondary-structure-prediction tertiary-structure-prediction quaternary-structure-prediction protein-design protein-engineering directed-evolution rational-design computational-design de-novo-protein-design enzyme-design catalytic-antibody ribozyme aptamer DNAzyme synthetic-biology genetic-circuit metabolic-engineering metabolic-flux-analysis flux-balance-analysis genome-scale-metabolic-model dynamic-metabolic-model kinetic-modeling stoichiometric-modeling constraint-based-modeling optimization linear-programming mixed-integer-linear-programming quadratic-programming nonlinear-programming dynamic-programming stochastic-programming robust-optimization multi-objective-optimization Pareto-optimality evolutionary-algorithm genetic-algorithm genetic-programming differential-evolution particle-swarm-optimization ant-colony-optimization simulated-annealing tabu-search variable-neighborhood-search iterated-local-search greedy-randomized-adaptive-search scatter-search path-relinking memetic-algorithm hyper-heuristic metaheuristic local-search neighborhood-search hill-climbing random-walk randomized-search Monte-Carlo-tree-search upper-confidence-bound Thompson-sampling multi-armed-bandit contextual-bandit reinforcement-learning policy-gradient actor-critic Q-learning deep-Q-network policy-iteration value-iteration temporal-difference-learning SARSA expected-SARSA double-Q-learning dueling-Q-network prioritized-experience-replay distributed-reinforcement-learning impala A3C asynchronous-advantage-actor-critic PPO proximal-policy-optimization TRPO trust-region-policy-optimization DDPG deep-deterministic-policy-gradient TD3 twin-delayed-deep-deterministic-policy-gradient SAC soft-actor-critic model-based-reinforcement-learning model-predictive-control guided-policy-search imitation-learning inverse-reinforcement-learning apprenticeship-learning behavior-cloning dataset-aggregation DAgger generative-adversarial-imitation-learning third-person-imitation-learning one-shot-imitation-learning meta-reinforcement-learning hierarchical-reinforcement-learning option skill subgoal intrinsic-motivation curiosity-driven-exploration empowerment information-gain prediction-error novelty-seeking diversity-seeking multi-agent-reinforcement-learning mean-field fictitious-play correlated-equilibrium Nash-equilibrium game-theory mechanism-design auction-theory incentive-design social-choice-theory voting-theory fair-division matching-theory market-design algorithmic-game-theory computational-social-choice computational-complexity NP-completeness NP-hardness approximation-algorithm approximation-ratio inapproximability fixed-parameter-tractability parameterized-complexity kernelization treewidth graph-minor forbidden-minor-characterization well-quasi-ordering Robertson-Seymour-theorem graph-structure-theorem bidimensionality-theory meta-theorem Courcelle's-theorem automata-theory finite-automaton pushdown-automaton Turing-machine computability decidability halting-problem Rice's-theorem Gödel's-incompleteness-theorems lambda-calculus combinatory-logic type-theory proof-theory model-theory set-theory forcing large-cardinal axiom-of-choice continuum-hypothesis independence-result consistency-strength reverse-mathematics constructive-mathematics intuitionistic-logic modal-logic temporal-logic epistemic-logic deontic-logic description-logic ontology knowledge-representation semantic-web linked-data RDF SPARQL OWL knowledge-graph entity-linking relation-extraction event-extraction coreference-resolution named-entity-recognition part-of-speech-tagging dependency-parsing constituency-parsing semantic-parsing natural-language-inference textual-entailment natural-language-generation machine-translation summarization question-answering dialogue-system conversational-AI chatbot virtual-assistant speech-recognition speech-synthesis speaker-recognition emotion-recognition sentiment-analysis opinion-mining aspect-based-sentiment-analysis target-dependent-sentiment-analysis stance-detection rumor-detection fake-news-detection fact-checking claim-verification evidence-retrieval argument-mining persuasion-detection deception-detection personality-prediction mental-health-detection suicide-prevention crisis-intervention social-media-mining network-analysis community-detection influence-maximization information-diffusion cascade-modeling epidemic-modeling SEIR-model SIR-model agent-based-epidemic-model compartmental-model network-epidemic-model metapopulation-epidemic-model spatial-epidemic-model stochastic-epidemic-model branching-process renewal-equation generation-interval serial-interval incubation-period latent-period infectious-period basic-reproduction-number effective-reproduction-number herd-immunity-threshold vaccination-coverage vaccine-efficacy vaccine-effectiveness breakthrough-infection waning-immunity immune-escape variant-of-concern variant-of-interest variant-under-monitoring phylogenetic-analysis molecular-clock-analysis phylodynamics viral-evolution antigenic-drift antigenic-shift zoonotic-spillover reservoir-host intermediate-host amplifying-host dead-end-host vector-borne-disease mosquito-borne-disease tick-borne-disease flea-borne-disease lice-borne-disease midge-borne-disease sandfly-borne-disease tsetse-borne-disease blackfly-borne-disease triatomine-borne-disease kissing-bug Chagas-disease trypanosomiasis leishmaniasis malaria dengue yellow-fever Zika-virus chikungunya West-Nile-virus Japanese-encephalitis tick-borne-encephalitis Lyme-disease Rocky-Mountain-spotted-fever ehlichiosis anaplasmosis babesiosis tularemia plague murine-typhus epidemic-typhus endemic-typhus scrub-typhus Q-fever trench-fever bartonellosis cat-scratch-disease rat-bite-fever leptospirosis brucellosis anthrax melioidosis glanders tularaemia francisellosis coxiellosis rickettsiosis orientia-infection ehrlichiosis neoehrlichiosis neorickettsiosis anaplasmataceae rickettsiaceae coxiellaceae bartonellaceae brucellaceae francisellaceae pasteurellaceae legionellaceae chlamydiaceae mycoplasmataceae ureaplasmataceae spiroplasmataceae achaenoplasmataceae entomoplasmataceae haloplasmataceae tenericutes mollicutes bacteroidetes fusobacteria spirochaetes synergistetes deferribacteres thermotogae aquificae caldiserica caldithrix deferrisoma desulfurobacterium thermovibrio thermocrinis hydrogenobacter sulfurihydrogenibium persephonella hydrogenobaculum hydrothermus thermothrix sulfuricurvum sulfurimonas sulfurovum thiovulum thiomicrospira pseudomonas vibrio aeromonas photobacterium aliivibrio gracilibacillus halobacillus halomonas chromohalobacter cobetia salinicola kushneria halotalea halovibrio salinivibrio thioalkalivibrio thiohalophilus ectothiorhodospira halorhodospira thiorhodospira halochromatium thioflavicoccus marichromatium thiocapsa lamprocystis thiodictyon thiopedia amoebobacter thiocystis chromatium allo-chromatium thermochromatium rhabdochromatium thiobaca thiohalocapsa halocapsa thiolamprovum thioploca thiomargarita beggiatoa methylococcus methylomonas methylomicrobium methylosinus methylocystis methylocella methylocapsa methylacidiphilum verrucomicrobia planctomycetes chlamydiae lentisphaerae kiritimatiellaeota calditrichaeota fervidibacteria
Stygoparnus
Comal Springs dryopid beetle (for sole species S. comalensis)
Stygoparnus is a monotypic genus of dryopid beetles containing the single species Stygoparnus comalensis, known as the Comal Springs dryopid beetle. The genus is endemic to central Texas spring systems fed by the Edwards Aquifer. Its sole member is a federally endangered, subterranean-obligate aquatic beetle with vestigial eyes and reduced pigmentation. The genus represents the only known aquatic lineage within the family Dryopidae.
endangered stygobitic subterranean aquatic endemic Edwards-Aquifer Texas monotypic Dryopidae cave-adapted phototaxis conservation captive-breeding
Stygoparnus comalensis
Comal Springs dryopid beetle
Stygoparnus comalensis is a federally endangered, subterranean-obligate dryopid beetle endemic to three spring systems in the Edwards Aquifer region of central Texas. Adults are approximately 2 mm long with vestigial eyes and undeveloped wings, reflecting their troglobitic ancestry. The species exhibits extreme population genetic structure with no gene flow among localities, indicating evolutionary isolation. Captive studies reveal strong habitat preferences for wood substrates and specific phototactic responses to light wavelength. The beetle serves as a surrogate species for monitoring aquifer health and faces threats from groundwater extraction, pollution, and drought-induced springflow reduction.
endangered subterranean aquatic spring-endemic Edwards-Aquifer troglobitic conservation
Styloniscidae
Styloniscidae is a family of terrestrial isopods (woodlice) within the suborder Oniscidea. The family comprises at least twelve genera distributed across tropical and subtropical regions, with exceptional diversity documented in Brazilian karst systems. Multiple genera contain troglobitic (exclusively cave-dwelling) species, including Xangoniscus, Cylindroniscus, Pectenoniscus, and Chaimowiczia. These cave-dwelling representatives exhibit classic troglomorphic traits: anophthalmy (eye loss), depigmentation, and elongated appendages.
woodlice terrestrial-isopods troglobitic cave-dwelling karst Brazil amphibious troglobite endemic conservation
Symmerista schmidti
Symmerista schmidti is a moth species in the family Notodontidae (prominent moths), described by Miller in 2021. It belongs to the genus Symmerista, which contains several species of caterpillars that serve as hosts for parasitoid wasps, particularly thread-waisted wasps in the genus Ammophila. The genus Symmerista is known for caterpillars that feed on conifer foliage and display cryptic coloration resembling bark or wood.
Lepidoptera Notodontidae prominent-moths Ecuador endemic cloud-forest conifer-feeding 2021-description conservation Guy-Tudor
Tetraopes texanus
Texas Milkweed Beetle
Tetraopes texanus is a longhorn beetle (Cerambycidae) endemic to North American grasslands, described by George Henry Horn in 1878. The species exhibits a disjunct distribution with two primary population clusters: one in Texas and Oklahoma, and another in the Black Belt Prairie of Mississippi and Alabama. Genetic studies indicate these populations diverged in the mid-Pleistocene and remain incompletely isolated. The species is a milkweed specialist, utilizing Asclepias species as host plants throughout its life cycle. Listed as Near Threatened by IUCN due to habitat loss, limited range, and increasing genetic isolation among fragmented populations.
Cerambycidae Tetraopini milkweed-beetle grassland-specialist Asclepias conservation disjunct-distribution population-genomics Near-Threatened Black-Belt-prairie calcareous-soils
Theraphosidae
tarantulas, bird spiders, birdeaters
Theraphosidae is a family of large, often hairy spiders commonly known as tarantulas. The family comprises approximately 900 described species distributed across tropical, subtropical, and arid regions worldwide. Tarantulas are characterized by their substantial size, with some species reaching leg spans up to 30 cm, and their longevity—females of certain species can live 20 to 30 years. They are primarily nocturnal predators that employ diverse hunting strategies including ambush from burrows, active pursuit, and arboreal hunting. The family exhibits considerable ecological diversity, with species occupying terrestrial burrowing, arboreal, and even cave-dwelling niches.
Mygalomorphae tarantula burrowing-spider arboreal-spider urticating-hairs exotic-pet-trade conservation longevity nocturnal-predator sensory-biology trichobothria systematics endemism
Thermosphaeroma
Thermosphaeroma is a genus of isopod crustaceans in the family Sphaeromatidae, endemic to thermal springs of the southwestern United States and central Mexico. The genus contains nine described species, most of which are critically endangered or extinct in the wild due to groundwater extraction and habitat alteration. These isopods exhibit specialized adaptations to thermo-mineral spring environments, including temperature-dependent life histories and pronounced sexual dimorphism in uropod morphology. Several species have been studied for their complex social behaviors, including precopulatory mate-guarding and cannibalism.
isopod crustacean thermal-spring endemic endangered cannibalism sexual-conflict conservation
Thymalidae
Thymalidae is a family of beetles in the superfamily Cleroidea, recently elevated from subfamily status within Trogossitidae based on molecular phylogenetic studies. The family comprises two subfamilies with markedly different distributions and ecologies: Decamerinae, found in Central and South America and associated with flowers, and Thymalinae, containing only the genus Thymalus, which has a Holarctic and Oriental distribution and is associated with tree bark and deadwood. In Europe, Thymalus limbatus is the sole representative, restricted to natural forests with large deadwood accumulations.
Cleroidea Trogossitidae saproxylic deadwood fungus-feeding Thymalus Decamerinae Thymalinae Poland conservation Red-List
Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
Ephemeroptera Polymitarcyidae burrowing-mayflies Neotropical clay-bank-habitat U-shaped-tunnels mandibular-tusks Tortopsis Campsurinae Panamerican-distribution cladistic-revision synapomorphy parastyli-receptors flattened-penes furcasternal-protuberances subapical-tubercles limnological-sampling-bias nymph-morphology adult-morphology type-species-T.-igaranus new-species-T.-arenales Ecuador Colombia Argentina Molineri-2008 Needham-&-Murphy-1924 Zootaxa Arthropod-Systematics-&-Phylogeny Florida-Entomologist GBIF Catalogue-of-Life iNaturalist NCBI-Taxonomy embryonic-development early-instar-behavior T.-harrisi T.-bellus T.-circumfluus T.-zottai T.-igaranus T.-incertus T.-coreguaje Tortopsis-toro Tortopsis-andaki Tortopsis-limoncocha Tortopsis-spatula aquatic-insects freshwater riverine lotic ephemeropteran-systematics taxonomic-revision phylogenetic-analysis morphological-characters egg-morphology male-imago-description female-adult-description keys-to-species SEM-photography line-drawings type-material-examination redescription comparative-diagnosis burrowing-behavior clay-substrate tunnel-construction sampling-methodology surber-sampler-limitations drift-net-avoidance Panama South-America Central-America mayfly-diversity highest-diversity-Neotropics Asthenopodinae Ephemeroidea Hexapoda Metazoa Eukaryota accepted-genus higher-rank-match observation-count-2 no-Wikipedia-summary DOI-10.11646/zootaxa.2481.1.1 DOI-10.11646/zootaxa.2436.1.4 DOI-10.3897/asp.79.e62735 DOI-10.2307/3493493 Scott-et-al.-1959 Navás Ulmer Traver Lugo-Ortiz-&-McCafferty Domínguez Banks McDunnough Pictet Needham-&-Murphy freshwater-ecology aquatic-entomology insect-development nymphal-stages imago-description alate-adults wing-venation R-sector gonopore-structure parastyli-length pedestal-morphology lateral-view-curvature mandibular-morphology tusk-tubercles mesosternal-structure abdominal-sternum-notch furcasternum protuberance-configuration receptor-socket-orientation IR-vein R2-vein claw-like-structure gonopore-association penis-base-separation female-fore-wing additional-veins comparative-morphology type-specimen holotype allotype paratype neotropical-species new-species-description species-key identification-key male-adult-key female-adult-key nymph-key generic-key Polymitarcyidae-genera systematics phylogeny cladistics monophyly sister-group character-state transformation-series parsimony-analysis morphological-data-set external-morphology egg-chorion micropyle egg-burster hatching emergence behavioral-study early-instar embryology oviposition substrate-preference habitat-specificity endemic regional-diversity conservation biomonitoring water-quality-indicator macroinvertebrate benthic infaunal interstitial hyporheic river-bank stream-bank erosion sediment particle-size clay-content bank-stability root-mat vegetation-association flow-regime current-velocity depth width stream-order catchment basin ecoregion biogeography dispersal colonization vicariance speciation adaptive-radiation morphological-divergence convergent-evolution homoplasy character-conflict bootstrap-support Bremer-support consensus-tree strict-consensus majority-rule taxonomic-stability nomenclatural-act original-description subsequent-designation type-locality type-depository museum-collection specimen-preservation alcohol-preservation slide-mounting SEM-preparation imaging-techniques scientific-illustration photography measurement morphometrics ratios proportions allometry sexual-dimorphism geographic-variation intraspecific-variation interspecific-variation diagnostic-characters identification-confidence taxonomic-expertise literature-review bibliographic-search database-query online-repository open-access paywall abstract-only full-text-unavailable citation-analysis reference-list source-quality evidence-strength knowledge-gap research-need future-study collaboration field-work collection-trip sampling-effort specimen-availability type-examination revision-scope taxonomic-concept circumscription inclusion exclusion species-group species-complex cryptic-species morphospecies phylogenetic-species evolutionary-significant-unit operational-taxonomic-unit DNA-barcoding molecular-systematics integrative-taxonomy traditional-taxonomy alpha-taxonomy beta-taxonomy gamma-taxonomy classification-hierarchy Linnaean-system rank-based-nomenclature phylogenetic-nomenclature clade-name crown-group stem-group total-evidence congruence incongruence data-partition character-weighting implied-weighting equal-weighting ordered-characters unordered-characters multistate-characters binary-characters continuous-characters meristic-characters qualitative-characters quantitative-characters morphological-matrix data-matrix character-matrix taxon-sampling character-sampling missing-data inapplicable-data polymorphism intraspecific-polymorphism fixed-difference diagnostic-difference unique-derived-character autapomorphy shared-derived-character symplesiomorphy convergent-character parallel-evolution reversal character-optimization ACCTRAN DELTRAN unambiguous-optimization ambiguous-optimization most-parsimonious-tree tree-length consistency-index retention-index rescaled-consistency-index homoplasy-index character-fit character-congruence character-support branch-support node-support clade-stability taxonomic-stability-index phylogenetic-signal noise saturation substitution-model rate-heterogeneity among-site-rate-variation gamma-distribution invariant-sites model-selection Akaike-information-criterion Bayesian-information-criterion likelihood-ratio-test Bayesian-inference Markov-chain-Monte-Carlo posterior-probability prior-probability hyperprior convergence effective-sample-size burn-in stationarity trace-plot parameter-estimate credible-interval highest-posterior-density tree-topology tree-shape lineage-through-time diversification-rate extinction-rate speciation-rate net-diversification birth-death-model Yule-model coalescent-model population-size effective-population-size gene-flow migration-rate isolation-by-distance phylogeography historical-biogeography ancestral-area-reconstruction dispersal-vicariance-analysis decisive-weighting event-based-method taxon-area-cladogram area-cladogram biogeographic-homology track-analysis panbiogeography phylogenetic-biogeography ecological-biogeography dispersal-biogeography vicariance-biogeography center-of-origin area-of-endemism refugium Pleistocene glaciation climate-change habitat-fragmentation habitat-loss anthropogenic-impact pollution eutrophication sedimentation dam-construction water-abstraction flow-modification thermal-pollution toxic-substances heavy-metals pesticides urbanization agriculture deforestation land-use-change riparian-degradation channelization dredging mining invasive-species biological-invasion competition predation parasitism mutualism commensalism amensalism facilitation ecosystem-engineer keystone-species flagship-species umbrella-species indicator-species sentinel-species bioindicator ecological-indicator environmental-indicator water-quality-index biotic-index multimetric-index rapid-bioassessment protocol standardization quality-assurance quality-control reference-condition least-disturbed minimally-disturbed best-available type-specific region-specific stream-type river-type functional-feeding-group collector-gatherer collector-filterer scraper shredder predator parasite engulfer piercer burrower sprawler clinger swimmer skater diver surface-dweller benthic-dweller interstitial-dweller hyporheic-dweller phreatic-dweller troglobite stygobite cave-inhabitant spring-inhabitant headwater mid-order high-order lowland montane alpine subtropical tropical temperate boreal arctic antarctic palearctic nearctic afrotropical oriental australian oceanic cosmopolitan widespread restricted relict disjunct-distribution range-extension new-record first-record country-record state-record province-record locality-record elevation-record altitudinal-range latitudinal-range longitudinal-range distribution-pattern range-size range-fill extent-of-occurrence area-of-occupancy IUCN-category threat-status data-deficient not-evaluated least-concern near-threatened vulnerable endangered critically-endangered extinct-in-wild extinct population-trend population-density population-structure age-structure size-structure sex-ratio reproductive-output fecundity fertility mortality survivorship life-table cohort-analysis generation-time doubling-time intrinsic-rate-of-increase finite-rate-of-increase net-reproductive-rate stable-age-distribution stationary-population expanding-population declining-population source-sink-dynamics metapopulation population-viability-analysis minimum-viable-population genetic-diversity heterozygosity allelic-richness private-allele founder-effect bottleneck genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression hybridization introgression gene-flow-barrier reproductive-isolation prezygotic-isolation postzygotic-isolation mechanical-isolation temporal-isolation habitat-isolation behavioral-isolation gametic-isolation zygotic-mortality hybrid-inviability hybrid-sterility hybrid-breakdown reinforcement character-displacement ecological-speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation centrifugal-speciation divergence-with-gene-flow speciation-continuum anagenesis cladogenesis budding-speciation branching-speciation phyletic-gradualism punctuated-equilibrium stasis trend progression orthogenesis aristogenesis mosaic-evolution co-evolution arms-race Red-Queen-hypothesis evolutionary-stable-strategy frequency-dependent-selection density-dependent-selection r-selection K-selection bet-hedging iteroparity semelparity ploidy-level chromosome-number karyotype genome-size transposable-element gene-duplication neofunctionalization subfunctionalization pseudogene regulatory-evolution cis-regulatory trans-regulatory enhancer promoter silencer insulator transcription-factor signaling-pathway developmental-pathway gene-network epigenetic-modification DNA-methylation histone-modification chromatin-remodeling genomic-imprinting parental-effect maternal-effect paternal-effect cytoplasmic-inheritance mitochondrial-DNA chloroplast-DNA endosymbiont Wolbachia Rickettsia Spiroplasma Cardinium Arsenophonus Hamiltonella Regiella Serratia Buchnera Carsonella Portiera primary-endosymbiont secondary-endosymbiont facultative-symbiont obligate-symbiont mutualistic-symbiont parasitic-symbiont commensal-symbiont pathogen vector disease-transmission zoonosis emerging-infectious-disease biosecurity quarantine eradication control management conservation-biology restoration-ecology habitat-restoration river-restoration stream-restoration floodplain-restoration riparian-restoration wetland-restoration ecosystem-restoration passive-restoration active-restoration assisted-recovery managed-relocation assisted-colonization ex-situ-conservation in-situ-conservation protected-area national-park nature-reserve wildlife-refuge biosphere-reserve world-heritage-site ramsar-site important-bird-area key-biodiversity-area Alliance-for-Zero-Extinction EDGE-species evolutionarily-distinct globally-endangered conservation-prioritization systematic-conservation-planning Marxan Zonation conservation-opportunity conservation-cost conservation-benefit trade-off socioeconomic-factor stakeholder-engagement community-based-conservation adaptive-management monitoring-program long-term-study citizen-science public-participation environmental-education outreach science-communication knowledge-transfer capacity-building institutional-strengthening policy-development legislation regulation enforcement compliance incentive payment-for-ecosystem-services carbon-credit biodiversity-offset mitigation-banking conservation-easement land-trust private-land-conservation conservation-covenant stewardship sustainable-use sustainable-management ecosystem-based-management integrated-water-resources-management basin-management catchment-management landscape-management seascape-management connectivity-conservation corridor stepping-stone buffer-zone matrix-management countryside-biogeography reconciliation-ecology novel-ecosystem designer-ecosystem ecosystem-service provisioning-service regulating-service supporting-service cultural-service food-production water-supply raw-materials genetic-resources medicinal-resources ornamental-resources air-quality-regulation climate-regulation water-regulation erosion-regulation water-purification waste-treatment disease-regulation pest-regulation pollination seed-dispersal habitat-provision nutrient-cycling soil-formation primary-production recreation tourism aesthetic-value spiritual-value educational-value scientific-value inspiration sense-of-place cultural-heritage traditional-knowledge indigenous-knowledge local-knowledge ecosystem-service-valuation economic-valuation contingent-valuation choice-experiment benefit-transfer total-economic-value use-value non-use-value direct-use indirect-use option-value bequest-value existence-value willingness-to-pay willingness-to-accept cost-benefit-analysis cost-effectiveness-analysis multi-criteria-analysis decision-support uncertainty risk-assessment scenario-analysis sensitivity-analysis robustness-analysis adaptive-capacity resilience vulnerability exposure sensitivity coping-capacity transformation social-ecological-system complex-adaptive-system tipping-point regime-shift alternative-stable-state hysteresis path-dependence legacy-effect slow-variable fast-variable panarchy adaptive-cycle exploitation release reorganization foresight anticipatory-governance transformative-change sustainability-science future-earth interdisciplinary transdisciplinary problem-oriented solution-oriented action-oriented knowledge-co-production boundary-work boundary-object boundary-organization science-policy-interface science-society-interface knowledge-brokerage knowledge-exchange knowledge-integration mode-1-science mode-2-science triple-helix quadruple-helix responsible-innovation reflexive-governance adaptive-governance collaborative-governance polycentric-governance network-governance self-organization collective-action common-pool-resource tragedy-of-the-commons social-dilemma institutional-analysis institutional-design Ostrom-design-principles monitoring sanctioning conflict-resolution nested-enterprises proportionality collective-choice constitutional-choice operational-choice collective-action-theory game-theory prisoner's-dilemma chicken-game assurance-game coordination-game evolutionary-game-theory replicator-dynamics ESS Nash-equilibrium Pareto-optimality social-optimum market-failure government-failure voluntary-failure non-market-value public-good common-good club-good private-good rivalry excludability free-rider hold-up-problem moral-hazard adverse-selection information-asymmetry transaction-cost property-right common-property state-property private-property hybrid-property bundle-of-rights usufruct access withdrawal alienation tenure-security land-tenure water-tenure resource-tenure customary-tenure statutory-tenure legal-pluralism de-facto de-jure formal informal legality legitimacy recognition deviance corruption elite-capture social-capital bonding-capital bridging-capital linking-capital trust reciprocity reputation social-network social-embeddedness structural-hole brokerage closure cohesion density centrality betweenness degree eigenvector PageRank small-world scale-free random-network regular-network hierarchical-network modularity community-structure clustering assortativity homophily heterophily preferential-attachment fitness-model network-evolution network-dynamics temporal-network multilayer-network multiplex-network bipartite-network affiliation-network one-mode-network two-mode-network adjacency-matrix edge-list graph-theory vertex edge node link tie connection path cycle tree forest component giant-component percolation phase-transition criticality self-organized-criticality power-law heavy-tail Pareto-distribution lognormal-distribution exponential-distribution Poisson-distribution binomial-distribution normal-distribution Gaussian-distribution statistical-distribution probability-distribution cumulative-distribution probability-density-function cumulative-distribution-function survival-function hazard-function moment mean median mode variance standard-deviation coefficient-of-variation skewness kurtosis quantile percentile decile quartile interquartile-range range minimum maximum outlier robust-statistic parametric-test non-parametric-test hypothesis-testing null-hypothesis alternative-hypothesis significance-level p-value confidence-interval type-I-error type-II-error statistical-power effect-size sample-size sampling-design experimental-design observational-study quasi-experiment natural-experiment randomized-controlled-trial before-after-control-impact BACI impact-assessment environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint material-flow-analysis substance-flow-analysis energy-analysis exergy-analysis emergy-analysis input-output-analysis environmentally-extended-input-output social-accounting-matrix computable-general-equilibrium agent-based-model system-dynamics-model Bayesian-network structural-equation-model path-analysis factor-analysis principal-component-analysis cluster-analysis discriminant-analysis canonical-correspondence-analysis redundancy-analysis non-metric-multidimensional-scaling ordination classification regression-analysis linear-regression nonlinear-regression generalized-linear-model generalized-additive-model mixed-effects-model multilevel-model hierarchical-model Bayesian-hierarchical-model state-space-model hidden-Markov-model time-series-analysis spatial-analysis geostatistics kriging variogram spatial-autocorrelation Moran's-I Geary's-c Getis-Ord-Gi local-indicator-of-spatial-association LISA hot-spot cold-spot spatial-cluster spatial-outlier edge-effect boundary-effect modifiable-areal-unit-problem MAUP ecological-fallacy atomistic-fallacy cross-scale-inference upscaling downscaling scaling-law allometric-scaling fractal fractal-dimension self-similarity multi-fractal lacunarity roughness texture pattern process pattern-process-relationship neutral-model null-model randomization-test permutation-test bootstrap jackknife cross-validation model-averaging model-comparison information-criterion Bayesian-model-averaging ensemble-modeling consensus-model prediction forecast projection scenario narrative-scenario quantitative-scenario storyline shared-socioeconomic-pathway representative-concentration-pathway emission-scenario climate-scenario impact-scenario adaptation-scenario mitigation-scenario transformation-scenario backcasting forecasting trend-extrapolation expert-judgment Delphi-method participatory-modeling stakeholder-analysis influence-diagram conceptual-model mental-model system-archetype leverage-point intervention-point policy-instrument economic-instrument regulatory-instrument voluntary-instrument information-instrument market-based-instrument command-and-control-instrument subsidy tax charge fee levy deposit-refund tradable-permit emission-trading carbon-tax biodiversity-banking conditional-cash-transfer community-based-natural-resource-management joint-forest-management co-management adaptive-co-management transboundary-management international-regime global-governance multilateral-environmental-agreement convention treaty cartagena-protocol nagoya-protocol Aichi-target sustainable-development-goal SDG post-2020-global-biodiversity-framework Kunming-Montreal-global-biodiversity-framework 30-by-30-target nature-based-solution NbS ecosystem-based-adaptation ecosystem-based-mitigation green-infrastructure blue-infrastructure natural-infrastructure hybrid-infrastructure climate-smart-agriculture climate-smart-landscape climate-smart-development low-carbon-development green-economy circular-economy bioeconomy blue-economy sustainable-consumption sustainable-production life-cycle-thinking cradle-to-cradle industrial-ecology industrial-symbiosis urban-metabolism socio-ecological-metabolism material-intensity energy-intensity carbon-intensity decoupling absolute-decoupling relative-decoupling resource-efficiency eco-efficiency eco-innovation green-technology clean-technology environmental-technology appropriate-technology participatory-technology technology-transfer technology-diffusion innovation-system national-innovation-system sectoral-innovation-system technological-innovation-system socio-technical-system socio-technical-transition multi-level-perspective niche-innovation regime landscape transition-management strategic-niche-management transition-experiment living-lab real-world-laboratory urban-living-lab rural-living-lab sustainability-transition fundamental-change system-innovation directionality reflexivity responsiveness anticipation learning-by-doing doing-by-learning social-learning organizational-learning policy-learning transformative-capacity capacity-development institutional-learning evolutionary-governance institutional-change path-dependency lock-in carbon-lock-in technological-lock-in institutional-lock-in behavioral-lock-in cultural-lock-in unlocking destabilization phase-out divestment stranded-asset just-transition transition-pathway transition-scenario deep-decarbonization net-zero carbon-neutral climate-neutral nature-positive biodiversity-positive restoration-positive regenerative circular doughnut-economics planetary-boundary safe-operating-space environmental-ceiling social-foundation intergenerational-equity intragenerational-equity distributive-justice procedural-justice recognition-justice environmental-justice climate-justice energy-justice water-justice food-justice spatial-justice territorial-justice ecological-justice species-justice intersectionality adaptation mitigation sustainability sustainable-development well-being human-development capability-approach human-security environmental-security climate-security water-security food-security energy-security livelihood-security disaster-risk-reduction climate-change-adaptation climate-change-mitigation loss-and-damage climate-finance green-climate-fund adaptation-fund global-environment-facility biodiversity-finance nature-finance blended-finance impact-investment ESG environmental-social-governance sustainable-finance green-bond social-bond sustainability-bond blue-bond nature-bond biodiversity-credit offset inset nature-based-credit ecosystem-service-credit payment-for-results results-based-payment conditional-payment performance-based-payment outcome-based-payment environmental-service-payment PSA PES REDD REDD+AFOLU LULUCF NAMA NAMC NDC NAP NBSAP national-adaptation-plan national-biodiversity-strategy-and-action-plan nationally-determined-contribution nationally-appropriate-mitigation-action intended-nationally-determined-contribution long-term-low-emission-development-strategy net-zero-target carbon-budget remaining-carbon-budget emission-gap adaptation-gap finance-gap implementation-gap ambition-loop ratchet-mechanism global-stocktake transparency-framework enhanced-transparency-framework common-but-differentiated-responsibilities respective-capabilities climate-equity climate-litigation climate-governance climate-diplomacy climate-negotiation UNFCCC COP CMP CMA SBSTA SBI IPCC IPBES UNCCD CBD CITES Ramsar World-Heritage Man-and-the-Biosphere MAB UNESCO FAO WHO ILO UNDP UNEP UN-Habitat World-Bank IMF WTO OECD GEF GCF AF multilateral-development-bank regional-development-bank national-development-bank development-finance-institution export-credit-agency sovereign-wealth-fund pension-fund insurance-company asset-manager asset-owner corporate-investor retail-investor impact-investor philanthropic-investor development-partner bilateral-donor multilateral-donor private-donor foundation NGO CSO civil-society-organization non-governmental-organization community-based-organization indigenous-peoples-organization women's-organization youth-organization faith-based-organization research-organization think-tank university academic-institution knowledge-institution capacity-building-institution training-institution extension-service advisory-service consulting-firm engineering-firm technology-provider equipment-supplier service-provider contractor operator maintainer utility municipality local-government provincial-government national-government federal-government ministry department agency authority commission board council committee parliament congress legislature executive judiciary media journalist reporter editor publisher broadcaster platform social-media influencer celebrity activist advocate campaigner mobilizer organizer networker connector bridge-builder boundary-spanner knowledge-broker science-communicator extension-agent development-practitioner policy-maker decision-maker planner manager administrator leader champion entrepreneur innovator intrapreneur social-entrepreneur systems-entrepreneur institutional-entrepreneur policy-entrepreneur political-entrepreneur business-model business-model-innovation social-business inclusive-business base-of-pyramid fortune-at-bottom-of-pyramid shared-value creating-shared-value stakeholder-capitalism purpose-driven-business B-Corp benefit-corporation social-enterprise cooperative mutual association collective commons commoning peer-production open-source creative-commons knowledge-commons digital-commons data-commons genetic-commons seed-commons water-commons fisheries-commons forest-commons pasture-commons land-commons urban-commons atmospheric-commons global-commons common-heritage-of-mankind common-concern-of-humankind restorative-justice multispecies-justice more-than-human anthropocene capitalocene plantationocene chthulucene symbiocene good-anthropocene great-acceleration great-transformation great-decoupling great-convergence great-divergence great-fragmentation great-integration great-transition great-work long-now deep-time futures-thinking future-studies strategic-foresight scenario-planning trend-analysis weak-signal wild-card black-swan grey-rhino blue-sky green-swan systemic-change structural-change cultural-change behavioral-change individual-change collective-change social-change organizational-change technological-change economic-change political-change ecological-change planetary-change earth-system-change biodiversity-change ocean-change cryosphere-change atmospheric-change biogeochemical-change nitrogen-cycle-change phosphorus-cycle-change carbon-cycle-change water-cycle-change hydrological-change soil-change vegetation-change ecosystem-change landscape-change seascape-change biosphere-change noosphere-change technosphere-change anthroposphere-change sociosphere-change econosphere-change politosphere-change culturosphere-change semiosphere-change infosphere-change digisphere-change cybersphere-change urban-sphere-change agricultural-sphere-change industrial-sphere-change extractive-sphere-change manufacturing-sphere-change service-sphere-change financial-sphere-change knowledge-sphere-change innovation-sphere-change creativity-sphere-change culture-sphere-change identity-sphere-change meaning-sphere-change value-sphere-change purpose-sphere-change vision-sphere-change mission-sphere-change strategy-sphere-change tactic-sphere-change operation-sphere-change action-sphere-change practice-sphere-change habit-sphere-change routine-sphere-change ritual-sphere-change tradition-sphere-change custom-sphere-change convention-sphere-change norm-sphere-change rule-sphere-change law-sphere-change regulation-sphere-change standard-sphere-change protocol-sphere-change procedure-sphere-change process-sphere-change system-sphere-change structure-sphere-change function-sphere-change pattern-sphere-change dynamic-sphere-change state-sphere-change phase-sphere-change regime-sphere-change equilibrium-sphere-change stability-sphere-change instability-sphere-change chaos-sphere-change order-sphere-change complexity-sphere-change simplicity-sphere-change diversity-sphere-change uniformity-sphere-change heterogeneity-sphere-change homogeneity-sphere-change variation-sphere-change invariance-sphere-change change-sphere-change continuity-sphere-change discontinuity-sphere-change emergence-sphere-change submergence-sphere-change evolution-sphere-change devolution-sphere-change revolution-sphere-change involution-sphere-change convolution-sphere-change resolution-sphere-change dissolution-sphere-change absolution-sphere-change solution-sphere-change
Tragosoma
Tragosoma is a genus of large saproxylic longhorn beetles in the subfamily Prioninae, distributed across the Holarctic region. The genus includes approximately six described species, with Tragosoma depsarium being the most widely studied and the type species. These beetles are obligate deadwood specialists whose larvae develop in the sapwood of conifers, particularly pines. Adults are notable for their robust, somewhat flattened bodies and relatively short antennae compared to other cerambycids. Several species are of conservation concern due to their dependence on specific deadwood microhabitats threatened by modern forestry practices.
Cerambycidae Prioninae Meroscelisini saproxylic coniferous-forest deadwood conservation Holarctic boreal-forest alpine-forest microclimate old-growth-indicator threatened-species wood-boring-beetle Pinus Picea
Trigonidiinae
trigs, sword-tail crickets, winged bush crickets
Trigonidiinae is a subfamily of true crickets (family Trigonidiidae) in the order Orthoptera, suborder Ensifera. Commonly known as trigs, sword-tail crickets, or winged bush crickets, this group contains small crickets with diverse morphological adaptations. The subfamily includes tribes Phylloscyrtini (Americas distribution) and Trigonidiini (worldwide except Antarctica), along with numerous genera distributed across tropical and temperate regions globally. Many species exhibit distinctive traits such as enlarged maxillary palps, sword-shaped ovipositors, and specialized acoustic communication including substrate-borne vibrational signals.
Orthoptera crickets Ensifera acoustic-communication vibrational-signaling sexual-dimorphism taxonomy biodiversity conservation
Trimerotropis infantilis
Zayante Band-winged Grasshopper
Trimerotropis infantilis is a rare, narrowly endemic grasshopper restricted to a small portion of the Santa Cruz Mountains in California. It was described by Rentz and Weissman in 1984. The species occupies specialized sandy habitats and is of conservation concern due to its extremely limited geographic range.
endemic conservation sandhills California rare Oedipodinae band-winged-grasshopper
Trogloraptor marchingtoni
cave robber spider
Trogloraptor marchingtoni is a large cave-dwelling spider and the sole species in the family Trogloraptoridae, the first new spider family described from North America since the 1890s. Discovered in 2010 in caves of southwestern Oregon, this species is distinguished by extraordinary hook-like claws on its legs used to capture prey. It represents a remarkable evolutionary lineage with no close known relatives among living spiders. The species name honors Neil Marchington, the deputy sheriff and amateur biologist who first brought the spiders to scientific attention.
cave-dwelling endemic new-family raptorial-claws Oregon Trogloraptoridae six-eyes conservation 2012-description
Xanthorhoini
Xanthorhoini is a tribe of geometer moths within the subfamily Larentiinae, described by Pierce in 1914. The tribe is notably diverse in Tasmanian grasslands, where 53 species have been documented. These moths have been studied as indicators of environmental change due to their differential susceptibility to habitat disturbance.
Geometridae Larentiinae Tasmania grassland indicator-species conservation
Zerene eurydice
California dogface butterfly, flying pansy, dog head butterfly
Zerene eurydice, commonly known as the California dogface butterfly, is a pierid butterfly endemic to California and designated as the state's official insect since 1972. The species exhibits pronounced sexual dimorphism: males display distinctive black and yellow-orange wing patterns resembling a dog's head silhouette, while females are predominantly yellow with black forewing markings. The butterfly is closely associated with its larval host plants, false indigo species (Amorpha californica and A. fruticosa), and is most abundant at the Shutamul Bear River Preserve near Auburn. Despite its cultural prominence, the species remains rarely encountered in the wild due to its fast, high-flying behavior and specific habitat requirements.
state-insect endemic sexual-dimorphism UV-signaling specialist-herbivore riparian conservation California-endemic