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Sticholotidinae
Sticholotidinae is a subfamily of lady beetles (family Coccinellidae). Phylogenetic analyses have found that Sticholotidinae is not a monophyletic group, rendering the subfamily invalid as currently circumscribed. The group has been the subject of recent taxonomic revisions, particularly for genera such as Stictobura and Sticholotis. Members of this subfamily are generally small, often flattened beetles with varied habits.
Coccinellidae lady-beetles invalid-subfamily non-monophyletic taxonomic-revision
Stictocephala stimulea
Stictocephala stimulea is a treehopper species in the family Membracidae, first described by Van Duzee in 1914. Treehoppers in this genus are characterized by their enlarged, often ornate pronotum that extends over the body like a helmet or thorn. The specific epithet 'stimulea' suggests possible resemblance to or association with stinging organisms, though this refers to appearance rather than actual sting. Like other membracids, this species likely feeds on plant phloem sap using piercing-sucking mouthparts.
Membracidae treehopper Auchenorrhyncha Cicadomorpha Stictocephala Ceresini Smiliinae Hemiptera sap-feeder pronotum helmet thorn-mimicry plant-feeding-insect phloem-feeder cryptic-coloration camouflage Van-Duzee-1914 Nearctic North-America Florida North-Carolina South-Dakota understudied-species data-deficient taxonomy entomology insect arthropod true-bug auchenorrhynchan cicadomorph leafhopper-relative plant-hopper ornate-pronotum pronotal-extension dorsal-helmet insect-morphology insect-systematics biodiversity natural-history conservation insect-ecology insect-behavior insect-development insect-distribution biogeography species-description type-specimen original-description historical-taxonomy museum-specimen collection-record occurrence-data GBIF iNaturalist citizen-science observation-record range-map distribution-data geographic-range endemic native-species regional-fauna local-biodiversity habitat-specificity host-plant feeding-guild phytophagous herbivore plant-insect-interaction coevolution mimicry defense crypsis evolutionary-adaptation morphological-adaptation exoskeleton-modification structural-coloration insect-identification field-guide diagnostic-character taxonomic-key species-complex cryptic-species morphological-variation geographic-variation phenotypic-plasticity life-history reproduction oviposition nymph instar metamorphosis hemimetabolous incomplete-metamorphosis population-dynamics community-ecology food-web trophic-level primary-consumer ecosystem-function nutrient-cycling energy-flow biological-control pest-status agricultural-relevance forestry urban-ecology invasive-potential climate-change habitat-loss conservation-status data-gap research-need taxonomic-revision phylogenetics molecular-systematics DNA-barcoding integrative-taxonomy natural-history-collection specimen-digitization open-data FAIR-data biodiversity-informatics biological-database taxonomic-authority nomenclature zoological-nomenclature International-Code-of-Zoological-Nomenclature type-locality type-specimen-deposition author-citation original-combination current-combination synonymy homonymy nomenclatural-act taxonomic-history classification-system phylogenetic-classification evolutionary-relationships monophyly clade lineage stem-group crown-group divergence-time molecular-clock fossil-record paleontology extant extinct living-fossil evolutionary-history biogeographic-history dispersal vicariance range-expansion range-contraction refugium glacial-relict postglacial-colonization anthropogenic-impact human-disturbance land-use-change urbanization agricultural-intensification pesticide-use habitat-fragmentation connectivity corridor protected-area conservation-planning systematic-conservation-planning priority-area hotspot endemic-area center-of-origin center-of-diversity evolutionary-significant-unit management-unit population metapopulation source-sink-dynamics demography vital-rates survival fecundity generation-time population-growth-rate carrying-capacity density-dependence regulation limiting-factor environmental-stochasticity demographic-stochasticity genetic-stochasticity inbreeding-depression outbreeding-depression genetic-diversity heterozygosity allelic-richness effective-population-size genetic-drift gene-flow migration isolation-by-distance isolation-by-resistance landscape-genetics population-genomics phylogeography historical-demography coalescent-theory approximate-Bayesian-computation species-distribution-model ecological-niche-model habitat-suitability bioclimatic-envelope climate-envelope niche-conservatism niche-evolution realized-niche fundamental-niche biotic-interaction competition predation parasitism mutualism commensalism amensalism facilitation apparent-competition indirect-interaction trophic-cascade top-down-control bottom-up-control mesopredator-release keystone-species ecosystem-engineer foundation-species dominant-species characteristic-species indicator-species umbrella-species flagship-species surrogate-species focal-species target-species pest-species invasive-species naturalized-species alien-species non-native-species introduced-species cryptogenic-species range-shift poleward-shift elevational-shift phenological-shift temporal-mismatch trophic-mismatch phenology seasonality voltinism univoltine bivoltine multivoltine partial-generation prolonged-diapause aestivation hibernation overwintering overwintering-strategy cold-hardiness freeze-tolerance freeze-avoidance supercooling cryoprotectant thermal-biology thermoregulation behavioral-thermoregulation microclimate-selection habitat-selection oviposition-site-selection host-selection mate-choice sexual-selection sperm-competition cryptic-female-choice reproductive-isolation prezygotic-isolation postzygotic-isolation speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation reinforcement hybridization introgression hybrid-zone cline ecotone contact-zone secondary-contact exogenous-selection endogenous-selection ecological-speciation mutation-order-speciation sexual-conflict sexual-dimorphism sexual-size-dimorphism sexual-dichromatism polymorphism color-polymorphism morph phase gregarious-phase solitary-phase density-dependent-polyphenism reaction-norm genotype-environment-interaction plasticity canalization genetic-assimilation acclimation acclimatization hardening stress-response heat-shock-protein oxidative-stress hypoxia desiccation starvation immune-response innate-immunity humoral-immunity cellular-immunity encapsulation nodulation phagocytosis melanization antimicrobial-peptide phenoloxidase prophenoloxidase lysozyme defensin cecropin diptericin attacin immune-gene immune-pathway Toll-pathway Imd-pathway JAK-STAT-pathway NF-κB relish dorsal immune-memory specific-immunity transgenerational-immune-priming gut-microbiome symbiosis obligate-symbiont facultative-symbiont primary-symbiont secondary-symbiont vertical-transmission horizontal-transmission mixed-mode-transmission nutritional-symbiosis defensive-symbiosis digestive-symbiosis detoxification metabolism biosynthesis vitamin-synthesis amino-acid-synthesis steroid-synthesis cuticle-synthesis chitin-synthesis molting ecdysis ecdysteroid juvenile-hormone insulin-signaling TOR-signaling nutrient-sensing energy-storage lipid-metabolism carbohydrate-metabolism protein-metabolism flight-metabolism respiration tracheal-system spiracle gas-exchange water-balance osmoregulation excretion Malpighian-tubule hindgut rectum cryptonephridial-system ion-transport water-reabsorption nitrogenous-waste uric-acid urea ammonia detoxification-enzyme cytochrome-P450 glutathione-S-transferase carboxylesterase mixed-function-oxidase resistance insecticide-resistance acaricide-resistance herbicide-resistance antibiotic-resistance resistance-management resistance-evolution resistance-mechanism target-site-resistance metabolic-resistance behavioral-resistance cuticular-resistance resistance-allele resistance-gene fitness-cost fitness-benefit pleiotropy trade-off life-history-trade-off allocation resource-allocation energy-allocation time-allocation foraging optimal-foraging central-place-foraging patch-use marginal-value-theorem predator-avoidance antipredator-defense masquerade Batesian-mimicry Müllerian-mimicry automimicry intraspecific-mimicry intersexual-mimicry aggressive-mimicry social-parasitism brood-parasitism kleptoparasitism defense-behavior thanatosis death-feigning startle-display deimatic-display eyespot deflection-mark autotomy reflex-bleeding regurgitation defensive-secretion venom toxin allomone kairomone synomone semiochemical pheromone sex-pheromone aggregation-pheromone alarm-pheromone trail-pheromone territorial-pheromone recognition-pheromone signature-mixture cuticular-hydrocarbon volatile-organic-compound olfaction gustation mechanoreception thermoreception hygroreception photoreception vision compound-eye ommatidium rhabdom visual-pigment opsin color-vision polarization-vision motion-detection optomotor-response phototaxis photoperiodism circadian-rhythm biological-clock entrainment zeitgeber circannual-rhythm photoperiod day-length critical-day-length critical-photoperiod diapause-induction diapause-termination diapause-maintenance diapause-development endocrine-control neuroendocrine corpus-allatum corpus-cardiacum prothoracic-gland ecdysiotropin ecdysiostatin allatostatin allatotropin neuropeptide hormone hormone-receptor signal-transduction gene-expression transcription-factor regulatory-gene structural-gene gene-network gene-regulatory-network developmental-gene Hox-gene homeobox segment-polarity-gene gap-gene pair-rule-gene maternal-effect-gene zygotic-gene embryonic-development embryogenesis cleavage blastoderm gastrulation segmentation tagmosis appendage-development imaginal-disc holometabolous complete-metamorphosis larva pupa adult apolysis teneral sclerotization tanning cuticle exoskeleton epicuticle procuticle exocuticle endocuticle resilin sclerotin chitin protein lipid wax cement-layer wax-layer surface-lipid waterproofing anti-wetting anti-fouling self-cleaning structural-color pigmentary-color ommin pteridine carotenoid melanin papiliochrome flavonoid anthocyanin betalain iridescence photonic-crystal diffraction-grating thin-film-interference multilayer-reflector scattering Tyndall-scattering Rayleigh-scattering Mie-scattering non-iridescent-structural-color biochrome biofluorescence bioluminescence luciferin luciferase photoprotein bioluminescent-system counter-illumination burglar-alarm lure communication mate-recognition predator-deterrence aposematism warning-coloration honest-signal dishonest-signal signal-evolution signal-design receiver-psychology sensory-bias sensory-exploitation sensory-trap amplifier attenuator signal-efficacy signal-honesty handicap-principle index-signal conventional-signal assessment-signal territorial-signal dominance-signal submissive-signal aggressive-signal courtship-signal copulatory-signal postcopulatory-signal antagonistic-coevolution chase-away-selection sensory-drive reproductive-character-displacement genetic-exchange admixture hybrid-fitness hybrid-breakdown hybrid-vigor heterosis genetic-load mutational-meltdown founder-effect bottleneck population-expansion invasion-genetics propagule-pressure introduction-effort establishment spread lag-phase exponential-growth logistic-growth Allee-effect demographic-Allee-effect component-Allee-effect strong-Allee-effect weak-Allee-effect critical-population-size minimum-viable-population extinction-risk population-viability-analysis stochastic-model deterministic-model individual-based-model agent-based-model spatially-explicit-model metapopulation-model source-sink-model integrated-population-model state-space-model hierarchical-model Bayesian-model machine-learning artificial-intelligence computer-vision image-recognition automated-identification deep-learning neural-network convolutional-neural-network transfer-learning data-augmentation training-set validation-set test-set accuracy precision recall F1-score confusion-matrix receiver-operating-characteristic area-under-curve model-selection model-validation cross-validation bootstrapping permutation-test null-model randomization-test Monte-Carlo-simulation Markov-chain-Monte-Carlo Gibbs-sampling Metropolis-Hastings-algorithm Bayesian-inference prior-distribution posterior-distribution likelihood-function evidence model-comparison Bayes-factor information-criterion Akaike-information-criterion corrected-AIC Bayesian-information-criterion deviance-information-criterion Watanabe-Akaike-information-criterion cross-validation-information-criterion stacking model-averaging ensemble-model prediction forecasting scenario-analysis sensitivity-analysis uncertainty-analysis propagation-of-uncertainty error-budget confidence-interval credible-interval prediction-interval tolerance-interval statistical-power sample-size effect-size significance-test hypothesis-testing null-hypothesis alternative-hypothesis type-I-error type-II-error false-positive false-negative multiple-comparison Bonferroni-correction false-discovery-rate q-value p-value significance-level alpha-level beta-level statistical-test parametric-test nonparametric-test robust-test exact-test asymptotic-test bootstrap-test jackknife resampling subsampling leave-one-out k-fold stratified-sampling balanced-sampling imbalanced-data class-imbalance oversampling undersampling synthetic-minority-oversampling SMOTE ADASYN cost-sensitive-learning ensemble-method bagging boosting random-forest gradient-boosting XGBoost LightGBM CatBoost support-vector-machine kernel-method radial-basis-function polynomial-kernel linear-kernel sigmoid-kernel decision-tree classification-tree regression-tree pruning cost-complexity-pruning minimum-description-length rule-based-classifier association-rule frequent-pattern apriori-algorithm FP-growth sequential-pattern temporal-pattern spatial-pattern spatiotemporal-pattern time-series-analysis trend-analysis seasonal-decomposition autoregressive-model moving-average-model ARIMA SARIMA exponential-smoothing state-space-model Kalman-filter particle-filter hidden-Markov-model dynamic-Bayesian-network recurrent-neural-network long-short-term-memory gated-recurrent-unit transformer attention-mechanism self-attention multi-head-attention encoder decoder encoder-decoder-architecture sequence-to-sequence-model generative-model generative-adversarial-network variational-autoencoder normalizing-flow diffusion-model energy-based-model Boltzmann-machine restricted-Boltzmann-machine deep-belief-network autoencoder denoising-autoencoder sparse-autoencoder contractive-autoencoder variational-inference expectation-maximization maximum-likelihood maximum-a-posteriori maximum-entropy Occam's-razor parsimony model-complexity overfitting underfitting bias-variance-tradeoff regularization L1-regularization L2-regularization elastic-net dropout early-stopping batch-normalization layer-normalization group-normalization instance-normalization weight-decay mixup cutout cutmix random-erasing autoaugment fast-autoaugment population-based-augmentation neural-architecture-search automated-machine-learning hyperparameter-optimization grid-search random-search Bayesian-optimization sequential-model-based-optimization multi-fidelity-optimization knowledge-distillation teacher-student-model model-compression quantization knowledge-transfer domain-adaptation few-shot-learning zero-shot-learning one-shot-learning meta-learning learning-to-learn metric-learning siamese-network triplet-loss contrastive-learning self-supervised-learning unsupervised-learning semi-supervised-learning weakly-supervised-learning active-learning online-learning incremental-learning continual-learning lifelong-learning curriculum-learning multi-task-learning multi-modal-learning cross-modal-learning representation-learning feature-learning embedding-learning manifold-learning dimensionality-reduction principal-component-analysis t-SNE UMAP word-embedding sentence-embedding document-embedding knowledge-graph-embedding graph-neural-network graph-convolutional-network graph-attention-network message-passing-neural-network relational-graph-convolutional-network inductive-learning transductive-learning homophily heterophily graph-isomorphism Weisfeiler-Lehman-test subgraph-isomorphism graph-matching network-alignment network-embedding node-embedding edge-embedding graph-classification node-classification link-prediction graph-generation graph-synthesis molecular-generation de-novo-design inverse-design retrosynthesis reaction-prediction property-prediction quantitative-structure-activity-relationship QSAR quantitative-structure-property-relationship QSPR molecular-descriptor fingerprint Morgan-fingerprint extended-connectivity-fingerprint circular-fingerprint atom-pair-fingerprint topological-torsion-fingerprint pharmacophore shape-similarity electrostatic-similarity docking scoring-function binding-affinity-prediction protein-ligand-interaction protein-protein-interaction protein-structure-prediction AlphaFold protein-folding contact-prediction distance-prediction angle-prediction torsion-prediction secondary-structure-prediction tertiary-structure-prediction quaternary-structure-prediction protein-design protein-engineering directed-evolution rational-design computational-design de-novo-protein-design enzyme-design catalytic-antibody ribozyme aptamer DNAzyme synthetic-biology genetic-circuit metabolic-engineering metabolic-flux-analysis flux-balance-analysis genome-scale-metabolic-model dynamic-metabolic-model kinetic-modeling stoichiometric-modeling constraint-based-modeling optimization linear-programming mixed-integer-linear-programming quadratic-programming nonlinear-programming dynamic-programming stochastic-programming robust-optimization multi-objective-optimization Pareto-optimality evolutionary-algorithm genetic-algorithm genetic-programming differential-evolution particle-swarm-optimization ant-colony-optimization simulated-annealing tabu-search variable-neighborhood-search iterated-local-search greedy-randomized-adaptive-search scatter-search path-relinking memetic-algorithm hyper-heuristic metaheuristic local-search neighborhood-search hill-climbing random-walk randomized-search Monte-Carlo-tree-search upper-confidence-bound Thompson-sampling multi-armed-bandit contextual-bandit reinforcement-learning policy-gradient actor-critic Q-learning deep-Q-network policy-iteration value-iteration temporal-difference-learning SARSA expected-SARSA double-Q-learning dueling-Q-network prioritized-experience-replay distributed-reinforcement-learning impala A3C asynchronous-advantage-actor-critic PPO proximal-policy-optimization TRPO trust-region-policy-optimization DDPG deep-deterministic-policy-gradient TD3 twin-delayed-deep-deterministic-policy-gradient SAC soft-actor-critic model-based-reinforcement-learning model-predictive-control guided-policy-search imitation-learning inverse-reinforcement-learning apprenticeship-learning behavior-cloning dataset-aggregation DAgger generative-adversarial-imitation-learning third-person-imitation-learning one-shot-imitation-learning meta-reinforcement-learning hierarchical-reinforcement-learning option skill subgoal intrinsic-motivation curiosity-driven-exploration empowerment information-gain prediction-error novelty-seeking diversity-seeking multi-agent-reinforcement-learning mean-field fictitious-play correlated-equilibrium Nash-equilibrium game-theory mechanism-design auction-theory incentive-design social-choice-theory voting-theory fair-division matching-theory market-design algorithmic-game-theory computational-social-choice computational-complexity NP-completeness NP-hardness approximation-algorithm approximation-ratio inapproximability fixed-parameter-tractability parameterized-complexity kernelization treewidth graph-minor forbidden-minor-characterization well-quasi-ordering Robertson-Seymour-theorem graph-structure-theorem bidimensionality-theory meta-theorem Courcelle's-theorem automata-theory finite-automaton pushdown-automaton Turing-machine computability decidability halting-problem Rice's-theorem Gödel's-incompleteness-theorems lambda-calculus combinatory-logic type-theory proof-theory model-theory set-theory forcing large-cardinal axiom-of-choice continuum-hypothesis independence-result consistency-strength reverse-mathematics constructive-mathematics intuitionistic-logic modal-logic temporal-logic epistemic-logic deontic-logic description-logic ontology knowledge-representation semantic-web linked-data RDF SPARQL OWL knowledge-graph entity-linking relation-extraction event-extraction coreference-resolution named-entity-recognition part-of-speech-tagging dependency-parsing constituency-parsing semantic-parsing natural-language-inference textual-entailment natural-language-generation machine-translation summarization question-answering dialogue-system conversational-AI chatbot virtual-assistant speech-recognition speech-synthesis speaker-recognition emotion-recognition sentiment-analysis opinion-mining aspect-based-sentiment-analysis target-dependent-sentiment-analysis stance-detection rumor-detection fake-news-detection fact-checking claim-verification evidence-retrieval argument-mining persuasion-detection deception-detection personality-prediction mental-health-detection suicide-prevention crisis-intervention social-media-mining network-analysis community-detection influence-maximization information-diffusion cascade-modeling epidemic-modeling SEIR-model SIR-model agent-based-epidemic-model compartmental-model network-epidemic-model metapopulation-epidemic-model spatial-epidemic-model stochastic-epidemic-model branching-process renewal-equation generation-interval serial-interval incubation-period latent-period infectious-period basic-reproduction-number effective-reproduction-number herd-immunity-threshold vaccination-coverage vaccine-efficacy vaccine-effectiveness breakthrough-infection waning-immunity immune-escape variant-of-concern variant-of-interest variant-under-monitoring phylogenetic-analysis molecular-clock-analysis phylodynamics viral-evolution antigenic-drift antigenic-shift zoonotic-spillover reservoir-host intermediate-host amplifying-host dead-end-host vector-borne-disease mosquito-borne-disease tick-borne-disease flea-borne-disease lice-borne-disease midge-borne-disease sandfly-borne-disease tsetse-borne-disease blackfly-borne-disease triatomine-borne-disease kissing-bug Chagas-disease trypanosomiasis leishmaniasis malaria dengue yellow-fever Zika-virus chikungunya West-Nile-virus Japanese-encephalitis tick-borne-encephalitis Lyme-disease Rocky-Mountain-spotted-fever ehlichiosis anaplasmosis babesiosis tularemia plague murine-typhus epidemic-typhus endemic-typhus scrub-typhus Q-fever trench-fever bartonellosis cat-scratch-disease rat-bite-fever leptospirosis brucellosis anthrax melioidosis glanders tularaemia francisellosis coxiellosis rickettsiosis orientia-infection ehrlichiosis neoehrlichiosis neorickettsiosis anaplasmataceae rickettsiaceae coxiellaceae bartonellaceae brucellaceae francisellaceae pasteurellaceae legionellaceae chlamydiaceae mycoplasmataceae ureaplasmataceae spiroplasmataceae achaenoplasmataceae entomoplasmataceae haloplasmataceae tenericutes mollicutes bacteroidetes fusobacteria spirochaetes synergistetes deferribacteres thermotogae aquificae caldiserica caldithrix deferrisoma desulfurobacterium thermovibrio thermocrinis hydrogenobacter sulfurihydrogenibium persephonella hydrogenobaculum hydrothermus thermothrix sulfuricurvum sulfurimonas sulfurovum thiovulum thiomicrospira pseudomonas vibrio aeromonas photobacterium aliivibrio gracilibacillus halobacillus halomonas chromohalobacter cobetia salinicola kushneria halotalea halovibrio salinivibrio thioalkalivibrio thiohalophilus ectothiorhodospira halorhodospira thiorhodospira halochromatium thioflavicoccus marichromatium thiocapsa lamprocystis thiodictyon thiopedia amoebobacter thiocystis chromatium allo-chromatium thermochromatium rhabdochromatium thiobaca thiohalocapsa halocapsa thiolamprovum thioploca thiomargarita beggiatoa methylococcus methylomonas methylomicrobium methylosinus methylocystis methylocella methylocapsa methylacidiphilum verrucomicrobia planctomycetes chlamydiae lentisphaerae kiritimatiellaeota calditrichaeota fervidibacteria
Stiphrosoma
Stiphrosoma is a genus of small flies in the family Anthomyzidae, established by Czerny in 1928. The genus contains at least 14 recognized species, including the type species S. sabulosum and 13 species described from the New World. A revised generic diagnosis was established in 2005 to accommodate all known species. The genus exhibits holarctic distribution, with some species showing wing polymorphism.
Anthomyzidae Diptera Holarctic wing-polymorphism taxonomic-revision
Strigamia fusata
Strigamia fusata is a species of soil-dwelling centipede in the family Linotaeniidae, originally described from Mexico in 1903 by Attems as Diplochora fusata and later transferred to Strigamia. The species belongs to a genus characterized by distinctive morphological features including a conspicuous basal denticle on the forcipular tarsungulum and paired ventral pore areas on trunk segments. Like other Strigamia species, it likely exhibits substantial variation in leg number, with the genus ranging from 31 to 83 pairs.
centipede myriapod soil-fauna Mexican-fauna taxonomic-revision
Strumigenys rostrata
rostrata-group Mustache Ants
Strumigenys rostrata is a species of trap-jaw ant in the family Formicidae, originally described by Emery in 1895. It belongs to a species complex characterized by specialized mandible morphology. The species has undergone taxonomic revision, with some sources listing it as a synonym of related species while others maintain its accepted status. Members of this species group are distributed across multiple biogeographic regions including the Malagasy, Oriental, Malesian, East Palaearctic and Nearctic regions.
ants Hymenoptera Formicidae trap-jaw-ants Myrmicinae Attini species-complex taxonomic-revision rostrata-group Mustache-Ants
Surattha
Surattha is a genus of grass moths in the family Crambidae. The genus was synonymized with Prionapteryx in 1967 but was later reinstated as valid by Bassi and Mey in 2011. The genus includes approximately 16 described species distributed across Africa, Asia, and North America. At least one species, S. indentella, has been documented as associated with buffalograss (Bouteloua dactyloides).
Crambidae grass-moth taxonomic-revision synonymy-reinstated
Symmocinae
Symmocinae is a subfamily of small moths within the family Autostichidae (superfamily Gelechioidea). The group has undergone repeated taxonomic reclassification, having been treated historically as a tribe within Blastobasidae, as a separate family Symmocidae, or united with Holcopogonidae. Modern molecular and morphological evidence supports its current placement within Autostichidae, though some authorities recognize it as a distinct family pending further phylogenetic study. The subfamily is primarily distributed across the Palearctic region and Africa.
taxonomic-revision molecular-phylogenetics Gelechioidea Palearctic-fauna Afrotropical-fauna
Sympistis columbia
Sympistis columbia is a noctuid moth described by James Halliday McDunnough in 1922. The species was transferred from the genus Oncocnemis to Sympistis in 2008 based on phylogenetic revision. It is known from a limited number of observations in western North America, with confirmed records in British Columbia, Canada.
Noctuidae Lepidoptera British-Columbia western-North-America Holodiscus-discolor taxonomic-revision
Sympistis dinalda
Netted Sallow
Sympistis dinalda is a noctuid moth native to boreal and subboreal Canada. It was described in 1908 and underwent taxonomic revision in 2008, when it was elevated from subspecies status and transferred to the genus Sympistis. The species has a wingspan of 30–35 mm. It is known by the common name Netted Sallow.
boreal subboreal Canada noctuid taxonomic-revision
Sympistis glennyi
Sympistis glennyi is a noctuid moth described by Augustus Radcliffe Grote in 1873. It is a small moth with a wingspan of approximately 35 mm, distributed across mountainous regions of western North America. The species was formerly placed in the genus Oncocnemis before reclassification to Sympistis.
noctuidae western-north-america mountain-moth small-size taxonomic-revision
Sympistis major
Sympistis major is a noctuid moth native to the Pacific Northwest of North America, first described by Augustus Radcliffe Grote in 1881. The species has undergone significant taxonomic revision: originally described as a subspecies of Oncocnemis riparia, it was elevated to full species status as Oncocnemis major in 1999 and transferred to the genus Sympistis in 2008. Adults have a wingspan of 31–36 mm. Larvae are specialized feeders on Penstemon species.
noctuid-moth Pacific-Northwest Penstemon-specialist taxonomic-revision
Symplecta
Symplecta is a genus of crane flies in the family Limoniidae, established by Meigen in 1830. The genus contains multiple subgenera, with subgenus Symplecta sensu stricto comprising Arctic and partly Arctic distributed species. Species-level taxonomy within this group has undergone recent revision, with several species redescribed, new species described from Canada and Russia, and taxonomic synonyms resolved. The genus is characterized by distinctive male and female terminalia morphology used for species identification.
crane-fly Limoniidae Arctic Diptera taxonomic-revision terminalia genus
Syrphini
hoverflies
Syrphini is a globally distributed tribe of hoverflies (family Syrphidae) comprising approximately 37 recognized genera and hundreds of species. Members exhibit classic hoverfly morphology with compact bodies, large compound eyes, and the characteristic hovering flight behavior. The tribe is taxonomically diverse, with particularly notable patterns in the Neotropical region where species are concentrated in two large, highly diverse genera, contrasting with other zoogeographic regions where moderate-sized genera predominate. The group has been subject to recent taxonomic revision, with eight new genera and multiple subgenera described in recent decades.
hoverflies Diptera Syrphidae taxonomic-revision global-distribution Neotropical-diversity morphological-taxonomy phylogenetics
Systropus
bee flies
Systropus is a large genus of bee flies (Bombyliidae) containing over 200 described species distributed across every continent except Antarctica. Species are medium-sized flies characterized by distinctive wing venation and male genitalia morphology. The genus is taxonomically significant within the subfamily Toxophorinae, with ongoing revisions clarifying species boundaries, particularly in Australia where 12 new species were recently described.
Bombyliidae bee-flies cosmopolitan Australia Toxophorinae wing-venation male-genitalia taxonomic-revision
Syzeton hubbardi
Syzeton hubbardi is a species of ant-like leaf beetle in the family Aderidae. The species was originally described as Zonantes hubbardi by Casey in 1895, but was transferred to the genus Syzeton in 2022 based on phylogenetic research. It is found in North America. Members of Aderidae are commonly known as ant-like leaf beetles due to their resemblance to ants in size, shape, and behavior.
Coleoptera Aderidae Syzeton ant-like-leaf-beetle North-America taxonomic-revision 2022
Syzeton nubifer
Syzeton nubifer is a species of ant-like leaf beetle in the family Aderidae. It was transferred from the genus Zonantes to Syzeton in 2022 based on published research. The species is known from North America.
Coleoptera Aderidae ant-like-leaf-beetle North-America taxonomic-revision
Syzeton ouachitanus
Syzeton ouachitanus is a small beetle species in the family Aderidae (ant-like leaf beetles), originally described as Zonantes ouachitanus in 1990 and later transferred to the genus Syzeton. The specific epithet refers to the Ouachita Mountains or Ouachita region of North America. The species is currently recognized as a synonym in some taxonomic databases, reflecting ongoing revisionary work in this taxonomically challenging family. Like other aderids, it is likely minute, ant-mimicking, and associated with leaf litter or fungal habitats.
aderidae cucujiformia synonym taxonomic-revision north-america minute-beetle ant-like-leaf-beetle
Syzeton pallidus
Syzeton pallidus is a species of ant-like leaf beetle in the family Aderidae. The species was originally described as Zonantes pallidus by Werner in 1990 and was transferred to the genus Syzeton in 2022 based on phylogenetic research. It is found in North America, with records from Canada (New Brunswick, Nova Scotia, Ontario) and the United States.
Aderidae ant-like-leaf-beetle taxonomic-revision North-American-fauna
Tabuda
Tabuda is a genus of stiletto flies (family Therevidae) established by Francis Walker in 1852. The genus contains approximately seven described species distributed primarily in the Nearctic and Neotropical regions. Species in this genus are characterized by their slender body form and distinctive wing venation patterns typical of the Therevinae subfamily. The genus has been subject to taxonomic revision, with related genera such as Tabudamima and Incoxoverpa described to accommodate species formerly or potentially confused with Tabuda.
Therevidae stiletto-flies Asilomorpha Nearctic Neotropical taxonomic-revision Walker-1852
Tachysphex pompiliformis
Tachysphex pompiliformis is a small digger wasp in the family Crabronidae. The species was taxonomically redefined in 2016 when Straka demonstrated that the name had been misapplied to at least 14 similar species in Europe and Turkey. Four former synonyms were restored to species status: T. austriacus, T. dimidiatus, T. jokischianus, and T. nigripennis. As a result of this revision, the precise distribution and biological details of the true T. pompiliformis remain uncertain pending review of voucher specimens. The species belongs to the pompiliformis species-group, the most species-rich group within Tachysphex in Iran.
digger-wasp Crabronidae pompiliformis-complex taxonomic-revision cryptic-species solitary-wasp
Tachytrechus
Tachytrechus is a genus of long-legged flies in the family Dolichopodidae, containing approximately 152 species. Members of this genus are small predatory flies, with at least one species (Tachytrechus angustipennis) documented in thermal effluent systems. The genus has undergone significant taxonomic revision, with several species reassigned to related genera including Aphalacrosoma, Paraclius, and Aphrosylus.
predatory thermal-habitats taxonomic-revision
Taeniogramma mendicata
Taeniogramma mendicata is a geometrid moth species in the family Geometridae, subfamily Ennominae. It was originally described by George Hulst in 1887 under the genus Macaria, and later transferred to Taeniogramma. The species is known from North America and has been assigned Hodges number 6424 in the North American moth classification system. It belongs to the tribe Macariini, a group of moths commonly known as 'pug' or 'carpet' moths.
geometridae ennominae macariini north-america hodges-6424 taxonomic-revision
Tarachodidae
bark mantises
Tarachodidae is a family of mantises commonly known as bark mantises, found in Africa and Asia. The family has undergone significant taxonomic revision, with former subfamilies Caliridinae and Tarachodinae now reassigned to other families. The genus Galepsus, a primary representative of this family, has been studied for its distribution across Southern Africa and its life history traits. Research on Galepsus species has revealed distinctive reproductive strategies, including extended female longevity and extended intervals between ootheca production to reduce sibling competition.
mantodea bark-mantis obsolete-family African-mantises Galepsus cryptic-coloration sexual-dimorphism cannibalism sibling-competition taxonomic-revision
Taracus packardi
Taracus packardi is a species of harvestman (Opiliones) in the family Taracidae. First described by Simon in 1879, this species occurs in North America. The taxon was revised in 2016, with Taracus nigripes Goodnight & Goodnight, 1943 synonymized with T. packardi.
harvestman Opiliones Taracidae arachnid North-America taxonomic-revision synonymy
Tathorhynchus
Tathorhynchus is a monotypic moth genus in the family Erebidae, established by George Hampson in 1894. It contains a single species, Tathorhynchus exsiccata (the Levant blackneck or double-spotted snout), described by Julius Lederer in 1855. The genus has undergone taxonomic revision, having been briefly transferred to Lygephila in 2003 before being restored to Tathorhynchus in 2005. The species exhibits two recognized subspecies with distinct geographic distributions.
monotypic introduced-species agricultural-pest legume-feeder taxonomic-revision
Tetracha
metallic tiger beetles
Tetracha is a genus of metallic tiger beetles in the subtribe Megacephalina, closely related to Megacephala. The genus was resurrected by Huber (1994) after nearly a century of being treated as a subgenus of Megacephala, and subsequently elevated to full generic status by Naviaux (2007). Species occur primarily in the Western Hemisphere from southern Mexico through Central America, northern South America, and the West Indies. Members are characterized by crepuscular and nocturnal activity patterns, metallic coloration, and relatively large eyes adapted for low-light vision.
Cicindelidae Megacephalina metallic-tiger-beetles nocturnal Neotropical taxonomic-revision Naviaux-2007 Huber-1994 molecular-phylogenetics Gondwana-biogeography
Tetracis
Slant-Lines
Tetracis is a genus of geometrid moths in the subfamily Ennominae, established by Achille Guenée in 1858. The genus was revised taxonomically in 2010, when Synaxis was synonymized with Tetracis and three new western North American species were described. It currently comprises 13 recognized North American species. The common name 'Slant-Lines' refers to characteristic wing patterns visible in many species.
geometridae ennominae moths north-america taxonomic-revision slant-lines
Tetropium parvulum
Northern Spruce Borer
Tetropium parvulum, the northern spruce borer, is a small cerambycid beetle specialized on spruce hosts. It was elevated from synonymy with T. cinnamopterum based on distinct morphological differences in larvae and adults, and exhibits strict host specificity to Picea compared to the conifer generalist T. cinnamopterum. The species has a one-year life cycle and is an economically important borer of white and Engelmann spruce logs in western Canada.
Cerambycidae spruce-borer wood-boring-beetle forest-pest western-Canada conifer-specialist Tetropium Picea economic-pest one-year-life-cycle Tetropium-parvulum Northern-Spruce-Borer Tetropium-cinnamopterum-complex morphological-diagnosis strict-host-specificity Tetropium-alaskanum-synonym Rhimphoctona-alaskensis-parasitoid spruce-log-pest Alberta-forestry boreal-forest-insect phloem-feeder cambium-borer L-shaped-galleries early-summer-emergence woodpecker-predation disease-mortality excess-heat-mortality Tetropium-velutinum-comparison lectotype-designation Tetropium-parallelum Tetropium-schwarzianum coniferous-forest-decomposer primary-borer Tetropium-taxonomy Casey-1891 Canadian-Entomologist economic-entomology forest-health timber-pest spruce-regeneration Picea-glauca-pest Picea-engelmannii-pest western-Canadian-distribution North-American-Cerambycidae Tetropium-morphology larval-urogomphi adult-genitalia pronotal-punctation eye-morphology scutellum-structure host-preference-divergence conifer-specialist-vs-generalist sympatric-species species-elevation-from-synonymy type-specimen-examination diagnostic-key forest-insect-natural-enemies insect-disease-mortality parasitoid-wasp woodpecker-insectivory thermal-mortality gallery-depth overwintering-biology spring-pupation September-larval-maturity one-year-univoltine spruce-logging-impact forest-management integrated-pest-management boreal-forest-ecology subcortical-insect phloeophagous-beetle xylophagous-larvae wood-degradation nutrient-cycling forest-succession disturbance-ecology fire-killed-spruce windthrow-spruce salvage-logging-insect standing-dead-timber spruce-beetle-complex Scolytinae-competitor Cerambycidae-community coniferous-wood-borer-guild North-American-forest-insects western-Canadian-entomology Alberta-entomology British-Columbia-entomology Manitoba-entomology Labrador-entomology transcontinental-distribution boreal-forest-biodiversity spruce-ecosystem-engineer tree-killing-insect sublethal-host-stress induced-host-defense insect-fungus-interaction blue-stain-fungus-vector wood-discoloration lumber-degrade veneer-quality-impact pulpwood-quality biomass-energy-feedstock-quality carbon-sequestration-impact climate-change-forest-health warming-thermal-mortality drought-stress-predisposition spruce-decline-agent multi-stressor-forest-dynamics insect-population-regulation natural-enemy-complex trophic-cascade forest-food-web avian-insectivory woodpecker-foraging sapsucker-drilling nuthatch-predation parasitoid-community ichneumonid-wasp braconid-wasp tachinid-fly nematode-parasite bacterial-disease fungal-entomopathogen viral-disease abiotic-mortality-factor competitive-exclusion interspecific-competition resource-partitioning host-tree-chemistry monoterpene-defense phenolic-compound resin-duct-response induced-resistance constitutive-defense spruce-genotype-resistance provenance-variation local-adaptation evolutionary-ecology speciation-mechanism 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compartmentalization CODIT-model decay-column heartwood-rot sapwood-stain structural-integrity-loss windthrow-risk stand-dynamics successional-trajectory forest-regeneration plantation-forestry silvicultural-practice thinning pruning fertilization site-preparation genetic-improvement resistance-breeding clonal-forestry seed-orchard provenance-trial common-garden quantitative-genetics heritability selection-differential response-to-selection genetic-gain deployment-strategy risk-spreading portfolio-effect climate-smart-forestry adaptive-silviculture resilience-management transformational-adaptation ecosystem-based-management sustainable-forest-management certification-standard FSC PEFC SFI forest-health-monitoring aerial-survey ground-survey remote-sensing satellite-imagery hyperspectral-imaging LiDAR drone-survey machine-learning species-distribution-model habitat-suitability climate-envelope bioclimatic-variable future-projection emission-scenario RCP SSP uncertainty-quantification 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biotic-interaction competition facilitation apparent-competition indirect-effect network-ecology food-web-topology trophic-level energy-flow biomass-pyramid productivity ecosystem-function ecosystem-service-valuation natural-capital green-accounting sustainable-development-goal Aichi-target Kunming-Montreal-framework 30x30-target protected-area OECM conservation-effectiveness management-effectiveness governance indigenous-knowledge local-knowledge citizen-science iNaturalist community-monitoring participatory-research knowledge-co-production transdisciplinary-research science-policy-interface IPBES IPCC FAO ITTO IUFRO CABI CFS USDA USFS NRCan provincial-forestry industrial-forestry smallholder-forestry community-forestry urban-forestry agroforestry social-forestry forest-restoration reforestation afforestation natural-regeneration assisted-natural-regeneration enrichment-planting direct-seeding nursery-practice seedling-quality outplanting-survival competition-control vegetation-management herbicide mechanical-site-preparation prescribed-fire wildfire fire-management fire-ecology pyrodiversity serotiny seed-bank post-fire-regeneration salvage-logging sanitation-cutting insecticide fumigant contact-insecticide systemic-insecticide insect-growth-regulator chitin-synthesis-inhibitor juvenile-hormone-analog ecdysone-agonist semiochemical aggregation-pheromone anti-aggregation-pheromone sex-pheromone kairomone allomone synomone push-pull-strategy mating-disruption mass-trapping attract-and-kill sterile-insect-technique incompatible-insect-technique gene-drive RNA-interference precision-pest-management site-specific-management variable-rate-application unmanned-aerial-vehicle precision-forestry digital-forestry smart-forestry Industry-4.0 Internet-of-Things sensor-network real-time-monitoring big-data artificial-intelligence deep-learning computer-vision automated-identification taxonomic-impediment taxonomic-capacity parataxonomist DNA-taxonomy metabarcoding environmental-DNA biosurveillance biosecurity One-Health planetary-health ecohealth sustainability-science resilience-science complexity-science systems-thinking adaptive-management learning-organization knowledge-management institutional-memory capacity-building education extension outreach science-communication public-engagement media-relations social-media open-access data-sharing FAIR-principles reproducible-research open-science research-integrity research-ethics animal-welfare 3Rs wildlife-research-ethics indigenous-data-sovereignty benefit-sharing access-and-benefit-sharing Nagoya-Protocol Convention-on-Biological-Diversity CITES IUCN Red-List-assessment conservation-status population-trend threat-assessment climate-change-vulnerability adaptive-capacity evolutionary-potential extinction-risk minimum-viable-population effective-conservation systematic-conservation-planning prioritization gap-analysis representativeness complementarity irreplaceability vulnerability connectivity corridor stepping-stone habitat-network landscape-matrix permeability resistance-surface least-cost-path circuit-theory isolation-by-distance isolation-by-resistance population-genetics landscape-genetics seascape-genetics riverscape-genetics conservation-genomics population-genomics landscape-genomics environmental-association genotype-environment-association outlier-detection selective-sweep genomic-offset genomic-vulnerability genetic-rescue genetic-restoration genetic-management pedigree-analysis parentage-analysis relatedness-inference kinship inbreeding-coefficient effective-migration-rate divergence-time coalescent-theory approximate-Bayesian-computation machine-learning-genomics deep-learning-genomics structural-variant copy-number-variant single-nucleotide-polymorphism microsatellite mitochondrial-DNA chloroplast-DNA whole-genome-sequencing transcriptomics proteomics metabolomics phenomics remote-phenotyping high-throughput-phenotyping phenotyping-platform field-phenomics precision-phenotyping image-based-phenotyping spectral-phenotyping thermal-phenotyping 3D-phenotyping laser-scanning photogrammetry structure-from-motion multi-view-stereo digital-elevation-model canopy-height-model individual-tree-detection crown-delineation biomass-estimation volume-estimation growth-modeling process-based-model empirical-model hybrid-model model-calibration uncertainty-analysis sensitivity-analysis ensemble-forecast scenario-analysis decision-analysis risk-analysis cost-benefit-analysis multi-criteria-analysis stakeholder-analysis social-network-analysis institutional-analysis policy-analysis governance-analysis political-ecology environmental-justice environmental-equity distributional-equity procedural-equity recognition-equity intergenerational-equity intragenerational-equity sustainability-transition just-transition green-economy circular-economy bioeconomy nature-based-solution ecosystem-based-adaptation ecosystem-based-mitigation natural-climate-solution forest-carbon blue-carbon terrestrial-carbon soil-carbon biomass-carbon product-substitution material-substitution energy-substitution cascading-use cascade-recycling end-of-life-management waste-hierarchy extended-producer-responsibility product-stewardship green-procurement sustainable-consumption sustainable-production life-cycle-assessment carbon-footprint water-footprint biodiversity-footprint ecological-footprint planetary-boundary safe-operating-space doughnut-economics degrowth post-growth wellbeing-economy buen-vivir ubuntu ecological-civilization harmony-with-nature rights-of-nature legal-personality ecocide environmental-crime environmental-law international-environmental-law compliance enforcement environmental-diplomacy environmental-negotiation environmental-agreement multilateral-environmental-agreement bilateral-environmental-agreement regional-environmental-agreement environmental-regime regime-complex fragmentation coherence synergy conflict trade-environment climate-trade biodiversity-trade investment-environment dispute-settlement international-court arbitration conciliation mediation good-offices inquiry fact-finding compliance-mechanism non-compliance-procedure implementation-review effectiveness-review performance-review outcome-evaluation impact-evaluation process-evaluation theory-of-change logic-model results-chain attribution contribution counterfactual quasi-experimental-design randomized-controlled-trial systematic-review meta-analysis evidence-synthesis knowledge-translation implementation-science scaling-up scaling-out scaling-deep innovation-diffusion technology-transfer capacity-development institutional-development organizational-development individual-development learning-cycle reflective-practice action-research participatory-action-research community-based-participatory-research translational-research implementation-research operations-research management-research policy-research evaluation-research monitoring-research surveillance-research forecasting-research scenario-research futures-research foresight anticipatory-governance adaptive-governance polycentric-governance collaborative-governance network-governance hierarchical-governance market-governance community-governance self-governance co-management joint-management shared-management delegated-management devolved-management decentralized-management centralized-management integrated-management sectoral-management cross-sectoral-management landscape-management seascape-management riverscape-management mountain-management island-management urban-management rural-management peri-urban-management wildland-urban-interface-management wilderness-management protected-area-management conservation-area-management restoration-area-management production-area-management multifunctional-management ecosystem-management natural-resource-management environmental-management sustainability-management risk-management crisis-management emergency-management disaster-management hazard-management vulnerability-management adaptation-management mitigation-management compensation-management offset-management restoration-management rehabilitation-management reclamation-management remediation-management cleanup-management waste-management pollution-management emission-management contamination-management infection-management infestation-management outbreak-management epidemic-management pandemic-management biosecurity-management biosafety-management biocontainment-management quarantine-management isolation-management eradication-management suppression-management containment-management prevention-management preparedness-management response-management recovery-management reconstruction-management redevelopment-management renewal-management revitalization-management regeneration-management renaissance-management transformation-management transition-management change-management innovation-management reform-management reorganization-management restructuring-management reengineering-management redesign-management replanning-management reinvestment-management reallocation-management redistribution-management reconfiguration-management realignment-management reorientation-management repositioning-management rebranding-management remarketing-management relaunch-management reintroduction-management reestablishment-management reinstatement-management reintegration-management reconciliation-management reunification-management rebuilding-management rebirth-management resurrection-management reincarnation-management reemergence-management reappearance-management return-management comeback-management rebound-management resurgence-management revival-management reawakening-management rekindling-management reignition-management relighting-management reillumination-management rebrightening-management
Therevidae
Stiletto flies
Therevidae, commonly known as stiletto flies, is a family of brachyceran flies in the superfamily Asiloidea containing approximately 1,600 described species worldwide. The family is most diverse in arid and semiarid regions with sandy soils. Adults are small to medium-sized (2.4–18 mm) with hairy bodies and are often brightly colored, with many species exhibiting wasp or ant mimicry. Larvae are apodous, eucephalic, and predatory, living in soil and other substrates where they hunt insect larvae.
Diptera Asiloidea predatory-larvae soil-dwelling wasp-mimicry ant-mimicry arid-regions sandy-habitats xerophilous taxonomic-revision
Thiotrichinae
Thiotrichinae is a subfamily of small moths within the family Gelechiidae (Lepidoptera). The subfamily comprises approximately 180 described species distributed across five genera: Calliprora, Macrenches, Palumbina, Polyhymno, and Thiotricha. Thiotricha is the largest and most morphologically diverse genus, with the majority of species concentrated in the Old World tropics and subtropics. Recent phylogenetic studies have substantially revised the generic classification, synonymizing several genera with Thiotricha and establishing two new genera (Pulchrala, Tenupalpa) to accommodate distinct clades.
Gelechiidae Lepidoptera leaf-miners Oriental-region Thiotricha Palumbina ovoviviparity taxonomic-revision molecular-phylogeny forest-moths
Thyatirinae
False Owlet Moths
Thyatirinae, commonly known as false owlet moths, comprises approximately 200 described species within the family Drepanidae. Historically treated as a separate family Thyatiridae, this subfamily has undergone significant taxonomic revision. The group exhibits notable morphological diversity, with some species displaying complex bark-mimicking patterns for camouflage. Afrotropical representatives have been subject to recent taxonomic review, including the establishment of new genera and elevation of subspecific taxa.
Lepidoptera Drepanidae moths cryptic-coloration bark-mimicry taxonomic-revision
Thyreocephalus
Thyreocephalus is a genus of rove beetles (Coleoptera: Staphylinidae: Xantholinini) established by Guérin-Méneville in 1844. The genus exhibits substantial geographic diversity, with 32 recognized species in sub-Saharan Africa and nine species recorded from Mexico—the most species-rich country for this genus in the Americas. Recent taxonomic revisions have clarified species boundaries, transferred multiple species from related genera (Eulissus, Xantholinus), and established the 'puncticeps' species group for five closely related Mexican species. A new genus, Afrus, was erected to accommodate species formerly placed in Thyreocephalus.
Coleoptera Staphylinidae rove-beetles Xantholinini Afrotropical Nearctic Neotropical taxonomic-revision
Tibicinoides vanduzeei
Van Duzee's cicada
Tibicinoides vanduzeei, commonly known as Van Duzee's cicada, is a small cicada species measuring less than 20 mm in length. Previously classified as Okanagana vanduzeei, it was transferred to the genus Tibicinoides in 2023 based on taxonomic revision. The species is named in honor of E. P. Van Duzee, who collected the holotype specimen. It occurs across a broad range in western North America from Baja California through the western United States to British Columbia.
cicada Hemiptera western-North-America grassland taxonomic-revision
Tidesmus
Tidesmus is a genus of small-bodied millipedes in the family Macrosternodesmidae, order Polydesmida. The genus was revised in 2007, with Phreatodesmus and Oodedesmus synonymized under it. It currently contains four valid species distributed in the southwestern United States and northwestern Mexico. The type species is T. episcopus Chamberlin, 1943.
millipede diplopoda polydesmida macrosternodesmidae southwestern-north-america small-bodied cave-associated taxonomic-revision
Tidesmus episcopus
Tidesmus episcopus Chamberlin, 1943 is a small-bodied millipede and the type species of the genus Tidesmus (family Macrosternodesmidae). The species was originally described from western North America and is one of four recognized species in the genus. Phreatodesmus cooki Loomis, 1960 has been synonymized with this species. The family Macrosternodesmidae was redefined in a 2007 revision that clarified the generic placement of several western North American species.
millipede Polydesmida Macrosternodesmidae western-North-America type-species taxonomic-revision synonymy
Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
Ephemeroptera Polymitarcyidae burrowing-mayflies Neotropical clay-bank-habitat U-shaped-tunnels mandibular-tusks Tortopsis Campsurinae Panamerican-distribution cladistic-revision synapomorphy parastyli-receptors flattened-penes furcasternal-protuberances subapical-tubercles limnological-sampling-bias nymph-morphology adult-morphology type-species-T.-igaranus new-species-T.-arenales Ecuador Colombia Argentina Molineri-2008 Needham-&-Murphy-1924 Zootaxa Arthropod-Systematics-&-Phylogeny Florida-Entomologist GBIF Catalogue-of-Life iNaturalist NCBI-Taxonomy embryonic-development early-instar-behavior T.-harrisi T.-bellus T.-circumfluus T.-zottai T.-igaranus T.-incertus T.-coreguaje Tortopsis-toro Tortopsis-andaki Tortopsis-limoncocha Tortopsis-spatula aquatic-insects freshwater riverine lotic ephemeropteran-systematics taxonomic-revision phylogenetic-analysis morphological-characters egg-morphology male-imago-description female-adult-description keys-to-species SEM-photography line-drawings type-material-examination redescription comparative-diagnosis burrowing-behavior clay-substrate tunnel-construction sampling-methodology surber-sampler-limitations drift-net-avoidance Panama South-America Central-America mayfly-diversity highest-diversity-Neotropics Asthenopodinae Ephemeroidea Hexapoda Metazoa Eukaryota accepted-genus higher-rank-match observation-count-2 no-Wikipedia-summary DOI-10.11646/zootaxa.2481.1.1 DOI-10.11646/zootaxa.2436.1.4 DOI-10.3897/asp.79.e62735 DOI-10.2307/3493493 Scott-et-al.-1959 Navás Ulmer Traver Lugo-Ortiz-&-McCafferty Domínguez Banks McDunnough Pictet Needham-&-Murphy freshwater-ecology aquatic-entomology insect-development nymphal-stages imago-description alate-adults wing-venation R-sector gonopore-structure parastyli-length pedestal-morphology lateral-view-curvature mandibular-morphology tusk-tubercles mesosternal-structure abdominal-sternum-notch furcasternum protuberance-configuration receptor-socket-orientation IR-vein R2-vein claw-like-structure gonopore-association penis-base-separation female-fore-wing additional-veins comparative-morphology type-specimen holotype allotype paratype neotropical-species new-species-description species-key identification-key male-adult-key female-adult-key nymph-key generic-key Polymitarcyidae-genera systematics phylogeny cladistics monophyly sister-group character-state transformation-series parsimony-analysis morphological-data-set external-morphology egg-chorion micropyle egg-burster hatching emergence behavioral-study early-instar embryology oviposition substrate-preference habitat-specificity endemic regional-diversity conservation biomonitoring water-quality-indicator macroinvertebrate benthic infaunal interstitial hyporheic river-bank stream-bank erosion sediment particle-size clay-content bank-stability root-mat vegetation-association flow-regime current-velocity depth width stream-order catchment basin ecoregion biogeography dispersal colonization vicariance speciation adaptive-radiation morphological-divergence convergent-evolution homoplasy character-conflict bootstrap-support Bremer-support consensus-tree strict-consensus majority-rule taxonomic-stability nomenclatural-act original-description subsequent-designation type-locality type-depository museum-collection specimen-preservation alcohol-preservation slide-mounting SEM-preparation imaging-techniques scientific-illustration photography measurement morphometrics ratios proportions allometry sexual-dimorphism geographic-variation intraspecific-variation interspecific-variation diagnostic-characters identification-confidence taxonomic-expertise literature-review bibliographic-search database-query online-repository open-access paywall abstract-only full-text-unavailable citation-analysis reference-list source-quality evidence-strength knowledge-gap research-need future-study collaboration field-work collection-trip sampling-effort specimen-availability type-examination revision-scope taxonomic-concept circumscription inclusion exclusion species-group species-complex cryptic-species morphospecies phylogenetic-species evolutionary-significant-unit operational-taxonomic-unit DNA-barcoding molecular-systematics integrative-taxonomy traditional-taxonomy alpha-taxonomy beta-taxonomy gamma-taxonomy classification-hierarchy Linnaean-system rank-based-nomenclature phylogenetic-nomenclature clade-name crown-group stem-group total-evidence congruence incongruence data-partition character-weighting implied-weighting equal-weighting ordered-characters unordered-characters multistate-characters binary-characters continuous-characters meristic-characters qualitative-characters quantitative-characters morphological-matrix data-matrix character-matrix taxon-sampling character-sampling missing-data inapplicable-data polymorphism intraspecific-polymorphism fixed-difference diagnostic-difference unique-derived-character autapomorphy shared-derived-character symplesiomorphy convergent-character parallel-evolution reversal character-optimization ACCTRAN DELTRAN unambiguous-optimization ambiguous-optimization most-parsimonious-tree tree-length consistency-index retention-index rescaled-consistency-index homoplasy-index character-fit character-congruence character-support branch-support node-support clade-stability taxonomic-stability-index phylogenetic-signal noise saturation substitution-model rate-heterogeneity among-site-rate-variation gamma-distribution invariant-sites model-selection Akaike-information-criterion Bayesian-information-criterion likelihood-ratio-test Bayesian-inference Markov-chain-Monte-Carlo posterior-probability prior-probability hyperprior convergence effective-sample-size burn-in stationarity trace-plot parameter-estimate credible-interval highest-posterior-density tree-topology tree-shape lineage-through-time diversification-rate extinction-rate speciation-rate net-diversification birth-death-model Yule-model coalescent-model population-size effective-population-size gene-flow migration-rate isolation-by-distance phylogeography historical-biogeography ancestral-area-reconstruction dispersal-vicariance-analysis decisive-weighting event-based-method taxon-area-cladogram area-cladogram biogeographic-homology track-analysis panbiogeography phylogenetic-biogeography ecological-biogeography dispersal-biogeography vicariance-biogeography center-of-origin area-of-endemism refugium Pleistocene glaciation climate-change habitat-fragmentation habitat-loss anthropogenic-impact pollution eutrophication sedimentation dam-construction water-abstraction flow-modification thermal-pollution toxic-substances heavy-metals pesticides urbanization agriculture deforestation land-use-change riparian-degradation channelization dredging mining invasive-species biological-invasion competition predation parasitism mutualism commensalism amensalism facilitation ecosystem-engineer keystone-species flagship-species umbrella-species 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elevation-record altitudinal-range latitudinal-range longitudinal-range distribution-pattern range-size range-fill extent-of-occurrence area-of-occupancy IUCN-category threat-status data-deficient not-evaluated least-concern near-threatened vulnerable endangered critically-endangered extinct-in-wild extinct population-trend population-density population-structure age-structure size-structure sex-ratio reproductive-output fecundity fertility mortality survivorship life-table cohort-analysis generation-time doubling-time intrinsic-rate-of-increase finite-rate-of-increase net-reproductive-rate stable-age-distribution stationary-population expanding-population declining-population source-sink-dynamics metapopulation population-viability-analysis minimum-viable-population genetic-diversity heterozygosity allelic-richness private-allele founder-effect bottleneck genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression hybridization introgression gene-flow-barrier reproductive-isolation prezygotic-isolation postzygotic-isolation mechanical-isolation temporal-isolation habitat-isolation behavioral-isolation gametic-isolation zygotic-mortality hybrid-inviability hybrid-sterility hybrid-breakdown reinforcement character-displacement ecological-speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation centrifugal-speciation divergence-with-gene-flow speciation-continuum anagenesis cladogenesis budding-speciation branching-speciation phyletic-gradualism punctuated-equilibrium stasis trend progression orthogenesis aristogenesis mosaic-evolution co-evolution arms-race Red-Queen-hypothesis evolutionary-stable-strategy frequency-dependent-selection density-dependent-selection r-selection K-selection bet-hedging iteroparity semelparity ploidy-level chromosome-number karyotype genome-size transposable-element gene-duplication neofunctionalization subfunctionalization pseudogene regulatory-evolution cis-regulatory trans-regulatory enhancer promoter silencer insulator transcription-factor signaling-pathway developmental-pathway gene-network epigenetic-modification DNA-methylation histone-modification chromatin-remodeling genomic-imprinting parental-effect maternal-effect paternal-effect cytoplasmic-inheritance mitochondrial-DNA chloroplast-DNA endosymbiont Wolbachia Rickettsia Spiroplasma Cardinium Arsenophonus Hamiltonella Regiella Serratia Buchnera Carsonella Portiera primary-endosymbiont secondary-endosymbiont facultative-symbiont obligate-symbiont mutualistic-symbiont parasitic-symbiont commensal-symbiont pathogen vector disease-transmission zoonosis emerging-infectious-disease biosecurity quarantine eradication control management conservation-biology restoration-ecology habitat-restoration river-restoration stream-restoration floodplain-restoration riparian-restoration wetland-restoration ecosystem-restoration passive-restoration active-restoration assisted-recovery managed-relocation assisted-colonization ex-situ-conservation in-situ-conservation protected-area national-park nature-reserve wildlife-refuge biosphere-reserve world-heritage-site ramsar-site important-bird-area key-biodiversity-area Alliance-for-Zero-Extinction EDGE-species evolutionarily-distinct globally-endangered conservation-prioritization systematic-conservation-planning Marxan Zonation conservation-opportunity conservation-cost conservation-benefit trade-off socioeconomic-factor stakeholder-engagement community-based-conservation adaptive-management monitoring-program long-term-study citizen-science public-participation environmental-education outreach science-communication knowledge-transfer capacity-building institutional-strengthening policy-development legislation regulation enforcement compliance incentive payment-for-ecosystem-services carbon-credit biodiversity-offset mitigation-banking conservation-easement land-trust private-land-conservation conservation-covenant stewardship 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ecosystem-service-valuation economic-valuation contingent-valuation choice-experiment benefit-transfer total-economic-value use-value non-use-value direct-use indirect-use option-value bequest-value existence-value willingness-to-pay willingness-to-accept cost-benefit-analysis cost-effectiveness-analysis multi-criteria-analysis decision-support uncertainty risk-assessment scenario-analysis sensitivity-analysis robustness-analysis adaptive-capacity resilience vulnerability exposure sensitivity coping-capacity transformation social-ecological-system complex-adaptive-system tipping-point regime-shift alternative-stable-state hysteresis path-dependence legacy-effect slow-variable fast-variable panarchy adaptive-cycle exploitation release reorganization foresight anticipatory-governance transformative-change sustainability-science future-earth interdisciplinary transdisciplinary problem-oriented solution-oriented action-oriented knowledge-co-production boundary-work boundary-object boundary-organization science-policy-interface science-society-interface knowledge-brokerage knowledge-exchange knowledge-integration mode-1-science mode-2-science triple-helix quadruple-helix responsible-innovation reflexive-governance adaptive-governance collaborative-governance polycentric-governance network-governance self-organization collective-action common-pool-resource tragedy-of-the-commons social-dilemma institutional-analysis institutional-design Ostrom-design-principles monitoring sanctioning conflict-resolution nested-enterprises proportionality collective-choice constitutional-choice operational-choice collective-action-theory game-theory prisoner's-dilemma chicken-game assurance-game coordination-game evolutionary-game-theory replicator-dynamics ESS Nash-equilibrium Pareto-optimality social-optimum market-failure government-failure voluntary-failure non-market-value public-good common-good club-good private-good rivalry excludability free-rider hold-up-problem moral-hazard adverse-selection information-asymmetry transaction-cost property-right common-property state-property private-property hybrid-property bundle-of-rights usufruct access withdrawal alienation tenure-security land-tenure water-tenure resource-tenure customary-tenure statutory-tenure legal-pluralism de-facto de-jure formal informal legality legitimacy recognition deviance corruption elite-capture social-capital bonding-capital bridging-capital linking-capital trust reciprocity reputation social-network social-embeddedness structural-hole brokerage closure cohesion density centrality betweenness degree eigenvector PageRank small-world scale-free random-network regular-network hierarchical-network modularity community-structure clustering assortativity homophily heterophily preferential-attachment fitness-model network-evolution network-dynamics temporal-network multilayer-network multiplex-network bipartite-network affiliation-network one-mode-network two-mode-network adjacency-matrix edge-list graph-theory vertex edge node link tie connection path cycle tree forest component giant-component percolation phase-transition criticality self-organized-criticality power-law heavy-tail Pareto-distribution lognormal-distribution exponential-distribution Poisson-distribution binomial-distribution normal-distribution Gaussian-distribution statistical-distribution probability-distribution cumulative-distribution probability-density-function cumulative-distribution-function survival-function hazard-function moment mean median mode variance standard-deviation coefficient-of-variation skewness kurtosis quantile percentile decile quartile interquartile-range range minimum maximum outlier robust-statistic parametric-test non-parametric-test hypothesis-testing null-hypothesis alternative-hypothesis significance-level p-value confidence-interval type-I-error type-II-error statistical-power effect-size sample-size sampling-design experimental-design observational-study quasi-experiment natural-experiment randomized-controlled-trial before-after-control-impact BACI impact-assessment environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint material-flow-analysis substance-flow-analysis energy-analysis exergy-analysis emergy-analysis input-output-analysis environmentally-extended-input-output social-accounting-matrix computable-general-equilibrium agent-based-model system-dynamics-model Bayesian-network structural-equation-model path-analysis factor-analysis principal-component-analysis cluster-analysis discriminant-analysis canonical-correspondence-analysis redundancy-analysis non-metric-multidimensional-scaling ordination classification regression-analysis linear-regression nonlinear-regression generalized-linear-model generalized-additive-model mixed-effects-model multilevel-model hierarchical-model Bayesian-hierarchical-model state-space-model hidden-Markov-model time-series-analysis spatial-analysis geostatistics kriging variogram spatial-autocorrelation Moran's-I Geary's-c Getis-Ord-Gi local-indicator-of-spatial-association LISA hot-spot cold-spot spatial-cluster spatial-outlier edge-effect boundary-effect modifiable-areal-unit-problem MAUP ecological-fallacy atomistic-fallacy cross-scale-inference upscaling downscaling scaling-law allometric-scaling fractal fractal-dimension self-similarity multi-fractal lacunarity roughness texture pattern process pattern-process-relationship neutral-model null-model randomization-test permutation-test bootstrap jackknife cross-validation model-averaging model-comparison information-criterion Bayesian-model-averaging ensemble-modeling consensus-model prediction forecast projection scenario narrative-scenario quantitative-scenario storyline shared-socioeconomic-pathway representative-concentration-pathway emission-scenario climate-scenario impact-scenario adaptation-scenario mitigation-scenario transformation-scenario backcasting forecasting trend-extrapolation expert-judgment Delphi-method participatory-modeling stakeholder-analysis influence-diagram conceptual-model mental-model system-archetype leverage-point intervention-point policy-instrument economic-instrument regulatory-instrument voluntary-instrument information-instrument market-based-instrument command-and-control-instrument subsidy tax charge fee levy deposit-refund tradable-permit emission-trading carbon-tax biodiversity-banking conditional-cash-transfer community-based-natural-resource-management joint-forest-management co-management adaptive-co-management transboundary-management international-regime global-governance multilateral-environmental-agreement convention treaty cartagena-protocol nagoya-protocol Aichi-target sustainable-development-goal SDG post-2020-global-biodiversity-framework Kunming-Montreal-global-biodiversity-framework 30-by-30-target nature-based-solution NbS ecosystem-based-adaptation ecosystem-based-mitigation green-infrastructure blue-infrastructure natural-infrastructure hybrid-infrastructure climate-smart-agriculture climate-smart-landscape climate-smart-development low-carbon-development green-economy circular-economy bioeconomy blue-economy sustainable-consumption sustainable-production life-cycle-thinking cradle-to-cradle industrial-ecology industrial-symbiosis urban-metabolism socio-ecological-metabolism material-intensity energy-intensity carbon-intensity decoupling absolute-decoupling relative-decoupling resource-efficiency eco-efficiency eco-innovation green-technology clean-technology environmental-technology appropriate-technology participatory-technology technology-transfer technology-diffusion innovation-system national-innovation-system sectoral-innovation-system technological-innovation-system socio-technical-system socio-technical-transition multi-level-perspective niche-innovation regime landscape transition-management strategic-niche-management transition-experiment living-lab real-world-laboratory urban-living-lab rural-living-lab sustainability-transition fundamental-change system-innovation directionality reflexivity responsiveness anticipation learning-by-doing doing-by-learning social-learning organizational-learning policy-learning transformative-capacity capacity-development institutional-learning evolutionary-governance institutional-change path-dependency lock-in carbon-lock-in technological-lock-in institutional-lock-in behavioral-lock-in cultural-lock-in unlocking destabilization phase-out divestment stranded-asset just-transition transition-pathway transition-scenario deep-decarbonization net-zero carbon-neutral climate-neutral nature-positive biodiversity-positive restoration-positive regenerative circular doughnut-economics planetary-boundary safe-operating-space environmental-ceiling social-foundation intergenerational-equity intragenerational-equity distributive-justice procedural-justice recognition-justice environmental-justice climate-justice energy-justice water-justice food-justice spatial-justice territorial-justice ecological-justice species-justice intersectionality adaptation mitigation sustainability sustainable-development well-being human-development capability-approach human-security environmental-security climate-security water-security food-security energy-security livelihood-security disaster-risk-reduction climate-change-adaptation climate-change-mitigation loss-and-damage climate-finance green-climate-fund adaptation-fund global-environment-facility biodiversity-finance nature-finance blended-finance impact-investment ESG environmental-social-governance sustainable-finance green-bond social-bond sustainability-bond blue-bond nature-bond biodiversity-credit offset inset nature-based-credit ecosystem-service-credit payment-for-results results-based-payment conditional-payment performance-based-payment outcome-based-payment environmental-service-payment PSA PES REDD REDD+AFOLU LULUCF NAMA NAMC NDC NAP NBSAP national-adaptation-plan national-biodiversity-strategy-and-action-plan nationally-determined-contribution nationally-appropriate-mitigation-action intended-nationally-determined-contribution long-term-low-emission-development-strategy net-zero-target carbon-budget remaining-carbon-budget emission-gap adaptation-gap finance-gap implementation-gap ambition-loop ratchet-mechanism global-stocktake transparency-framework enhanced-transparency-framework common-but-differentiated-responsibilities respective-capabilities climate-equity climate-litigation climate-governance climate-diplomacy climate-negotiation UNFCCC COP CMP CMA SBSTA SBI IPCC IPBES UNCCD CBD CITES Ramsar World-Heritage Man-and-the-Biosphere MAB UNESCO FAO WHO ILO UNDP UNEP UN-Habitat World-Bank IMF WTO OECD GEF GCF AF multilateral-development-bank regional-development-bank national-development-bank development-finance-institution export-credit-agency sovereign-wealth-fund pension-fund insurance-company asset-manager asset-owner corporate-investor retail-investor impact-investor philanthropic-investor development-partner bilateral-donor multilateral-donor private-donor foundation NGO CSO civil-society-organization non-governmental-organization community-based-organization indigenous-peoples-organization women's-organization youth-organization faith-based-organization research-organization think-tank university academic-institution knowledge-institution capacity-building-institution training-institution extension-service advisory-service consulting-firm engineering-firm technology-provider equipment-supplier service-provider contractor operator maintainer utility municipality local-government provincial-government national-government federal-government ministry department agency authority commission board council committee parliament congress legislature executive judiciary media journalist reporter editor publisher broadcaster platform social-media influencer celebrity activist advocate campaigner mobilizer organizer networker connector bridge-builder boundary-spanner knowledge-broker science-communicator extension-agent development-practitioner policy-maker decision-maker planner manager administrator leader champion entrepreneur innovator intrapreneur social-entrepreneur systems-entrepreneur institutional-entrepreneur policy-entrepreneur political-entrepreneur business-model business-model-innovation social-business inclusive-business base-of-pyramid fortune-at-bottom-of-pyramid shared-value creating-shared-value stakeholder-capitalism purpose-driven-business B-Corp benefit-corporation social-enterprise cooperative mutual association collective commons commoning peer-production open-source creative-commons knowledge-commons digital-commons data-commons genetic-commons seed-commons water-commons fisheries-commons forest-commons pasture-commons land-commons urban-commons atmospheric-commons global-commons common-heritage-of-mankind common-concern-of-humankind restorative-justice multispecies-justice more-than-human anthropocene capitalocene plantationocene chthulucene symbiocene good-anthropocene great-acceleration great-transformation great-decoupling great-convergence great-divergence great-fragmentation great-integration great-transition great-work long-now deep-time futures-thinking future-studies strategic-foresight scenario-planning trend-analysis weak-signal wild-card black-swan grey-rhino blue-sky green-swan systemic-change structural-change cultural-change behavioral-change individual-change collective-change social-change organizational-change technological-change economic-change political-change ecological-change planetary-change earth-system-change biodiversity-change ocean-change cryosphere-change atmospheric-change biogeochemical-change nitrogen-cycle-change phosphorus-cycle-change carbon-cycle-change water-cycle-change hydrological-change soil-change vegetation-change ecosystem-change landscape-change seascape-change biosphere-change noosphere-change technosphere-change anthroposphere-change sociosphere-change econosphere-change politosphere-change culturosphere-change semiosphere-change infosphere-change digisphere-change cybersphere-change urban-sphere-change agricultural-sphere-change industrial-sphere-change extractive-sphere-change manufacturing-sphere-change service-sphere-change financial-sphere-change knowledge-sphere-change innovation-sphere-change creativity-sphere-change culture-sphere-change identity-sphere-change meaning-sphere-change value-sphere-change purpose-sphere-change vision-sphere-change mission-sphere-change strategy-sphere-change tactic-sphere-change operation-sphere-change action-sphere-change practice-sphere-change habit-sphere-change routine-sphere-change ritual-sphere-change tradition-sphere-change custom-sphere-change convention-sphere-change norm-sphere-change rule-sphere-change law-sphere-change regulation-sphere-change standard-sphere-change protocol-sphere-change procedure-sphere-change process-sphere-change system-sphere-change structure-sphere-change function-sphere-change pattern-sphere-change dynamic-sphere-change state-sphere-change phase-sphere-change regime-sphere-change equilibrium-sphere-change stability-sphere-change instability-sphere-change chaos-sphere-change order-sphere-change complexity-sphere-change simplicity-sphere-change diversity-sphere-change uniformity-sphere-change heterogeneity-sphere-change homogeneity-sphere-change variation-sphere-change invariance-sphere-change change-sphere-change continuity-sphere-change discontinuity-sphere-change emergence-sphere-change submergence-sphere-change evolution-sphere-change devolution-sphere-change revolution-sphere-change involution-sphere-change convolution-sphere-change resolution-sphere-change dissolution-sphere-change absolution-sphere-change solution-sphere-change
Toxocampinae
Toxocampinae is a subfamily of moths within the family Erebidae, superfamily Noctuoidea. The subfamily was previously classified as the tribe Toxocampini within the former subfamily Catocalinae, but phylogenetic analysis supports its recognition as a distinct subfamily outside the Erebinae (formerly Catocalinae). Males possess a primitive form of genital claspers resembling those found in some Noctuidae subfamilies. The subfamily includes approximately 200 species across six genera, with the genus Lygephila being among the most studied.
Lepidoptera Erebidae Noctuoidea moths genitalia taxonomic-revision Palearctic genus-Lygephila genus-Autophila genus-Anumeta genus-Apopestes genus-Tathorhynchus
Trichosirocalus
rosette weevils, crown weevils
A genus of small true weevils native to the Palearctic region. Several species have been widely introduced to North America, Australia, and New Zealand as biological control agents for invasive thistles. The genus was historically treated as containing a single species, T. horridus, but integrative taxonomic studies using molecular and morphological data have revealed multiple distinct species with different host associations.
weevil biological-control thistle invasive-species-management Ceutorhynchinae rosette-feeder Ceutorhynchini Baridinae taxonomic-revision classical-biological-control Carduus Cirsium Onopordum Palearctic-native introduced-species molecular-systematics integrative-taxonomy univoltine leaf-miner agricultural-pest-control pasture-weed-management species-complex host-specificity musk-thistle plumeless-thistle Scotch-thistle Illyrian-thistle welted-thistle Italian-thistle crown-weevil rosette-weevil oviposition-behavior overwintering egg-stage larval-stage adult-stage decaying-organic-matter soil-surface non-photosynthetic-leaves green-leaves leaf-midribs petioles leaf-mining notching-damage rosette-survival flowering-stem-reduction seed-production-reduction rosette-mortality population-density-effects mark-recapture dispersal-patterns artificial-diet-rearing laboratory-culture vitamin-supplementation lyophilized-thistle-roots ergosterol casein-hydrolysate 7-dehydrocholesterol sucrose larval-survival adult-longevity oviposition female-fecundity field-cage-experiments individual-cage-experiments growth-parameter-reduction pre-release-host-specificity-testing post-release-monitoring establishment-success range-expansion forested-areas agricultural-land pasture small-farms crops disturbed-habitats meadows dense-patches rosette-size-preference spatial-distribution selective-oviposition autumn-egg-laying winter-larval-development spring-pupation late-summer-adult-emergence soil-overwintering plant-debris-overwintering rachis leaf-dissection destructive-sampling non-destructive-sampling sampling-bias life-stage-detection established-populations mitochondrial-COI nuclear-EF-1α morphological-characters host-plant-associations synonymy species-validity introduction-history release-records establishment-records Old-World-native New-World-introduction Southern-Hemisphere-introduction Australia New-Zealand North-America Virginia Southwest-Virginia Europe Spain France Germany Italy Switzerland Austria Scandinavia Denmark Norway Sweden Colonnelli-1979 Panzer-1801 Alonso-Zarazaga-&-Sánchez-Ruiz-2002 Ceutorhynchinae-vs-Baridinae subfamily-placement tribe-placement true-weevils Curculionidae Coleoptera Insecta Arthropoda Animalia 882-iNaturalist-observations GBIF-records Catalogue-of-Life-accepted NCBI-Taxonomy Bulletin-of-Entomological-Research Journal-of-Applied-Ecology Journal-of-Entomological-Science Annals-of-the-Entomological-Society-of-America Environmental-Entomology DOI-10.1017/s000748531500084x DOI-10.1046/j.1365-2664.2002.00747.x DOI-10.18474/0749-8004-22.4.324 DOI-10.18474/0749-8004-22.4.330 DOI-10.18474/0749-8004-24.4.465 DOI-10.1093/aesa/73.6.694 DOI-10.1093/ee/10.5.691
Trigonogenius
Trigonogenius is a genus of spider beetles in the family Ptinidae, established by Hagedorn in 1912. The genus contains five to six described species, including the globular spider beetle (Trigonogenius globulum). A new species was described from Peru in 2025, indicating ongoing taxonomic work and expanding knowledge of the genus's Neotropical diversity.
spider-beetles Ptinidae Ptininae stored-product-pests taxonomic-revision Neotropical
Trogoderma grassmani
Trogoderma grassmani is a species of dermestid beetle in the genus Trogoderma, described by Beal in 1954. Like other members of its genus, it belongs to a group of beetles commonly known as cabinet beetles or warehouse beetles, many of which are significant pests of stored products. The species is native to North America. Information specific to this species is limited compared to better-known congeners such as Trogoderma granarium (khapra beetle) and Trogoderma variabile (warehouse beetle).
Dermestidae Trogoderma stored-product-beetles North-American-fauna Coleoptera polyphyletic-genus taxonomic-revision
Trogossitidae
bark-gnawing beetles
Trogossitidae is a small family of beetles in the superfamily Cleroidea, commonly known as bark-gnawing beetles. The family has undergone significant taxonomic revision, with many former members reassigned to separate families (Lophocateridae, Peltidae, Protopeltidae, Rentoniidae, and Thymalidae) as of 2019. Under the modern circumscription, the family contains approximately 400 species in 25 genera, reduced from roughly 600 species in over 50 genera under previous definitions. Members are predominantly predatory and/or mycophagous, with both adults and larvae associated with wood habitats.
Coleoptera Cleroidea bark-beetles predatory-beetles wood-associated-beetles fire-associated-insects mycophagy biological-control taxonomic-revision Cretaceous-fossil-record
Tvetenia
non-biting midges
Tvetenia is a genus of non-biting midges in the family Chironomidae, subfamily Orthocladiinae. Species occur across northern and eastern Europe, extending into Russia and Central Asia. The genus was established by Kieffer in 1922. Taxonomic revisions have clarified species boundaries, including the synonymy of T. vitracies with T. tshernovskii.
Chironomidae Orthocladiinae non-biting-midges aquatic-insects freshwater Europe Asia Kieffer-1922 Tvetenia-tshernovskii Tvetenia-vitracies taxonomic-revision
Tychobythinus
Tychobythinus is a genus of small rove beetles in the subfamily Pselaphinae (Staphylinidae). The genus is widely distributed across the Palearctic region, including North Africa, southern and Central Europe, the Caucasus, Japan, far eastern Russia, and southeastern China. Several species exhibit troglobitic adaptations, including reduced pigmentation, microphthalmia, winglessness, and elongated appendages. The genus contains at least 44 species and subspecies in Italy alone, with ongoing taxonomic revisions clarifying species boundaries and synonymies.
rove-beetle pselaphine cave-beetle troglobite Palearctic endemic gypsum-karst microphthalmia aptery Staphylinidae Sicily Italy North-Africa taxonomic-revision
Tymnes oregonensis
Oregon Leaf Beetle
Tymnes oregonensis is a species of leaf beetle in the family Chrysomelidae, tribe Alticini. It is native to western North America. The species was originally described by Crotch in 1873 and has been historically treated under the genus Phyllotreta as a synonym. Available records indicate presence in the Canadian provinces of Alberta and Manitoba, with broader distribution across North America. Like other members of the Alticini, it possesses enlarged hind femora adapted for jumping.
leaf-beetle flea-beetle Alticini Chrysomelidae Coleoptera North-America western-North-America Oregon Alberta Manitoba rarely-collected taxonomic-revision jumping-beetle
Underwoodia
Underwoodia is a genus of ascomycete fungi in the family Helvellaceae, described by Charles Horton Peck in 1890 to honor mycologist Lucien Marcus Underwood. The genus contains three species with widespread distribution. Northern Hemisphere species (U. columnaris) occur in North America, while Southern Hemisphere species previously classified in Underwoodia have been reclassified to the resurrected genus Geomorium (family Geomoriaceae). Underwoodia species are ectomycorrhizal fungi associated with forest ecosystems.
ectomycorrhizal Pezizales Helvellaceae taxonomic-revision Geomorium Gondwanan-distribution
Wadotes hybridus
hacklemesh weaver
Wadotes hybridus is a species of funnel weaver spider in the family Agelenidae, found in the United States and Canada. It is among the larger spiders in its family, reaching up to 14 mm in body length at maturity. The species has been documented as prey for the spider wasp Priocnemis minorata. Taxonomic placement has shifted historically, with the genus Wadotes formerly classified in Amaurobiidae before returning to Agelenidae.
spider funnel-weaver large North-America predated-by-wasps taxonomic-revision
Wroughtonia
Wroughtonia is a genus of braconid wasps in the family Braconidae, subfamily Helconinae. The genus contains over 40 described species, with recent taxonomic work describing new species from Vietnam. As members of Braconidae, these wasps are presumed to be parasitoids, though specific host associations remain poorly documented for most species.
Braconidae Helconinae parasitoid-wasps Hymenoptera taxonomic-revision Vietnam Palearctic Oriental
Xantholininae
Xantholininae is a subfamily of rove beetles (Staphylinidae) comprising approximately 28 genera and over 100 species in North America alone, with additional diversity in the Palearctic and other regions. The group has been subject to extensive taxonomic revision due to historical confusion in genus-level classification, with multiple synonymizations required to stabilize nomenclature. Members are characterized by specific morphological features that distinguish them from other staphylinid subfamilies, though detailed biological studies remain limited compared to taxonomic treatments.
Staphylinidae rove-beetles taxonomic-revision Holarctic North-America Himalaya Coleoptera morphology genus-synonymization
Xanthopygina
Xanthopygina is a subtribe of rove beetles (Staphylinidae) containing approximately 334 species distributed across the Neotropical region from the southern United States to Argentina. Species within this subtribe are frequently encountered in tropical forests and can be abundant in various microhabitats. The subtribe has been the subject of extensive taxonomic revision due to historically poorly defined genera, particularly the genus Gastrisus, which accumulated morphologically disparate species without clear diagnostic boundaries. Recent phylogenomic studies have resolved relationships within Xanthopygina, identifying nine major lineages and clarifying generic limits.
Coleoptera Staphylinidae Neotropical rove-beetles phylogenomics taxonomy tropical-forest Staphylinini subtribe hyperdiverse taxonomic-revision Chatzimanolis Sharp-1884
Xanthopygus
Xanthopygus is a genus of rove beetles (Staphylinidae: Staphylininae) established by Kraatz in 1857. The genus belongs to the subtribe Xanthopygina and is distributed across Mexico, Central America, the West Indies, and South America. A 2022 taxonomic revision by Chatzimanolis updated distributional records, resolved nomenclatural issues including synonymies and type designations, and provided an illustrated identification key for all species. The genus is characterized by distinctive morphological features typical of the Xanthopygina, though specific diagnostic traits vary among species.
rove-beetle Staphylinidae Coleoptera taxonomic-revision Neotropical Xanthopygina