Asiloidea
Guides
Asiloidea
asiloid flies
Asiloidea is a large superfamily of true flies (Diptera: Brachycera) with cosmopolitan distribution. It comprises approximately 11 families including the well-known Asilidae (robber/assassin flies), Bombyliidae (bee flies), Therevidae (stiletto flies), Mydidae (mydas flies), and Scenopinidae (window flies). Adult morphology is characterized by antennae with no more than 4 flagellomeres, leg empodium usually setiform or absent, and wing venation featuring an elongate cell cup with vein CuA2 ending freely or meeting A1 near the wing margin. Larval synapomorphies include posterior spiracles arising dorsally from the penultimate abdominal segment and, in most families except Bombyliidae and Hilarimorphidae, a modified cranium forming a hinged metacephalic rod.
DipteraBrachyceraAsiloidearobber-fliesbee-fliesstiletto-fliesmydas-flieswindow-fliescosmopolitanarid-habitatspredatory-larvaeflower-visitorsparasitoidsbiological-controlAsilidaeBombyliidaeTherevidaeMydidaeScenopinidaeApioceridaeApsilocephalidaeApystomyiidaeEvocoidaeHilarimorphidaeMythicomyiidaeProtapioceridaephylogenysystematicsvenomouspredatorymystaxmetacephalic-roddichopticholopticJurassicCretaceoustherevoid-cladeNamib-DesertCanary-IslandsSaudi-ArabiaDominicaChileGermanyfossilcybertaxonomyopen-accessZooKeysBohart-MuseumWorld-Robber-Fly-DayHilarimorphidae
hilarimorphid flies
Hilarimorphidae is a small family of flies (Diptera) with uncertain phylogenetic placement. They have been variously associated with Acroceridae or Bombyliidae within the superfamily Asiloidea. The family is primarily Nearctic in distribution and includes both extant and fossil species.
Mydidae
Mydas flies, Mydaid flies
Mydidae, or mydas flies, are a small cosmopolitan family of large to very large flies within the superfamily Asiloidea. With approximately 471 described species, they represent one of the most distinctive dipteran groups due to their exceptional size—Gauromydas heros is the largest known fly species—and their frequent mimicry of stinging hymenopterans, particularly spider wasps. Most species inhabit arid and semiarid regions globally. The family is poorly studied, with larval biology documented for only a handful of species.
Mythicomyiidae
micro bee flies, mythicomyiids, microbombyliids
Mythicomyiidae are minute flies (0.5–5.0 mm) historically treated as a subfamily of Empididae or Bombyliidae, but elevated to family rank based on morphological and phylogenetic evidence. The family comprises 35 genera and over 450 described species, with hundreds more awaiting description. They exhibit highest diversity in Africa and are especially abundant in desert and semi-desert regions globally. The lineage dates to the Middle Jurassic, making it older than any known Bombyliidae.
Therevidae
Stiletto flies
Therevidae, commonly known as stiletto flies, is a family of brachyceran flies in the superfamily Asiloidea containing approximately 1,600 described species worldwide. The family is most diverse in arid and semiarid regions with sandy soils. Adults are small to medium-sized (2.4–18 mm) with hairy bodies and are often brightly colored, with many species exhibiting wasp or ant mimicry. Larvae are apodous, eucephalic, and predatory, living in soil and other substrates where they hunt insect larvae.