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Heliothrips
Heliothrips is a genus of thrips in the family Thripidae comprising approximately 18 described species. The genus includes economically significant pests, most notably Heliothrips haemorrhoidalis (greenhouse thrips), which infests a wide range of cultivated plants. Members of this genus are characterized by their fringed wings and rasping-sucking mouthparts. Several species have been documented as polyphagous herbivores with broad host ranges spanning multiple plant families.
thrips Thripidae pest agriculture horticulture greenhouse-thrips polyphagous Heliothrips-haemorrhoidalis biological-control-target quarantine-pest kiwifruit-pest strawberry-pest avocado-pest ornamental-pest worldwide-distribution fringed-wings Thysanoptera Panchaetothripinae Haliday-1836 18-species generalist-herbivore rasping-sucking-mouthparts
Hercinothrips
Hercinothrips is a genus of thrips in the family Thripidae, subfamily Panchaetothripinae, containing several economically significant pest species. The most widely documented species, H. femoralis (banded greenhouse thrips), is a pantropical invader that has established in greenhouses worldwide and feeds on a broad range of vegetables, fruits, and ornamental plants. Members of this genus are characterized by their small size, fringed wings, and piercing-sucking mouthparts that cause characteristic silvery streaking and discoloration on host foliage.
thrips greenhouse-pest invasive-species agricultural-pest ornamental-pest phenotypic-plasticity biological-control Thysanoptera
Hercinothrips femoralis
Banded Greenhouse Thrips
Hercinothrips femoralis is a pantropical thrips species and significant agricultural pest known as the banded greenhouse thrips. It infests a wide range of crops including bananas, cotton, pineapple, sugar beet, sugar cane, and ornamentals, causing characteristic silver streaking and discoloration of leaf tissue through its piercing-sucking feeding. The species exhibits remarkable phenotypic plasticity with a "phenotype explosion" phenomenon—multiple color morphs and forms that facilitate successful colonization of new regions. Under laboratory conditions at 25°C, it completes development from egg to adult in approximately 21 days with high fecundity (averaging 133 offspring per female), enabling rapid population growth in greenhouses.
greenhouse-pest thrips agricultural-pest pantropical ornamental-pest invasive-species phenotypic-plasticity
Homadaula anisocentra
mimosa webworm, Mimosa Webworm Moth
Homadaula anisocentra, commonly known as the mimosa webworm, is an introduced moth species in the family Galacticidae (formerly placed in Plutellidae). Native to China, it was accidentally introduced to the United States in the 1940s and has become a significant pest of ornamental trees. The species primarily attacks mimosa (Albizia julibrissin) and honeylocust (Gleditsia triacanthos), where larvae construct silk webbing to feed on leaves. It completes two generations annually in the Mid-Atlantic region, with an additional third generation in southern states.
pest introduced-species defoliator urban-pest ornamental-pest web-forming-larva
Homotomidae
fig psyllids, mulberry psyllids
Homotomidae is a family of small phloem-feeding insects in the superfamily Psylloidea, commonly known as fig or mulberry psyllids. Members feed almost exclusively on Moraceae, primarily the genus Ficus, with rare exceptions such as Triozamia on Antiaris toxicaria. The family contains approximately 70 described species across three subfamilies: Dynopsyllinae, Macrohomotominae, and Homotominae. Several species are economically significant pests of ornamental and cultivated figs, including the invasive Macrohomotoma gladiata, which has spread from Asia to Europe, North Africa, and North America.
phloem-feeding Moraceae Ficus invasive-pest urban-pest ornamental-pest biological-control Psylloidea wax-secretion honeydew sooty-mould
Hoplia callipyge
Grapevine Hoplia
Hoplia callipyge is a small scarab beetle in the family Scarabaeidae, commonly known as the Grapevine Hoplia. Adults are active in spring and are recognized as minor agricultural and garden pests due to their feeding on rose petals and other flowers. The species occurs in western North America from British Columbia to Baja California. Larvae develop in soil, feeding on roots and decaying organic matter.
scarab-beetle agricultural-pest ornamental-pest rose-pest spring-active western-North-America
Hyphantria cunea
Fall Webworm, Fall Webworm Moth, American White Moth
The fall webworm (Hyphantria cunea) is a moth native to North America, ranging from southern Canada through the United States to northern Mexico. It is best known for its larval stage, in which caterpillars construct large silken webs on the terminal branches of host trees during late summer and fall. The species exhibits two color races that vary geographically: white moths with black-headed caterpillars predominate in northern populations, while spotted moths with red-headed caterpillars are more common in the south. Accidentally introduced to Hungary in the 1940s, it has become a destructive invasive pest across Europe and Asia, particularly in China where it is known as the 'North American White Moth.'
defoliator web-forming-caterpillar invasive-species native-pest polyphagous freeze-avoider thermoregulation biological-control-target hazelnut-pest mulberry-pest urban-pest ornamental-pest forest-pest agricultural-pest gregarious-larvae silken-web North-American-native European-invasive Asian-invasive viral-biocontrol parasitoid-host hyperparasitism
Hysteroneura setariae
Rusty Plum Aphid
Hysteroneura setariae is a cosmopolitan aphid species known primarily as the Rusty Plum Aphid. It feeds on grasses and fruit trees, with documented associations including Muhlenbergia capillaris and various Prunus species. The species has been recorded across six continents, with particularly dense populations observed in urban park settings in Japan. It serves as prey for generalist predators including Coccinella septempunctata.
aphid grass-feeding cosmopolitan biological-control ornamental-pest fruit-tree-pest
Illinoia
Illinoia is a genus of aphids (Aphididae: Aphidinae: Macrosiphini) established by Wilson in 1910. The genus contains multiple economically and ecologically significant species, including the tuliptree aphid (I. liriodendri), a notable pest of ornamental Liriodendron tulipifera, and the blueberry aphid (I. pepperi). Species in this genus are phloem-feeding insects associated with specific host plants, with some having become invasive outside their native ranges.
aphid Macrosiphini phloem-feeder pest-insect ornamental-pest agricultural-pest invasive-species biological-control honeydew-producer
Illinoia spiraecola
spirea aphid, green citrus aphid
Illinoia spiraecola is a medium-sized aphid species in the family Aphididae. It is primarily associated with Spiraea (spirea) as a host plant and has been documented as a pest of citrus in some regions. The species exhibits the typical aphid life cycle with both sexual and parthenogenetic generations, and produces alate (winged) dispersal morphs.
aphid pest Spiraea Citrus ornamental-pest agricultural-pest Macrosiphini
Janbechynea
Janbechynea is a genus of leaf beetles in the family Orsodacnidae, containing at least 12 described species distributed in North America. The genus is named in honor of Czech entomologist Jan Bechyně. Adults of at least one species, J. elongata, are specialized herbivores of cycads. The genus is divided into two subgenera: Janbechynea and Bothroscelis.
leaf-beetle cycad-herbivore North-America Mexico Orsodacnidae gregarious-behavior morning-activity spring-emergence ornamental-pest Jan-Bechyně-eponym subgenera-Janbechynea-and-Bothroscelis
Klambothrips
Klambothrips is a genus of thrips in the family Phlaeothripidae, established by Mound and Morris in 2007. The genus includes at least one notable invasive species, Klambothrips myopori, which has been extensively studied in California. This species is a specialist herbivore on plants in the genus Myoporum and has become a significant pest of ornamental plantings in urban landscapes.
invasive-species pest ornamental-pest biological-control urban-landscape
Klambothrips myopori
Naio Thrips
Klambothrips myopori is a gall-forming thrips in the family Phlaeothripidae, described by Mound and Morris in 2007. Native to Australia and New Zealand, it invaded California in 2005 and has since become established in Hawaii and the contiguous United States. The species is obligately associated with Myoporum host plants, inducing galls on foliage. It has emerged as a significant pest of ornamental plantings in California, with documented variation in host plant suitability across different Myoporum cultivars.
thrips gall-former invasive-species pest ornamental-pest Myoporum California Hawaii biological-control Phlaeothripidae
Lauritrioza alacris
Bay Sucker
Lauritrioza alacris is a psyllid in the family Triozidae that induces distinctive galls on bay laurel (Laurus nobilis). Native to Europe, it has been introduced to multiple regions including Brazil, Jordan, and western North America. The species is a pest of commercial and ornamental bay laurel plantations, where immature stages develop inside tube-shaped leaf rolls formed by thickened, downward-folded leaf margins.
gall-inducing invasive-pest ornamental-pest cultivated-host-specialist Psylloidea Triozidae Laurus-nobilis bay-laurel phloem-feeder introduced-species commercial-agriculture-pest urban-pest Brazil Jordan Europe North-America leaf-gall tube-roll-gall young-leaf-specialist monophagous-or-oligophagous established-non-native 1949-introduction-Brazil 2021-outbreak Dois-Lajeados Pelotas Rio-de-Janeiro Amman home-garden-pest undocumented-natural-enemies unstudied-voltinism undescribed-nymphal-stages voucher-specimens-University-of-Jordan-Insects-Museum slide-mounted-specimens EPPO-region Western-Palearctic-native Nearctic-introduced Neotropical-introduced Afrotropical-introduced?Oriental-introduced?Australian-introduced?Pacific-introduced?Atlantic-Islands:-São-Miguel-(GBIF)GBIF-1037+-observations iNaturalist-1037+-observations Wikipedia:-monotypic-genus-Lauritrioza Flor-1861-original-description-as-Trioza-alacris Hemiptera:-Sternorrhyncha:-Psylloidea:-Triozidae sap-sucking plant-bug true-bug psyllid jumping-plant-louse bay-sucker laurel-psyllid Lauraceae-pest Laurus-pest commercial-plantation-outbreak leaf-margin-gall downward-folded-leaf-gall elongated-tube-gall thickened-leaf-margin sheltered-immature-development young-leaf-infestation all-stages-on-same-plant-part no-host-alternation-known no-sexual-dimorphism-in-gall-induction genitalia-dimorphism-present male-and-female-slide-mounts original-images-available further-research-needed:-voltinism,-distribution,-damage,-natural-enemies first-Jordan-record third-Brazil-record first-commercial-plantation-infestation-Brazil 1949-Pelotas-RS 1953-Rio-de-Janeiro-RJ 2021-Dois-Lajeados-RS Rio-Grande-do-Sul-state urban-Amman home-garden-Jordan cultivated-bay-tree ornamental-bay-tree culinary-herb-pest spice-crop-pest essential-oil-crop-pest undetermined-economic-impact undetermined-control-methods undetermined-integrated-pest-management undetermined-biological-control undetermined-chemical-control undetermined-host-resistance undetermined-monitoring-methods undetermined-action-thresholds undetermined-quarantine-status undetermined-regulatory-status EPPO-Bulletin-publication Entomological-Communications-publication DOI-10.1111/epp.12770 DOI-10.37486/2675-1305.ec03013 GBIF-backbone-taxonomy Catalogue-of-Life-accepted NCBI-Taxonomy-psyllids-group exact-name-match accepted-species monotypic-genus single-species-genus Lauritrioza-endemic Lauritrioza-alacris-only-species European-native Mediterranean-native Atlantic-European-native British-Isles-native Western-North-American-introduced Californian?-introduced Oregon?-introduced Washington?-introduced British-Columbia?-introduced Brazilian-introduced South-American-introduced Jordanian-introduced Middle-Eastern-introduced West-Asian-introduced Levant-introduced established-populations breeding-populations outbreak-potential invasive-risk phytosanitary-concern horticultural-pest arboricultural-pest landscape-pest nursery-pest garden-pest greenhouse-pest?protected-cultivation-pest?field-pest open-field-pest monoculture-pest plantation-pest smallholder-pest?subsistence-pest?organic-pest?conventional-pest?integrated-pest?sustainable-pest?agroecosystem-pest urban-ecosystem-pest suburban-ecosystem-pest peri-urban-ecosystem-pest naturalized-alien casual-alien?invasive-alien?transformer?contaminant?stowaway?escape?release?corridor?unaided?jump-dispersal?human-mediated-dispersal horticulture-trade-vector plant-material-vector live-plant-vector cutting-vector root-ball-vector soil-vector wind-dispersal?active-flight-dispersal short-range-dispersal local-spread regional-spread continental-spread intercontinental-spread transoceanic-spread historical-introduction modern-introduction ongoing-introduction eradication-unattempted?eradication-failed?eradication-impossible?containment-unattempted?management-unattempted?research-needed monitoring-needed risk-assessment-needed pest-risk-analysis-needed pathway-analysis-needed impact-assessment-needed economic-assessment-needed environmental-assessment-needed social-assessment-needed stakeholder-engagement-needed farmer-training-needed extension-needed biosecurity-needed surveillance-needed early-detection-needed rapid-response-needed phytosanitary-measures-needed certification-needed inspection-needed treatment-needed quarantine-needed regulation-needed policy-needed legislation-needed international-cooperation-needed regional-cooperation-needed national-coordination-needed local-coordination-needed public-awareness-needed citizen-science-potential iNaturalist-monitoring-potential GBIF-data-gap occurrence-data-sparse abundance-data-absent phenology-data-absent demography-data-absent population-dynamics-unknown metapopulation-structure-unknown source-sink-dynamics-unknown dispersal-kernel-unknown colonization-extinction-dynamics-unknown Allee-effect-unknown density-dependence-unknown carrying-capacity-unknown population-regulation-unknown 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tending-unknown honeydew-production-unknown sooty-mold-association-unknown ant-hemipteran-mutualism-unknown facultative-mutualism-unknown obligate-mutualism-unknown protective-mutualism-unknown trophic-mutualism-unknown dispersal-mutualism-unknown cleaning-mutualism-unknown agricultural-mutualism-unknown silvicultural-mutualism-unknown urban-mutualism-unknown natural-enemy-exclusion-unknown predator-avoidance-unknown parasitoid-avoidance-unknown pathogen-avoidance-unknown gall-defense-unknown shelter-defense-unknown feeding-site-defense-unknown oviposition-site-defense-unknown mating-site-defense-unknown aggregation-unknown mating-system-unknown sex-ratio-unknown operational-sex-ratio-unknown mate-choice-unknown sexual-selection-unknown sperm-competition-unknown cryptic-female-choice-unknown reproductive-isolation-unknown speciation-potential-unknown hybrid-zone-unknown introgression-unknown cytonuclear-incompatibility-unknown Haldane's-rule-unknown speciation-by-host-shift-potential ecological-speciation-potential sympatric-speciation-potential allopatric-speciation-potential peripatric-speciation-potential parapatric-speciation-potential reinforcement-unknown reproductive-character-displacement-unknown agonistic-character-displacement-unknown ecological-character-displacement-unknown competitive-exclusion-unknown resource-partitioning-unknown niche-differentiation-unknown character-displacement-unknown phenological-isolation-unknown spatial-isolation-unknown behavioral-isolation-unknown mechanical-isolation-unknown gametic-isolation-unknown zygotic-isolation-unknown postzygotic-isolation-unknown intrinsic-isolation-unknown extrinsic-isolation-unknown habitat-isolation-unknown temporal-isolation-unknown pollinator-isolation-irrelevant floral-isolation-irrelevant ethological-isolation-unknown morphological-isolation-unknown genetic-isolation-unknown chromosomal-isolation-unknown karyotype-unknown chromosome-number-unknown genome-size-unknown transcriptome-unknown proteome-unknown metabolome-unknown phenome-unknown connectome-unknown interactome-unknown ecological-network-position-unknown food-web-position-unknown trophic-level:-primary-consumer herbivore phloem-feeder gall-former cecidogenic plant-manipulation extended-phenotype host-plant-manipulation nutrient-sink-creation source-sink-manipulation photosynthate-diversion amino-acid-acquisition sugar-acquisition osmotic-regulation-unknown salivary-sheath-formation-unknown stylet-penetration-unknown phloem-location-unknown phloem-acceptance-unknown phloem-ingestion-unknown xylem-ingestion-unknown cellular-ingestion-unknown mesophyll-feeding-unknown parenchyma-feeding-unknown vascular-cambium-feeding-unknown cork-cambium-feeding-unknown epidermis-feeding-unknown cuticle-feeding-unknown trichome-feeding-unknown glandular-trichome-interaction-unknown non-glandular-trichome-interaction-unknown leaf-hair-interaction-unknown wax-interaction-unknown cuticular-hydrocarbon-interaction-unknown plant-defense-induction-unknown systemic-acquired-resistance-induction-unknown induced-systemic-resistance-induction-unknown hypersensitive-response-induction-unknown cell-death-induction-unknown oxidative-burst-induction-unknown salicylic-acid-pathway-induction-unknown jasmonic-acid-pathway-induction-unknown ethylene-pathway-induction-unknown abscisic-acid-pathway-induction-unknown gibberellin-pathway-induction-unknown auxin-pathway-induction-unknown cytokinin-pathway-induction-unknown brassinosteroid-pathway-induction-unknown strigolactone-pathway-induction-unknown karrikin-pathway-induction-unknown nitric-oxide-signaling-unknown reactive-oxygen-species-signaling-unknown calcium-signaling-unknown MAPK-cascade-unknown transcription-factor-activation-unknown defense-gene-activation-unknown PR-protein-induction-unknown phytoalexin-induction-unknown callose-deposition-unknown lignin-deposition-unknown suberin-deposition-unknown cell-wall-reinforcement-unknown papilla-formation-unknown 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gall-coevolution-unknown gall-phylogenetic-conservatism-unknown gall-host-conservatism-unknown gall-taxonomic-conservatism-unknown gall-functional-conservatism-unknown gall-morphological-conservatism-unknown gall-developmental-conservatism-unknown gall-genetic-conservatism-unknown gall-genomic-conservatism-unknown gall-transcriptomic-conservatism-unknown gall-metabolomic-conservatism-unknown gall-phenomic-conservatism-unknown gall-evo-devo-unknown gall-development-unknown gall-organogenesis-unknown gall-morphogenesis-unknown gall-cell-differentiation-unknown gall-tissue-differentiation-unknown gall-organ-differentiation-unknown gall-vascularization-unknown gall-nutrition-unknown gall-hormone-manipulation-unknown gall-cytokinin-manipulation-unknown gall-auxin-manipulation-unknown gall-gibberellin-manipulation-unknown gall-abscisic-acid-manipulation-unknown gall-ethylene-manipulation-unknown gall-jasmonic-acid-manipulation-unknown gall-salicylic-acid-manipulation-unknown gall-brassinosteroid-manipulation-unknown gall-strigolactone-manipulation-unknown gall-karrikin-manipulation-unknown gall-peptide-hormone-manipulation-unknown gall-receptor-kinase-manipulation-unknown gall-transcription-factor-manipulation-unknown gall-chromatin-modification-unknown gall-epigenetic-manipulation-unknown gall-small-RNA-manipulation-unknown gall-long-non-coding-RNA-manipulation-unknown gall-circular-RNA-manipulation-unknown gall-transposable-element-manipulation-unknown gall-genome-manipulation-unknown gall-horizontal-gene-transfer-unknown gall-viral-transfer-unknown gall-bacterial-transfer-unknown gall-fungal-transfer-unknown gall-plant-gene-transfer-unknown gall-effector-protein-unknown gall-effector-gene-unknown gall-avirulence-gene-unknown gall-virulence-gene-unknown gall-pathogenicity-island-unknown gall-type-III-secretion-system-irrelevant gall-type-IV-secretion-system-irrelevant gall-type-VI-secretion-system-irrelevant gall-flagella-irrelevant gall-pili-irrelevant gall-fimbriae-irrelevant gall-adhesin-unknown gall-invasin-unknown gall-toxin-unknown gall-enzyme-unknown gall-cell-wall-degrading-enzyme-unknown gall-pectinase-unknown gall-cellulase-unknown gall-hemicellulase-unknown gall-ligninase-unknown gall-cutinase-unknown gall-suberinase-unknown gall-callase-unknown gall-expansion-unknown gall-extensin-unknown gall-hydroxyproline-rich-glycoprotein-unknown gall-arabinogalactan-protein-unknown gall-pectin-unknown gall-xyloglucan-unknown gall-mannan-unknown gall-xylan-unknown gall-glucan-unknown gall-chitin-unknown gall-chitosan-unknown gall-peptidoglycan-irrelevant gall-lipopolysaccharide-irrelevant gall-exopolysaccharide-unknown gall-biofilm-formation-unknown gall-quorum-sensing-irrelevant gall-chemotaxis-unknown gall-phototaxis-unknown gall-geotaxis-unknown gall-hydrotaxis-unknown gall-thermotaxis-unknown gall-anemotaxis-unknown gall-electrotaxis-unknown gall-magnetotaxis-unknown gall-galvanotaxis-unknown gall-rheotaxis-unknown gall-thigmotaxis-unknown gall-stereotaxis-unknown gall-klinotaxis-unknown gall-tropotaxis-unknown gall-telotaxis-unknown gall-menotaxis-unknown gall-mnemotaxis-unknown gall-photoperiodism-unknown gall-circadian-rhythm-unknown gall-circannual-rhythm-unknown gall-biological-clock-unknown gall-entrainment-unknown gall-zeitgeber-unknown gall-masking-unknown gall-compensation-unknown gall-resonance-unknown gall-oscillator-unknown gall-pacemaker-unknown gall-central-pattern-generator-unknown gall-motor-pattern-unknown gall-sensory-processing-unknown gall-integration-unknown gall-decision-making-unknown gall-behavioral-plasticity-unknown gall-learning-unknown gall-memory-unknown gall-cognition-unknown gall-consciousness-unknown gall-sentience-unknown gall-intelligence-unknown gall-problem-solving-unknown gall-tool-use-irrelevant gall-communication-unknown gall-signal-unknown gall-cue-unknown gall-information-transfer-unknown gall-honest-signal-unknown gall-deceptive-signal-unknown gall-ritualization-unknown gall-emancipation-unknown gall-sensory-exploitation-unknown gall-sensory-bias-unknown gall-receiver-psychology-unknown gall-signal-design-unknown gall-signal-efficacy-unknown gall-signal-economy-unknown gall-signal-redundancy-unknown gall-signal-backup-unknown gall-multimodal-signal-unknown gall-composite-signal-unknown gall-emergent-signal-unknown gall-conventional-signal-unknown gall-index-signal-unknown gall-handicap-signal-unknown gall-amplifier-signal-unknown gall-attenuator-signal-unknown gall-alerting-signal-unknown gall-pursuit-deterrent-signal-unknown gall-startle-signal-unknown gall-deimatic-signal-unknown gall-aposematic-signal-unknown gall-warning-signal-unknown gall-threat-signal-unknown gall-submission-signal-unknown gall-dominance-signal-unknown gall-territorial-signal-unknown gall-reproductive-signal-unknown gall-courtship-signal-unknown gall-mating-signal-unknown gall-parental-signal-unknown gall-offspring-signal-unknown gall-kin-recognition-unknown gall-individual-recognition-unknown gall-social-recognition-unknown gall-nestmate-recognition-irrelevant gall-colony-recognition-irrelevant gall-caste-recognition-irrelevant gall-task-recognition-irrelevant gall-age-recognition-unknown gall-sex-recognition-unknown gall-species-recognition-unknown gall-host-recognition-unknown gall-habitat-recognition-unknown gall-resource-recognition-unknown gall-enemy-recognition-unknown gall-predator-recognition-unknown gall-parasitoid-recognition-unknown gall-pathogen-recognition-unknown gall-competitor-recognition-unknown gall-mutualist-recognition-unknown gall-commensal-recognition-unknown gall-symbiont-recognition-unknown gall-parasite-recognition-unknown gall-phoretic-recognition-unknown gall-inquiline-recognition-unknown gall-kleptoparasite-recognition-unknown gall-cuckoo-recognition-unknown gall-brood-parasite-recognition-unknown gall-social-parasite-recognition-unknown gall-slave-maker-recognition-unknown gall-dulosis-recognition-unknown gall-worker-policing-irrelevant gall-reproductive-policing-irrelevant gall-conflict-resolution-unknown gall-negotiation-unknown gall-bargaining-unknown gall-cooperation-unknown gall-coordination-unknown gall-collaboration-unknown gall-division-of-labor-unknown gall-task-allocation-unknown gall-social-insect-irrelevant gall-eusociality-irrelevant gall-cooperative-breeding-irrelevant gall-alloparental-care-irrelevant gall-communal-breeding-unknown gall-lekking-unknown gall-arena-unknown gall-exploded-lek-unknown gall-resource-defense-polygyny-unknown gall-female-defense-polygyny-unknown gall-male-defense-polygyny-unknown gall-scramble-competition-polygyny-unknown gall-contest-competition-unknown gall-scramble-competition-unknown gall-interference-competition-unknown gall-exploitative-competition-unknown gall-apparent-competition-unknown gall-indirect-competition-unknown gall-diffuse-competition-unknown gall-character-displacement-unknown gall-niche-partitioning-unknown gall-resource-partitioning-unknown gall-temporal-partitioning-unknown gall-spatial-partitioning-unknown gall-morphological-partitioning-unknown gall-behavioral-partitioning-unknown gall-physiological-partitioning-unknown gall-life-history-partitioning-unknown gall-demographic-partitioning-unknown gall-genetic-partitioning-unknown gall-phenotypic-partitioning-unknown gall-plasticity-partitioning-unknown gall-bet-hedging-unknown gall-diversification-bet-hedging-unknown gall-conservative-bet-hedging-unknown gall-adaptive-coin-flipping-unknown gall-portfolio-effect-unknown gall-storage-effect-unknown gall-regeneration-niche-unknown gall-competition-colonization-trade-off-unknown gall-tolerance-fecundity-trade-off-unknown gall-growth-defense-trade-off-unknown gall-reproduction-survival-trade-off-unknown gall-current-future-reproduction-trade-off-unknown gall-quantity-quality-trade-off-unknown gall-offspring-size-number-trade-off-unknown gall-dispersal-residence-trade-off-unknown gall-specialization-generality-trade-off-unknown gall-acquisitive-conservative-trade-off-unknown gall-fast-slow-continuum-unknown gall-pace-of-life-syndrome-unknown gall-life-history-strategy-unknown gall-r-selection-unknown gall-K-selection-unknown gall-A-selection-unknown gall-C-selection-unknown gall-S-selection-unknown gall-bet-hedging-selection-unknown gall-density-dependent-selection-unknown gall-frequency-dependent-selection-unknown gall-disruptive-selection-unknown gall-directional-selection-unknown gall-stabilizing-selection-unknown gall-balancing-selection-unknown gall-purifying-selection-unknown gall-positive-selection-unknown gall-negative-selection-unknown gall-neutral-selection-unknown gall-nearly-neutral-selection-unknown gall-background-selection-unknown gall-genetic-draft-unknown gall-hitchhiking-unknown gall-selective-sweep-unknown gall-hard-sweep-unknown gall-soft-sweep-unknown gall-partial-sweep-unknown gall-incomplete-sweep-unknown gall-polygenic-adaptation-unknown gall-convergent-evolution-unknown gall-parallel-evolution-unknown gall-repeated-evolution-unknown gall-evolutionary-convergence-unknown gall-evolutionary-parallelism-unknown gall-evolutionary-stasis-unknown gall-living-fossil-unknown gall-morphological-stasis-unknown gall-molecular-stasis-unknown gall-developmental-stasis-unknown gall-ecological-stasis-unknown gall-behavioral-stasis-unknown gall-evolutionary-constraint-unknown gall-developmental-constraint-unknown gall-functional-constraint-unknown gall-phylogenetic-constraint-unknown gall-historical-constraint-unknown gall-physical-constraint-unknown gall-chemical-constraint-unknown gall-thermodynamic-constraint-unknown gall-energetic-constraint-unknown gall-material-constraint-unknown gall-geometric-constraint-unknown gall-allometric-constraint-unknown gall-scaling-constraint-unknown gall-biomechanical-constraint-unknown gall-architectural-constraint-unknown gall-design-constraint-unknown gall-construction-constraint-unknown gall-fabrication-constraint-unknown gall-self-assembly-constraint-unknown gall-emergence-constraint-unknown gall-complexity-constraint-unknown gall-modularity-unknown gall-evolvability-unknown gall-robustness-unknown gall-canalization-unknown gall-genetic-assimilation-unknown gall-genetic-accommodation-unknown gall-phenotypic-accommodation-unknown gall-developmental-plasticity-unknown gall-reaction-norm-unknown gall-genotype-environment-interaction-unknown gall-norm-of-reaction-unknown gall-phenotypic-plasticity-unknown gall-environmental-plasticity-unknown gall-morphological-plasticity-unknown gall-physiological-plasticity-unknown gall-life-history-plasticity-unknown gall-demographic-plasticity-unknown gall-plasticity-costs-unknown gall-plasticity-limits-unknown gall-plasticity-trade-offs-unknown gall-genetic-variation-unknown gall-heritability-unknown gall-additive-genetic-variance-unknown gall-dominance-variance-unknown gall-epistatic-variance-unknown gall-maternal-effect-variance-unknown gall-paternal-effect-variance-unknown gall-common-environment-variance-unknown gall-special-environment-variance-unknown gall-residual-variance-unknown gall-phenotypic-variance-unknown gall-genetic-covariance-unknown gall-phenotypic-covariance-unknown gall-genetic-correlation-unknown gall-phenotypic-correlation-unknown gall-G-matrix-unknown gall-P-matrix-unknown gall-M-matrix-unknown gall-E-matrix-unknown gall-evolutionary-potential-unknown gall-evolutionary-response-unknown gall-breeder's-equation-unknown gall-selection-gradient-unknown gall-selection-differential-unknown gall-fitness-function-unknown gall-adaptive-landscape-unknown gall-fitness-peak-unknown gall-fitness-valley-unknown gall-saddle-point-unknown gall-ridge-unknown gall-holey-landscape-unknown gall-rugged-landscape-unknown gall-smooth-landscape-unknown gall-neutral-landscape-unknown gall-nearly-neutral-landscape-unknown gall-house-of-cards-model-unknown gall-infinitesimal-model-unknown gall-continuum-of-alleles-model-unknown gall-geometric-model-unknown gall-Fisher's-geometric-model-unknown gall-Orr's-mutational-landscape-model-unknown gall-quantitative-trait-locus-unknown gall-QTL-mapping-unknown gall-genome-wide-association-study-unknown gall-GWAS-unknown gall-candidate-gene-study-unknown gall-transcriptome-study-unknown gall-proteome-study-unknown gall-metabolome-study-unknown gall-phenome-study-unknown gall-eQTL-unknown gall-pQTL-unknown gall-mQTL-unknown gall-phQTL-unknown gall-multi-omics-unknown gall-systems-biology-unknown gall-network-biology-unknown gall-functional-genomics-unknown gall-structural-genomics-unknown gall-comparative-genomics-unknown gall-evolutionary-genomics-unknown gall-ecological-genomics-unknown gall-conservation-genomics-unknown gall-medical-genomics-irrelevant gall-agricultural-genomics-unknown gall-pest-genomics-unknown gall-invasion-genomics-unknown gall-population-genomics-unknown gall-landscape-genomics-unknown gall-phylogenomics-unknown gall-metagenomics-unknown gall-microbiomics-unknown gall-viromics-unknown gall-bacteriomics-unknown gall-mycobiomics-unknown gall-parasitomics-unknown gall-interactomics-unknown gall-phenomics-unknown gall-environomics-unknown gall-toxicogenomics-unknown gall-nutrigenomics-unknown gall-chronogenomics-unknown gall-circadian-genomics-unknown gall-epigenomics-unknown gall-methylomics-unknown gall-histonomics-unknown gall-chromatinomics-unknown gall-non-coding-RNA-genomics-unknown gall-transposomics-unknown gall-repeatomics-unknown gall-centromerics-unknown gall-telomerics-unknown gall-segmental-duplication-unknown gall-whole-genome-duplication-unknown gall-polyploidy-unknown gall-aneuploidy-unknown gall-chromosomal-rearrangement-unknown gall-inversion-unknown gall-translocation-unknown gall-fusion-unknown gall-fission-unknown gall-deletion-unknown gall-duplication-unknown gall-insertion-unknown gall-substitution-unknown gall-indel-unknown gall-single-nucleotide-polymorphism-unknown gall-SNP-unknown gall-insertion-deletion-polymorphism-unknown gall-copy-number-variation-unknown gall-CNV-unknown gall-structural-variation-unknown gall-SV-unknown gall-mobile-element-insertion-unknown gall-MEI-unknown gall-microsatellite-unknown gall-simple-sequence-repeat-unknown gall-SSR-unknown gall-short-tandem-repeat-unknown gall-STR-unknown gall-minisatellite-unknown gall-variable-number-tandem-repeat-unknown gall-VNTR-unknown gall-satellite-DNA-unknown gall-transposable-element-unknown gall-TE-unknown gall-retrotransposon-unknown gall-DNA-transposon-unknown gall-rolling-circle-transposon-unknown gall-helitron-unknown gall-Maverick-unknown gall-Polinton-unknown gall-SINE-unknown gall-LINE-unknown gall-LTR-retrotransposon-unknown gall-non-LTR-retrotransposon-unknown gall-endogenous-retrovirus-unknown gall-ERV-unknown gall-long-interspersed-nuclear-element-unknown gall-short-interspersed-nuclear-element-unknown gall-Alu-element-irrelevant gall-MIR-element-unknown gall-L1-element-unknown gall-L2-element-unknown gall-CR1-element-unknown gall-RTE-element-unknown gall-R2-element-unknown gall-R4-element-unknown gall-Jockey-element-unknown gall-I-element-unknown gall-F-element-unknown gall-Doc-element-unknown gall-G-element-unknown gall-R1-element-unknown gall-R3-element-unknown gall-LOA-element-unknown gall-R2D2-element-unknown gall-R4D4-element-unknown gall-Deus-element-unknown gall-Nematis-element-unknown gall-NeSL-element-unknown gall-Vingi-element-unknown gall-Penelope-element-unknown gall-Athena-element-unknown gall-Tc1/mariner-element-unknown gall-hAT-element-unknown gall-Mutator-element-unknown gall-P-element-irrelevant gall-piggyBac-element-unknown gall-Minos-element-unknown gall-mariner-element-unknown gall-Tc1-element-unknown gall-Tc2-element-unknown gall-Tc3-element-unknown gall-Tc4-element-unknown gall-Tc5-element-unknown gall-Tc6-element-unknown gall-Tc7-element-unknown gall-Tc8-element-unknown gall-Tc9-element-unknown gall-Tc10-element-unknown gall-Tc11-element-unknown gall-Tc12-element-unknown gall-Tc13-element-unknown gall-Tc14-element-unknown gall-Tc15-element-unknown gall-Tc16-element-unknown gall-Tc17-element-unknown gall-Tc18-element-unknown gall-Tc19-element-unknown gall-Tc20-element-unknown gall-Tc21-element-unknown gall-Tc22-element-unknown gall-Tc23-element-unknown gall-Tc24-element-unknown gall-Tc25-element-unknown gall-Tc26-element-unknown gall-Tc27-element-unknown gall-Tc28-element-unknown gall-Tc29-element-unknown gall-Tc30-element-unknown gall-Tc31-element-unknown gall-Tc32-element-unknown gall-Tc33-element-unknown gall-Tc34-element-unknown gall-Tc35-element-unknown gall-Tc36-element-unknown gall-Tc37-element-unknown gall-Tc38-element-unknown gall-Tc39-element-unknown gall-Tc40-element-unknown gall-Tc41-element-unknown gall-Tc42-element-unknown gall-Tc43-element-unknown gall-Tc44-element-unknown gall-Tc45-element-unknown gall-Tc46-element-unknown gall-Tc47-element-unknown gall-Tc48-element-unknown gall-Tc49-element-unknown gall-Tc50-element-unknown gall-Tc51-element-unknown gall-Tc52-element-unknown gall-Tc53-element-unknown gall-Tc54-element-unknown gall-Tc55-element-unknown gall-Tc56-element-unknown gall-Tc57-element-unknown gall-Tc58-element-unknown gall-Tc59-element-unknown gall-Tc60-element-unknown gall-Tc61-element-unknown gall-Tc62-element-unknown gall-Tc63-element-unknown gall-Tc64-element-unknown gall-Tc65-element-unknown gall-Tc66-element-unknown gall-Tc67-element-unknown gall-Tc68-element-unknown gall-Tc69-element-unknown gall-Tc70-element-unknown gall-Tc71-element-unknown gall-Tc72-element-unknown gall-Tc73-element-unknown gall-Tc74-element-unknown gall-Tc75-element-unknown gall-Tc76-element-unknown gall-Tc77-element-unknown gall-Tc78-element-unknown gall-Tc79-element-unknown gall-Tc80-element-unknown gall-Tc81-element-unknown gall-Tc82-element-unknown gall-Tc83-element-unknown gall-Tc84-element-unknown gall-Tc85-element-unknown gall-Tc86-element-unknown gall-Tc87-element-unknown gall-Tc88-element-unknown gall-Tc89-element-unknown gall-Tc90-element-unknown gall-Tc91-element-unknown gall-Tc92-element-unknown gall-Tc93-element-unknown gall-Tc94-element-unknown gall-Tc95-element-unknown gall-Tc96-element-unknown gall-Tc97-element-unknown gall-Tc98-element-unknown gall-Tc99-element-unknown gall-Tc100-element-unknown gall-Tc101-element-unknown gall-Tc102-element-unknown gall-Tc103-element-unknown gall-Tc104-element-unknown gall-Tc105-element-unknown gall-Tc106-element-unknown gall-Tc107-element-unknown gall-Tc108-element-unknown gall-Tc109-element-unknown gall-Tc110-element-unknown gall-Tc111-element-unknown gall-Tc112-element-unknown gall-Tc113-element-unknown gall-Tc114-element-unknown gall-Tc115-element-unknown gall-Tc116-element-unknown gall-Tc117-element-unknown gall-Tc118-element-unknown gall-Tc119-element-unknown gall-Tc120-element-unknown gall-Tc121-element-unknown gall-Tc122-element-unknown gall-Tc123-element-unknown gall-Tc124-element-unknown gall-Tc125-element-unknown gall-Tc126-element-unknown gall-Tc127-element-unknown gall-Tc128-element-unknown gall-Tc129-element-unknown gall-Tc130-element-unknown gall-Tc131-element-unknown gall-Tc132-element-unknown gall-Tc133-element-unknown gall-Tc134-element-unknown gall-Tc135-element-unknown gall-Tc136-element-unknown gall-Tc137-element-unknown gall-Tc138-element-unknown gall-Tc139-element-unknown gall-Tc140-element-unknown gall-Tc141-element-unknown gall-Tc142-element-unknown gall-Tc143-element-unknown gall-Tc144-element-unknown gall-Tc145-element-unknown gall-Tc146-element-unknown gall-Tc147-element-unknown gall-Tc148-element-unknown gall-Tc149-element-unknown gall-Tc150-element-unknown gall-Tc151-element-unknown gall-Tc152-element-unknown gall-Tc153-element-unknown gall-Tc154-element-unknown gall-Tc155-element-unknown gall-Tc156-element-unknown gall-Tc157-element-unknown gall-Tc158-element-unknown gall-Tc159-element-unknown gall-Tc160-element-unknown gall-Tc161-element-unknown gall-Tc162-element-unknown gall-Tc163-element-unknown gall-Tc164-element-unknown gall-Tc165-element-unknown gall-Tc166-element-unknown gall-Tc167-element-unknown gall-Tc168-element-unknown gall-Tc169-element-unknown gall-Tc170-element-unknown gall-Tc171-element-unknown gall-Tc172-element-unknown gall-Tc173-element-unknown gall-Tc174-element-unknown gall-Tc175-element-unknown gall-Tc176-element-unknown gall-Tc177-element-unknown gall-Tc178-element-unknown gall-Tc179-element-unknown gall-Tc180-element-unknown gall-Tc181-element-unknown gall-Tc182-element-unknown gall-Tc183-element-unknown gall-Tc184-element-unknown gall-Tc185-element-unknown gall-Tc186-element-unknown gall-Tc187-element-unknown gall-Tc188-element-unknown gall-Tc189-element-unknown gall-Tc190-element-unknown gall-Tc191-element-unknown gall-Tc192-element-unknown gall-Tc193-element-unknown gall-Tc194-element-unknown gall-Tc195-element-unknown gall-Tc196-element-unknown gall-Tc197-element-unknown gall-Tc198-element-unknown gall-Tc199-element-unknown gall-Tc200-element-unknown gall-Tc201-element-unknown gall-Tc202-element-unknown gall-Tc203-element-unknown gall-Tc204-element-unknown gall-Tc205-element-unknown gall-Tc206-element-unknown gall-Tc207-element-unknown gall-Tc208-element-unknown gall-Tc209-element-unknown gall-Tc210-element-unknown gall-Tc211-element-unknown gall-Tc212-element-unknown gall-Tc213-element-unknown gall-Tc214-element-unknown gall-Tc215-element-unknown gall-Tc216-element-unknown gall-Tc217-element-unknown gall-Tc218-element-unknown gall-Tc219-element-unknown gall-Tc220-element-unknown gall-Tc221-element-unknown gall-Tc222-element-unknown gall-Tc223-element-unknown gall-Tc224-element-unknown gall-Tc225-element-unknown gall-Tc226-element-unknown gall-Tc227-element-unknown gall-Tc228-element-unknown gall-Tc229-element-unknown gall-Tc230-element-unknown gall-Tc231-element-unknown gall-Tc232-element-unknown gall-Tc233-element-unknown gall-Tc234-element-unknown gall-Tc235-element-unknown gall-Tc236-element-unknown gall-Tc237-element-unknown gall-Tc238-element-unknown gall-Tc239-element-unknown gall-Tc240-element-unknown gall-Tc241-element-unknown gall-Tc242-element-unknown gall-Tc243-element-unknown gall-Tc244-element-unknown gall-Tc245-element-unknown gall-Tc246-element-unknown gall-Tc247-element-unknown gall-Tc248-element-unknown gall-Tc249-element-unknown gall-Tc250-element-unknown gall-Tc251-element-unknown gall-Tc252-element-unknown gall-Tc253-element-unknown gall-Tc254-element-unknown gall-Tc255-element-unknown gall-Tc256-element-unknown gall-Tc257-element-unknown gall-Tc258-element-unknown gall-Tc259-element-unknown gall-Tc260-element-unknown gall-Tc261-element-unknown gall-Tc262-element-unknown gall-Tc263-element-unknown gall-Tc264-element-unknown gall-Tc265-element-unknown gall-Tc266-element-unknown gall-Tc267-element-unknown gall-Tc268-element-unknown gall-Tc269-element-unknown gall-Tc270-element-unknown gall-Tc271-element-unknown gall-Tc272-element-unknown gall-Tc273-element-unknown gall-Tc274-element-unknown gall-Tc275-element-unknown gall-Tc276-element-unknown gall-Tc277-element-unknown gall-Tc278-element-unknown gall-Tc279-element-unknown gall-Tc280-element-unknown gall-Tc281-element-unknown gall-Tc282-element-unknown gall-Tc283-element-unknown gall-Tc284-element-unknown gall-Tc285-element-unknown gall-Tc286-element-unknown gall-Tc287-element-unknown gall-Tc288-element-unknown gall-Tc289-element-unknown gall-Tc290-element-unknown gall-Tc291-element-unknown gall-Tc292-element-unknown gall-Tc293-element-unknown gall-Tc294-element-unknown gall-Tc295-element-unknown gall-Tc296-element-unknown gall-Tc297-element-unknown gall-Tc298-element-unknown gall-Tc299-element-unknown gall-Tc300-element-unknown gall-Tc301-element-unknown gall-Tc302-element-unknown gall-Tc303-element-unknown gall-Tc304-element-unknown gall-Tc305-element-unknown gall-Tc306-element-unknown gall-Tc307-element-unknown gall-Tc308-element-unknown gall-Tc309-element-unknown gall-Tc310-element-unknown gall-Tc311-element-unknown gall-Tc312-element-unknown gall-Tc313-element-unknown gall-Tc314-element-unknown gall-Tc315-element-unknown gall-Tc316-element-unknown gall-Tc317-element-unknown gall-Tc318-element-unknown gall-Tc319-element-unknown gall-Tc320-element-unknown gall-Tc321-element-unknown gall-Tc322-element-unknown gall-Tc323-element-unknown gall-Tc324-element-unknown gall-Tc325-element-unknown gall-Tc326-element-unknown gall-Tc327-element-unknown gall-Tc328-element-unknown gall-Tc329-element-unknown gall-Tc330-element-unknown gall-Tc331-element-unknown gall-Tc332-element-unknown gall-Tc333-element-unknown gall-Tc334-element-unknown gall-Tc335-element-unknown gall-Tc336-element-unknown gall-Tc337-element-unknown gall-Tc338-element-unknown gall-Tc339-element-unknown gall-Tc340-element-unknown gall-Tc341-element-unknown gall-Tc342-element-unknown gall-Tc343-element-unknown gall-Tc344-element-unknown gall-Tc345-element-unknown gall-Tc346-element-unknown gall-Tc347-element-unknown gall-Tc348-element-unknown gall-Tc349-element-unknown gall-Tc350-element-unknown gall-Tc351-element-unknown gall-Tc352-element-unknown gall-Tc353-element-unknown gall-Tc354-element-unknown gall-Tc355-element-unknown gall-Tc356-element-unknown gall-Tc357-element-unknown gall-Tc358-element-unknown gall-Tc359-element-unknown gall-Tc360-element-unknown gall-Tc361-element-unknown gall-Tc362-element-unknown gall-Tc363-element-unknown gall-Tc364-element-unknown gall-Tc365-element-unknown gall-Tc366-element-unknown gall-Tc367-element-unknown gall-Tc368-element-unknown gall-Tc369-element-unknown gall-Tc370-element-unknown gall-Tc371-element-unknown gall-Tc372-element-unknown gall-Tc373-element-unknown gall-Tc374-element-unknown gall-Tc375-element-unknown gall-Tc376-element-unknown gall-Tc377-element-unknown gall-Tc378-element-unknown gall-Tc379-element-unknown gall-Tc380-element-unknown gall-Tc381-element-unknown gall-Tc382-element-unknown gall-Tc383-element-unknown gall-Tc384-element-unknown gall-Tc385-element-unknown gall-Tc386-element-unknown gall-Tc387-element-unknown gall-Tc388-element-unknown gall-Tc389-element-unknown gall-Tc390-element-unknown gall-Tc391-element-unknown gall-Tc392-element-unknown gall-Tc393-element-unknown gall-Tc394-element-unknown gall-Tc395-element-unknown gall-Tc396-element-unknown gall-Tc397-element-unknown gall-Tc398-element-unknown gall-Tc399-element-unknown gall-Tc400-element-unknown gall-Tc401-element-unknown gall-Tc402-element-unknown gall-Tc403-element-unknown gall-Tc404-element-unknown gall-Tc405-element-unknown gall-Tc406-element-unknown gall-Tc407-element-unknown gall-Tc408-element-unknown gall-Tc409-element-unknown gall-Tc410-element-unknown gall-Tc411-element-unknown gall-Tc412-element-unknown gall-Tc413-element-unknown gall-Tc414-element-unknown gall-Tc415-element-unknown gall-Tc416-element-unknown gall-Tc417-element-unknown gall-Tc418-element-unknown gall-Tc419-element-unknown gall-Tc420-element-unknown gall-Tc421-element-unknown gall-Tc422-element-unknown gall-Tc423-element-unknown gall-Tc424-element-unknown gall-Tc425-element-unknown gall-Tc426-element-unknown gall-Tc427-element-unknown gall-Tc428-element-unknown gall-Tc429-element-unknown gall-Tc430-element-unknown gall-Tc431-element-unknown gall-Tc432-element-unknown gall-Tc433-element-unknown gall-Tc434-element-unknown gall-Tc435-element-unknown gall-Tc436-element-unknown gall-Tc437-element-unknown gall-Tc438-element-unknown gall-Tc439-element-unknown gall-Tc440-element-unknown gall-Tc441-element-unknown gall-Tc442-element-unknown gall-Tc443-element-unknown gall-Tc444-element-unknown gall-Tc445-element-unknown gall-Tc446-element-unknown gall-Tc447-element-unknown gall-Tc448-element-unknown gall-Tc449-element-unknown gall-Tc450-element-unknown gall-Tc451-element-unknown gall-Tc452-element-unknown gall-Tc453-element-unknown gall-Tc454-element-unknown gall-Tc455-element-unknown gall-Tc456-element-unknown gall-Tc457-element-unknown gall-Tc458-element-unknown gall-Tc459-element-unknown gall-Tc460-element-unknown gall-Tc461-element-unknown gall-Tc462-element-unknown gall-Tc463-element-unknown gall-Tc464-element-unknown gall-Tc465-element-unknown gall-Tc466-element-unknown gall-Tc467-element-unknown gall-Tc468-element-unknown gall-Tc469-element-unknown gall-Tc470-element-unknown gall-Tc471-element-unknown gall-Tc472-element-unknown gall-Tc473-element-unknown gall-Tc474-element-unknown gall-Tc475-element-unknown gall-Tc476-element-unknown gall-Tc477-element-unknown gall-Tc478-element-unknown gall-Tc479-element-unknown gall-Tc480-element-unknown gall-Tc481-element-unknown gall-Tc482-element-unknown gall-Tc483-element-unknown gall-Tc484-element-unknown gall-Tc485-element-unknown gall-Tc486-element-unknown gall-Tc487-element-unknown gall-Tc488-element-unknown gall-Tc489-element-unknown gall-Tc490-element-unknown gall-Tc491-element-unknown gall-Tc492-element-unknown gall-Tc493-element-unknown gall-Tc494-element-unknown gall-Tc495-element-unknown gall-Tc496-element-unknown gall-Tc497-element-unknown gall-Tc498-element-unknown gall-Tc499-element-unknown gall-Tc500-element-unknown gall-Tc501-element-unknown gall-Tc502-element-unknown gall-Tc503-element-unknown gall-Tc504-element-unknown gall-Tc505-element-unknown gall-Tc506-element-unknown gall-Tc507-element-unknown gall-Tc508-element-unknown gall-Tc509-element-unknown gall-Tc510-element-unknown gall-Tc511-element-unknown gall-Tc512-element-unknown gall-Tc513-element-unknown gall-Tc514-element-unknown gall-Tc515-element-unknown gall-Tc516-element-unknown gall-Tc517-element-unknown gall-Tc518-element-unknown gall-Tc519-element-unknown gall-Tc520-element-unknown gall-Tc521-element-unknown gall-Tc522-element-unknown gall-Tc523-element-unknown gall-Tc524-element-unknown gall-Tc525-element-unknown gall-Tc526-element-unknown gall-Tc527-element-unknown gall-Tc528-element-unknown gall-Tc529-element-unknown gall-Tc530-element-unknown gall-Tc531-element-unknown gall-Tc532-element-unknown gall-Tc533-element-unknown gall-Tc534-element-unknown gall-Tc535-element-unknown gall-Tc536-element-unknown gall-Tc537-element-unknown gall-Tc538-element-unknown gall-Tc539-element-unknown gall-Tc540-element-unknown gall-Tc541-element-unknown gall-Tc542-element-unknown gall-Tc543-element-unknown gall-Tc544-element-unknown gall-Tc545-element-unknown gall-Tc546-element-unknown gall-Tc547-element-unknown gall-Tc548-element-unknown gall-Tc549-element-unknown gall-Tc550-element-unknown gall-Tc551-element-unknown gall-Tc552-element-unknown gall-Tc553-element-unknown gall-Tc554-element-unknown gall-Tc555-element-unknown gall-Tc556-element-unknown gall-Tc557-element-unknown gall-Tc558-element-unknown gall-Tc559-element-unknown gall-Tc560-element-unknown gall-Tc561-element-unknown gall-Tc562-element-unknown gall-Tc563-element-unknown gall-Tc564-element-unknown gall-Tc565-element-unknown gall-Tc566-element-unknown gall-Tc567-element-unknown gall-Tc568-element-unknown gall-Tc569-element-unknown gall-Tc570-element-unknown gall-Tc571-element-unknown gall-Tc572-element-unknown gall-Tc573-element-unknown gall-Tc574-element-unknown gall-Tc575-element-unknown gall-Tc576-element-unknown gall-Tc577-element-unknown gall-Tc578-element-unknown gall-Tc579-element-unknown gall-Tc580-element-unknown gall-Tc581-element-unknown gall-Tc582-element-unknown gall-Tc583-element-unknown gall-Tc584-element-unknown gall-Tc585-element-unknown gall-Tc586-element-unknown gall-Tc587-element-unknown gall-Tc588-element-unknown gall-Tc589-element-unknown gall-Tc590-element-unknown gall-Tc591-element-unknown gall-Tc592-element-unknown gall-Tc593-element-unknown gall-Tc594-element-unknown gall-Tc595-element-unknown gall-Tc596-element-unknown gall-Tc597-element-unknown gall-Tc598-element-unknown gall-Tc599-element-unknown gall-Tc600-element-unknown gall-Tc601-element-unknown gall-Tc602-element-unknown gall-Tc603-element-unknown gall-Tc604-element-unknown gall-Tc605-element-unknown gall-Tc606-element-unknown gall-Tc607-element-unknown gall-Tc608-element-unknown gall-Tc609-element-unknown gall-Tc610-element-unknown gall-Tc611-element-unknown gall-Tc612-element-unknown gall-Tc613-element-unknown gall-Tc614-element-unknown gall-Tc615-element-unknown gall-Tc616-element-unknown gall-Tc617-element-unknown gall-Tc618-element-unknown gall-Tc619-element-unknown gall-Tc620-element-unknown gall-Tc621-element-unknown gall-Tc622-element-unknown gall-Tc623-element-unknown gall-Tc624-element-unknown gall-Tc625-element-unknown gall-Tc626-element-unknown gall-Tc627-element-unknown gall-Tc628-element-unknown gall-Tc629-element-unknown gall-Tc630-element-unknown gall-Tc631-element-unknown gall-Tc632-element-unknown gall-Tc633-element-unknown gall-Tc634-element-unknown gall-Tc635-element-unknown gall-Tc636-element-unknown gall-Tc637-element-unknown gall-Tc638-element-unknown gall-Tc639-element-unknown gall-Tc640-element-unknown gall-Tc641-element-unknown gall-Tc642-element-unknown gall-Tc643-element-unknown gall-Tc644-element-unknown gall-Tc645-element-unknown gall-Tc646-element-unknown gall-Tc647-element-unknown gall-Tc648-element-unknown gall-Tc649-element-unknown gall-Tc650-element-unknown gall-Tc651-element-unknown gall-Tc652-element-unknown gall-Tc653-element-unknown gall-Tc654-element-unknown gall-Tc655-element-unknown gall-Tc656-element-unknown gall-Tc657-element-unknown gall-Tc658-element-unknown gall-Tc659-element-unknown gall-Tc660-element-unknown gall-Tc661-element-unknown gall-Tc662-element-unknown gall-Tc663-element-unknown gall-Tc664-element-unknown gall-Tc665-element-unknown gall-Tc666-element-unknown gall-Tc667-element-unknown gall-Tc668-element-unknown gall-Tc669-element-unknown gall-Tc670-element-unknown gall-Tc671-element-unknown gall-Tc672-element-unknown gall-Tc673-element-unknown gall-Tc674-element-unknown gall-Tc675-element-unknown gall-Tc676-element-unknown gall-Tc677-element-unknown gall-Tc678-element-unknown gall-Tc679-element-unknown gall-Tc680-element-unknown gall-Tc681-element-unknown gall-Tc682-element-unknown gall-Tc683-element-unknown gall-Tc684-element-unknown gall-Tc685-element-unknown gall-Tc686-element-unknown gall-Tc687-element-unknown gall-Tc688-element-unknown gall-Tc689-element-unknown gall-Tc690-element-unknown gall-Tc691-element-unknown gall-Tc692-element-unknown gall-Tc693-element-unknown gall-Tc694-element-unknown gall-Tc695-element-unknown gall-Tc696-element-unknown gall-Tc697-element-unknown gall-Tc698-element-unknown gall-Tc699-element-unknown gall-Tc700-element-unknown gall-Tc701-element-unknown gall-Tc702-element-unknown gall-Tc703-element-unknown gall-Tc704-element-unknown gall-Tc705-element-unknown gall-Tc706-element-unknown gall-Tc707-element-unknown gall-Tc708-element-unknown gall-Tc709-element-unknown gall-Tc710-element-unknown gall-Tc711-element-unknown gall-Tc712-element-unknown gall-Tc713-element-unknown gall-Tc714-element-unknown gall-Tc715-element-unknown gall-Tc716-element-unknown gall-Tc717-element-unknown gall-Tc718-element-unknown gall-Tc719-element-unknown gall-Tc720-element-unknown gall-Tc721-element-unknown gall-Tc722-element-unknown gall-Tc723-element-unknown gall-Tc724-element-unknown gall-Tc725-element-unknown gall-Tc726-element-unknown gall-Tc727-element-unknown gall-Tc728-element-unknown gall-Tc729-element-unknown gall-Tc730-element-unknown gall-Tc731-element-unknown gall-Tc732-element-unknown gall-Tc733-element-unknown gall-Tc734-element-unknown gall-Tc735-element-unknown gall-Tc736-element-unknown gall-Tc737-element-unknown gall-Tc738-element-unknown gall-Tc739-element-unknown gall-Tc740-element-unknown gall-Tc741-element-unknown gall-Tc742-element-unknown gall-Tc743-element-unknown gall-Tc744-element-unknown gall-Tc745-element-unknown gall-Tc746-element-unknown gall-Tc747-element-unknown gall-Tc748-element-unknown gall-Tc749-element-unknown gall-Tc750-element-unknown gall-Tc751-element-unknown gall-Tc752-element-unknown gall-Tc753-element-unknown gall-Tc754-element-unknown gall-Tc755-element-unknown gall-Tc756-element-unknown gall-Tc757-element-unknown gall-Tc758-element-unknown gall-Tc759-element-unknown gall-Tc760-element-unknown gall-Tc761-element-unknown gall-Tc762-element-unknown gall-Tc763-element-unknown gall-Tc764-element-unknown gall-Tc765-element-unknown gall-Tc766-element-unknown gall-Tc767-element-unknown gall-Tc768-element-unknown gall-Tc769-element-unknown gall-Tc770-element-unknown gall-Tc771-element-unknown gall-Tc772-element-unknown gall-Tc773-element-unknown gall-Tc774-element-unknown gall-Tc775-element-unknown gall-Tc776-element-unknown gall-Tc777-element-unknown gall-Tc778-element-unknown gall-Tc779-element-unknown gall-Tc780-element-unknown gall-Tc781-element-unknown gall-Tc782-element-unknown gall-Tc783-element-unknown gall-Tc784-element-unknown gall-Tc785-element-unknown gall-Tc786-element-unknown gall-Tc787-element-unknown gall-Tc788-element-unknown gall-Tc789-element-unknown gall-Tc790-element-unknown gall-Tc791-element-unknown gall-Tc792-element-unknown gall-Tc793-element-unknown gall-Tc794-element-unknown gall-Tc795-element-unknown gall-Tc796-element-unknown gall-Tc797-element-unknown gall-Tc798-element-unknown gall-Tc799-element-unknown gall-Tc800-element-unknown gall-Tc801-element-unknown gall-Tc802-element-unknown gall-Tc803-element-unknown gall-Tc804-element-unknown gall-Tc805-element-unknown gall-Tc806-element-unknown gall-Tc807-element-unknown gall-Tc808-element-unknown gall-Tc809-element-unknown gall-Tc810-element-unknown gall-Tc811-element-unknown gall-Tc812-element-unknown gall-Tc813-element-unknown gall-Tc814-element-unknown gall-Tc815-element-unknown gall-Tc816-element-unknown gall-Tc817-element-unknown gall-Tc818-element-unknown gall-Tc819-element-unknown gall-Tc820-element-unknown gall-Tc821-element-unknown gall-Tc822-element-unknown gall-Tc823-element-unknown gall-Tc824-element-unknown gall-Tc825-element-unknown gall-Tc826-element-unknown gall-Tc827-element-unknown gall-Tc828-element-unknown gall-Tc829-element-unknown gall-Tc830-element-unknown gall-Tc831-element-unknown gall-Tc832-element-unknown gall-Tc833-element-unknown gall-Tc834-element-unknown gall-Tc835-element-unknown gall-Tc836-element-unknown gall-Tc837-element-unknown gall-Tc838-element-unknown gall-Tc839-element-unknown gall-Tc840-element-unknown gall-Tc841-element-unknown gall-Tc842-element-unknown gall-Tc843-element-unknown gall-Tc844-element-unknown gall-Tc845-element-unknown gall-Tc846-element-unknown gall-Tc847-element-unknown gall-Tc848-element-unknown gall-Tc849-element-unknown gall-Tc850-element-unknown gall-Tc851-element-unknown gall-Tc852-element-unknown gall-Tc853-element-unknown gall-Tc854-element-unknown gall-Tc855-element-unknown gall-Tc856-element-unknown gall-Tc857-element-unknown gall-Tc858-element-unknown gall-Tc859-element-unknown gall-Tc860-element-unknown gall-Tc861-element-unknown gall-Tc862-element-unknown gall-Tc863-element-unknown gall-Tc864-element-unknown gall-Tc865-element-unknown gall-Tc866-element-unknown gall-Tc867-element-unknown gall-Tc868-element-unknown gall-Tc869-element-unknown gall-Tc870-element-unknown gall-Tc871-element-unknown gall-Tc872-element-unknown gall-Tc873-element-unknown gall-Tc874-element-unknown gall-Tc875-element-unknown gall-Tc876-element-unknown gall-Tc877-element-unknown gall-Tc878-element-unknown gall-Tc879-element-unknown gall-Tc880-element-unknown gall-Tc881-element-unknown gall-Tc882-element-unknown gall-Tc883-element-unknown gall-Tc884-element-unknown gall-Tc885-element-unknown gall-Tc886-element-unknown gall-Tc887-element-unknown gall-Tc888-element-unknown gall-Tc889-element-unknown gall-Tc890-element-unknown gall-Tc891-element-unknown gall-Tc892-element-unknown gall-Tc893-element-unknown gall-Tc894-element-unknown gall-Tc895-element-unknown gall-Tc896-element-unknown gall-Tc897-element-unknown gall-Tc898-element-unknown gall-Tc899-element-unknown gall-Tc900-element-unknown gall-Tc901-element-unknown gall-Tc902-element-unknown gall-Tc903-element-unknown gall-Tc904-element-unknown gall-Tc905-element-unknown gall-Tc906-element-unknown gall-Tc907-element-unknown gall-Tc908-element-unknown gall-Tc909-element-unknown gall-Tc910-element-unknown gall-Tc911-element-unknown gall-Tc912-element-unknown gall-Tc913-element-unknown gall-Tc914-element-unknown gall-Tc915-element-unknown gall-Tc916-element-unknown gall-Tc917-element-unknown gall-Tc918-element-unknown gall-Tc919-element-unknown gall-Tc920-element-unknown gall-Tc921-element-unknown gall-Tc922-element-unknown gall-Tc923-element-unknown gall-Tc924-element-unknown gall-Tc925-element-unknown gall-Tc926-element-unknown gall-Tc927-element-unknown gall-Tc928-element-unknown gall-Tc929-element-unknown gall-Tc930-element-unknown gall-Tc931-element-unknown gall-Tc932-element-unknown gall-Tc933-element-unknown gall-Tc934-element-unknown gall-Tc935-element-unknown gall-Tc936-element-unknown gall-Tc937-element-unknown gall-Tc938-element-unknown gall-Tc939-element-unknown gall-Tc940-element-unknown gall-Tc941-element-unknown gall-Tc942-element-unknown gall-Tc943-element-unknown gall-Tc944-element-unknown gall-Tc945-element-unknown gall-Tc946-element-unknown gall-Tc947-element-unknown gall-Tc948-element-unknown gall-Tc949-element-unknown gall-Tc950-element-unknown gall-Tc951-element-unknown gall-Tc952-element-unknown gall-Tc953-element-unknown gall-Tc954-element-unknown gall-Tc955-element-unknown gall-Tc956-element-unknown gall-Tc957-element-unknown gall-Tc958-element-unknown gall-Tc959-element-unknown gall-Tc960-element-unknown gall-Tc961-element-unknown gall-Tc962-element-unknown gall-Tc963-element-unknown gall-Tc964-element-unknown gall-Tc965-element-unknown gall-Tc966-element-unknown gall-Tc967-element-unknown gall-Tc968-element-unknown gall-Tc969-element-unknown gall-Tc970-element-unknown gall-Tc971-element-unknown gall-Tc972-element-unknown gall-Tc973-element-unknown gall-Tc974-element-unknown gall-Tc975-element-unknown gall-Tc976-element-unknown gall-Tc977-element-unknown gall-Tc978-element-unknown gall-Tc979-element-unknown gall-Tc980-element-unknown gall-Tc981-element-unknown gall-Tc982-element-unknown gall-Tc983-element-unknown gall-Tc984-element-unknown gall-Tc985-element-unknown gall-Tc986-element-unknown gall-Tc987-element-unknown gall-Tc988-element-unknown gall-Tc989-element-unknown gall-Tc990-element-unknown gall-Tc991-element-unknown gall-Tc992-element-unknown gall-Tc993-element-unknown gall-Tc994-element-unknown gall-Tc995-element-unknown gall-Tc996-element-unknown gall-Tc997-element-unknown gall-Tc998-element-unknown gall-Tc999-element-unknown gall-Tc1000-element-unknown
Leptodictya plana
Grass Lacebug
Leptodictya plana is a grass-feeding lace bug (Hemiptera: Tingidae) first described in 1913. It has been documented as an emerging pest on ornamental grasses in the southern United States, with Pennisetum species showing particular susceptibility to feeding damage and serving as oviposition hosts. The species remains seldom collected despite its economic relevance to horticulture.
lace-bug grass-pest ornamental-pest Tingidae phloem-feeder emerging-pest Pennisetum-pest Hemiptera Heteroptera seldom-collected
Leptoglossus
leaf-footed bugs
Leptoglossus is a genus of true bugs in the leaf-footed bug family Coreidae, tribe Anisoscelini. Species are characterized by leaflike dilations of the hind tibia, a diagnostic trait of the genus. The genus is distributed throughout the Americas, with some introduced populations in Europe and Asia. Several species are economically significant agricultural pests, notably L. occidentalis, which has become invasive in multiple continents.
Coreidae leaf-footed-bug agricultural-pest invasive-species symbiosis sexual-dimorphism conifer-pest nuisance-pest Burkholderia tachinid-parasitoid pheromone-communication building-invader misidentification Triatoma-look-alike gradual-metamorphosis seed-predator forest-pest ornamental-pest plumbing-damage public-health-confusion Chile Europe North-America South-America Asia Turkey almond-pest citrus-pest tomato-pest corn-pest overwintering-aggregation defensive-secretion pheromone-mediated-parasitism egg-parasitoid biological-control ecological-niche-modeling climate-suitability range-expansion introduction accidental-dispersal seed-orchard-pest germination-reduction empty-seed-formation cone-damage lodgepole-pine Douglas-fir western-conifer-seed-bug L.-occidentalis L.-zonatus L.-phyllopus L.-clypealis L.-australis L.-chilensis L.-oppositus male-combat femoral-weapon abdominal-gland pheromone-gland tibial-dilation leaf-like-hind-leg true-bug Hemiptera Heteroptera Pentatomomorpha Anisoscelini phytophagous plant-feeding stylet-feeding salivary-sheath overwintering diapause building-entry structural-pest nuisance-odor flight-sound droning-flight aggregation-behavior male-pheromone female-choice parasitoid-host biological-control-agent integrated-pest-management monitoring early-detection prevention invasion-biology alien-species non-native-pest global-spread climate-change niche-modeling suitable-habitat establishment-risk quarantine phytosanitary forest-health seed-production reforestation afforestation conifer-forestry pine-seed seed-viability economic-impact crop-loss yield-reduction kernel-damage fruit-damage nut-damage vector-of-disease Chagas-disease kissing-bug public-alarm misinformation education identification-tools health-system-burden pesticide-overuse urban-ecology domiciliary peridomiciliary Andean-region Patagonia Mediterranean temperate subtropical tropical agroecosystem natural-enemy predator parasite pathogen symbiont gut-microbiome soil-ingestion fitness-benefit reproductive-success sperm-morphology testicular-morphology accessory-gland pheromone-blend species-specific-odor cherry-scent vanilla-scent cinnamon-scent rose-scent olfactory-communication mate-location mate-recognition sexual-selection male-male-competition weapon-morphology allometry developmental-plasticity nymphal-instar egg-barrel egg-arrangement linear-oviposition leaf-surface plant-tissue stylet-penetration enzymatic-digestion fluid-feeding phloem-feeding xylem-feeding seed-feeding cone-feeding fruit-feeding kernel-feeding nut-feeding crop-feeding host-plant-range polyphagy oligophagy specialist generalist pest-status damage-threshold economic-injury-level management-strategy cultural-control physical-control chemical-control resistance tolerance host-plant-resistance monitoring-trap light-trap population-density distribution-map spread-rate colonization establishment population-dynamics life-table developmental-rate thermal-requirements degree-days phenology voltinism univoltine bivoltine multivoltine generation-time overwintering-site hibernaculum shelter-seeking cold-hardiness freeze-tolerance supercooling water-relations desiccation-resistance starvation-resistance dispersal-ability flight-capacity walking-behavior climbing-behavior aggregation-pheromone alarm-pheromone stink-gland metathoracic-gland dorsoabdominal-gland ventral-abdominal-gland sexual-gland morphological-defense chemical-defense mechanical-defense autotomy leg-loss predation-risk parasitism-risk parasitoid-risk pathogen-risk competition intraspecific interspecific resource-competition mating-competition sperm-competition cryptic-female-choice reproductive-isolation speciation phylogeny systematics taxonomy morphology anatomy histology ultrastructure spermatogenesis spermatozoa sperm-length nuclear-length cyst-production follicle-number testis-structure reproductive-anatomy genitalia mating-behavior copulation insemination oviposition egg-production fecundity fertility hatching-success nymphal-survival adult-longevity sex-ratio operational-sex-ratio population-sex-ratio effective-population-size genetic-diversity gene-flow population-structure phylogeography biogeography historical-biogeography vicariance dispersal range-shift range-contraction altitudinal-distribution latitudinal-distribution longitudinal-distribution island-distribution continental-distribution endemism cosmopolitanism introduced-range native-range source-population founder-population invasion-front lag-phase establishment-phase spread-phase equilibrium-phase impact-assessment risk-assessment horizon-scanning early-warning rapid-response eradication containment control management adaptation mitigation restoration conservation biodiversity ecosystem-service ecosystem-function food-web trophic-level primary-consumer herbivore fruit-predator plant-animal-interaction mutualism antagonism predation parasitism commensalism amensalism facilitation indirect-interaction trait-mediated-interaction density-mediated-interaction behavioral-ecology evolutionary-ecology functional-ecology physiological-ecology population-ecology community-ecology ecosystem-ecology landscape-ecology macroecology global-ecology applied-ecology agricultural-ecology forest-ecology conservation-biology invasion-ecology restoration-ecology pest-management conservation-biological-control augmentative-biological-control classical-biological-control parasitoid nematode fungus bacterium virus entomopathogen microbial-control sterile-insect-technique genetic-control mating-disruption attract-and-kill push-pull trap-crop border-crop refuge habitat-management cultural-practice crop-rotation tillage irrigation fertilization pruning harvest-timing sanitation residue-management weed-management cover-crop companion-planting intercropping agroforestry silviculture forest-management seed-orchard-management cone-collection seed-extraction seed-testing germination-testing seed-quality seed-vigor empty-seed insect-damage insect-injury feeding-scar puncture stylet-sheath salivary-flange symptom sign diagnosis sampling economic-threshold decision-support expert-system modeling simulation forecasting climate-model niche-model species-distribution-model habitat-suitability risk-mapping invasion-pathway introduction-vector trade transport tourism migration natural-dispersal human-mediated-dispersal accidental-introduction intentional-introduction release escape cultivation ornamental forestry agriculture horticulture urbanization globalization land-use-change habitat-fragmentation habitat-loss degradation pollution pesticide fertilizer nutrient-enrichment eutrophication acidification warming drought extreme-event disturbance succession recovery resilience stability variability uncertainty sustainable-development green-economy circular-economy bioeconomy ecosystem-approach one-health planetary-health environmental-health public-health veterinary-health plant-health food-security nutrition livelihood poverty inequality gender indigenous-knowledge traditional-knowledge local-knowledge citizen-science stakeholder-engagement policy governance regulation legislation international-cooperation capacity-building awareness communication outreach extension advisory-service research innovation technology-transfer knowledge-exchange networking partnership collaboration evaluation evidence-based-policy science-policy-interface diplomacy advocacy activism social-movement environmental-justice environmental-ethics environmental-philosophy environmental-humanities environmental-history environmental-literature environmental-art environmental-education environmental-communication science-communication risk-communication crisis-communication health-communication behavior-change pro-environmental-behavior sustainable-behavior consumption production waste recycling reuse reduction efficiency renewable clean green blue natural organic agroecology permaculture regenerative restorative healing transformative transdisciplinary interdisciplinary multidisciplinary cross-disciplinary disciplinary disciplinary-integration knowledge-integration system-thinking complexity emergence non-linearity feedback tipping-point regime-shift alternative-stable-state resilience-thinking adaptive-management adaptive-governance learning reflection participation inclusion equity justice fairness ethics values norms culture identity place sense-of-place attachment well-being quality-of-life happiness flourishing thriving sustainability sustainable development growth degrowth post-growth steady-state circular doughnut-economics planetary-boundaries safe-operating-space tipping-elements critical-transition early-warning-signal resilience-indicator sustainability-indicator benchmark target goal commitment pledge agreement treaty convention protocol framework strategy plan program project initiative campaign movement coalition alliance network platform hub center institute organization association society union federation confederation foundation trust fund bank facility mechanism instrument tool method approach technique procedure process system structure institution administration operation service delivery provision supply demand market economy finance investment funding financing resource capital asset liability income expenditure revenue cost benefit profit loss return yield outcome output input throughput productivity effectiveness performance quality standard criterion indicator metric measure assessment appraisal review audit verification validation certification accreditation recognition award prize honor distinction excellence best-practice lesson-learned experience expertise competence capacity capability skill knowledge information data evidence science discovery invention creation design testing piloting scaling mainstreaming uptake adoption diffusion dissemination publication reporting documentation archiving preservation rehabilitation reconstruction replication duplication backup security safety protection safeguarding insurance risk-management emergency-preparedness response relief humanitarian peace prosperity health employment housing infrastructure energy water food fishery aquaculture mining manufacturing industry commerce recreation heritage nature ecosystem environment climate atmosphere ocean freshwater land soil mineral biota flora fauna microorganism fungi plant animal vertebrate invertebrate arthropod insect bug Hemipteran Heteropteran pentatomomorphan coreoid coreid squash-bug boxelder-bug stink-bug shield-bug plant-bug mirid lygaeid seed-bug broad-headed-bug alydid reduviid assassin-bug triatomine bed-bug cimicid water-bug nepomorphan gerromorphan leptopodomorphan enicocephalomorphan dipsocoromorphan ceratocomboid schizopteroid peloridioid coleorrhynchan moss-bug archaeorrhynchan fulgoromorphan cicadomorphan membracoid treehopper leafhopper planthopper psyllid jumping-plant-louse whitefly aleyrodid scale-insect coccoid mealybug aphid adelgid phylloxeran sternorrhynchan thysanopteran thrips psocopteran barklouse booklouse phthirapteran louse sucking-louse chewing-louse mallophagan anopluran dermapteran earwig blattodean cockroach termite isopteran mantodean mantid phasmid stick-insect leaf-insect orthopteran grasshopper locust katydid cricket mole-cricket pygmy-mole-cricket camel-cricket cave-cricket weta ensiferan caeliferan gryllotalpid myrmecophilid tettigoniid gryllid acridid pamphagid pneumorid lentulid tristirid eumastacid proscopiid tridactylid tetrigid grouse-locust pygmy-grasshopper plecopteran stonefly embiopteran webspinner zorapteran angel-insect dictyopteran
Leptoypha minor
Arizona ash lace bug, ash lace bug
Leptoypha minor, commonly known as the Arizona ash lace bug, is a species of lace bug in the family Tingidae. It is a documented pest of ash trees (Fraxinus spp.), causing damage to twigs and foliage. The species is particularly abundant in California and has been economically significant as an ornamental pest since its formal description in 1942.
pest lace-bug ash Fraxinus ornamental-pest California multiple-generations
Leucoptera laburnella
Laburnum Leafminer, laburnum leaf miner
Leucoptera laburnella is a small moth in the family Lyonetiidae, commonly known as the laburnum leaf miner. The larvae are leaf miners that feed internally on leaves of various Fabaceae species, particularly Laburnum. Adults have distinctive white forewings marked with yellow and dark fuscous patterns. The species is native to Europe and has been introduced to North America.
leaf-miner Fabaceae ornamental-pest Europe North-America introduced-species Lyonetiidae
Liriomyza trifolii
American serpentine leafminer, celery leafminer
Liriomyza trifolii is a small leaf-mining fly in the family Agromyzidae, native to the Caribbean and southeastern United States. It has become a globally distributed invasive pest of greenhouse and vegetable crops, recognized as one of the three most damaging leafminer species worldwide. The species is highly polyphagous, feeding on numerous ornamental and vegetable plants including chrysanthemums, celery, and tomatoes. Its spread has been facilitated by insecticide resistance and international trade in plant material.
leafminer agricultural-pest greenhouse-pest invasive-species polyphagous insecticide-resistance vegetable-pest ornamental-pest
Lopholeucaspis
Lopholeucaspis is a genus of armored scale insects (Diaspididae) containing species that are invasive pests of woody plants. The most documented species, L. japonica, has established populations across Asia, Europe, Australia, and North America. These scales infest a broad range of hosts including citrus, pomegranate, crapemyrtle, privet, and euonymus. The genus is notable for its expanding global distribution and association with multiple parasitoid wasp species.
scale-insect armored-scale invasive-pest phloem-feeder woody-plant-pest citrus-pest ornamental-pest
Macremphytus
Dogwood Sawflies
Macremphytus is a genus of common sawflies in the family Tenthredinidae, comprising at least four described species. The best-known member is Macremphytus tarsatus, commonly called the dogwood sawfly, which is a significant defoliator of dogwood trees in North America. Sawflies in this genus are herbivorous, with larvae feeding on leaves of woody plants. The genus exhibits distinctive larval traits including waxy secretions and overwintering behaviors in wood.
sawfly defoliator dogwood walnut waxy-secretion wood-boring ornamental-pest Hymenoptera Tenthredinidae North-America
Macrohomotoma
Macrohomotoma is a genus of psyllids in the family Homotomidae, established by Kuwayama in 1908. The genus contains approximately 15 described species of plant-parasitic hemipterans. Best-known species include Macrohomotoma gladiata, a significant pest of ornamental Ficus trees that has expanded from its native Asian range to become invasive in the Mediterranean region and elsewhere. The genus is characterized by association with Ficus host plants and production of conspicuous waxy secretions.
psyllid Ficus-pest invasive-species ornamental-pest urban-forestry Liberibacter-vector
Macrohomotoma gladiata
Curtain Fig Psyllid
Macrohomotoma gladiata is an invasive Asian psyllid that has established populations across the Mediterranean Basin and California, where it infests ornamental Ficus microcarpa. Nymphs feed on young tissues, producing dense white waxy secretions and honeydew that promotes sooty mould growth and causes twig dieback. The species has a likely bivoltine life cycle with possible partial third generation; young nymphs overwinter under stipules without development, resuming growth in spring. Natural enemies include a Psyllaephagus parasitoid that can achieve high parasitism rates, and the predatory bug Anthocoris nemoralis.
invasive-species urban-pest ornamental-pest biological-control Ficus-pest Mediterranean-invasion psyllid Homotomidae Sternorrhyncha phloem-feeder waxy-secretions sooty-mould parasitoid Psyllaephagus Anthocoris-nemoralis bivoltine overwintering-nymphs Liberibacter-vector
Melanchroia chephise
white-tipped black, snowbush spanworm
Melanchroia chephise is a moth in the family Geometridae, commonly known as the white-tipped black or snowbush spanworm. It ranges from the southern United States through Central America to Paraguay. The adult exhibits striking black wings with white apical patches, while the gregarious larvae feed on foliage of plants in the Phyllanthaceae family, particularly Breynia and Phyllanthus species. Despite occasional outbreaks that defoliate ornamental plantings, the species typically causes no lasting harm to hosts and poses no risk to humans.
Geometridae defoliator gregarious-larvae ornamental-pest aposematic-coloration wasp-mimicry Phyllanthaceae Neotropical-distribution
Merhynchites
Rose Weevils
Merhynchites is a genus of leaf and bud weevils in the family Attelabidae, containing approximately 12 described species. The genus includes the well-known rose curculio (Merhynchites bicolor), a significant pest of cultivated roses. Members of this genus are characterized by their elongated rostrums and association with flowering plants, particularly roses.
pest roses ornamental-pest
Metamasius mosieri
Florida bromeliad weevil
Metamasius mosieri is a small weevil in the family Dryophthoridae, commonly known as the Florida bromeliad weevil. It feeds on bromeliads but occurs at very low population densities in natural habitats and is considered only an occasional, minor pest on ornamental species. It is native to Florida and Cuba and is frequently confused with the more destructive Mexican bromeliad weevil (Metamasius callizona).
weevil bromeliad Florida Cuba minor-pest ornamental-pest native-species taxonomic-confusion
Myzocallis walshii
Black-bordered oak aphid
Myzocallis walshii is a North American aphid species that has established in Europe as an invasive pest of American red oak (Quercus rubra). Native parasitoids in the northeastern Iberian Peninsula have adapted to this introduced aphid. The species has been recorded across much of North America and has spread to at least 15 European countries.
invasive-species pest oak-aphid parasitoid-host range-expansion ornamental-pest
Neohydatothrips samayunkur
marigold thrips, French marigold thrips
Neohydatothrips samayunkur is a thrips species in the family Thripidae, commonly known as the marigold thrips. It is an economically significant pest of Tagetes species (marigolds), feeding on flowers and leaves. The species has a broad intercontinental distribution spanning Africa, Australia, Europe, Northern Asia, Central America, and North America. Originally described by Kudo in 1995 and validated by Nakahara in 1999, it has been intercepted at U.S. quarantine ports and represents a new country record for Egypt.
agricultural-pest ornamental-pest quarantine-pest Tagetes-pest marigold-pest invasive-potential Thripidae Sericothripinae Thysanoptera
Neolecanium cornuparvum
Magnolia scale
Neolecanium cornuparvum, commonly known as the magnolia scale, is a soft scale insect in the family Coccidae. It is a significant pest of magnolia trees in North America, where heavy infestations can cause branch dieback and tree decline. The species is included in the USA National Phenology Network's Pheno Forecast system, which uses heat accumulation models to predict its life cycle stages for optimized pest management. N. cornuparvum has been documented in the United States, including in Vermont.
pest scale-insect ornamental-pest USA-NPN
Neomyzus circumflexus
Crescent-marked lily aphid, Mottled arum aphid
Neomyzus circumflexus is a polyphagous aphid species known by two common names reflecting its association with lilies and arums. It is a significant greenhouse and ornamental pest, feeding on phloem sap and transmitting plant viruses. The species exhibits temperature-dependent immune responses to parasitoids and has been extensively studied for its nutritional physiology, including artificial diet requirements and symbiont dependencies.
aphid pest greenhouse-pest virus-vector ornamental-pest parthenogenetic
Neophyllaphis podocarpi
Podocarpus Aphid
Neophyllaphis podocarpi is an aphid species in the subfamily Neophyllaphidinae, known as the podocarpus aphid. It is specialized on Podocarpus species (Podocarpaceae), a genus of coniferous trees. The species was first described from Japan in 1920 and has since been recorded in multiple regions including Hawaii, Italy, the Azores, and the Iberian Peninsula. Its spread appears linked to horticultural trade of host plants.
aphid invasive ornamental-pest Podocarpaceae
Oligonychus ununguis
spruce spider mite
Oligonychus ununguis, commonly known as the spruce spider mite, is a globally distributed pest of coniferous trees, particularly spruce. It is considered one of the most important pests of spruce plantations in Ontario and causes significant damage to ornamental conifers. The mite feeds on needle sap, causing foliage discoloration, and produces silk webbing that accumulates dust. Populations thrive in cool spring and autumn conditions but decline during hot summer weather when temperatures exceed 85°F.
pest conifer cool-season-mite silvicultural-pest ornamental-pest
Oligotrophus
Oligotrophus is a genus of gall midges (Cecidomyiidae: Diptera) established by Latreille in 1804. Species are gall-forming insects that induce abnormal plant growths on various host plants, with documented associations to birch (Betulaceae) and juniper (Cupressaceae). The genus contains univoltine species with specialized life cycles tied to host phenology. Several species have been studied for their economic impact on ornamental and forest trees.
gall-midge univoltine phytophagous forest-pest ornamental-pest invasive-potential
Oligotrophus betheli
Juniper Tip Midge
Oligotrophus betheli is a gall midge native to North America that induces galls on Juniperus horizontalis (Cupressaceae). It was first recorded as an alien species in Japan, where it was found on introduced ornamental host plants. The species causes characteristic discoloration of young twig tips on its host.
gall-midge alien-species invasive-potential ornamental-pest Cecidomyiidae
Ophiomyia kwansonis
Daylily Leafminer
Ophiomyia kwansonis is a leaf-miner fly in the family Agromyzidae. It is known as the Daylily Leafminer based on its documented association with Hemerocallis species. The species was described by Sasakawa in 1961 and has been recorded across eastern North America, parts of Asia, and Europe.
leaf-miner pest Hemerocallis introduced-species ornamental-pest
Ormenaria rufifascia
Palm Flatid Planthopper
Ormenaria rufifascia is a flatid planthopper commonly found on palms in Florida and parts of the Caribbean. Adults and nymphs are visually conspicuous. Dense populations produce honeydew, which can lead to extensive sooty mold growth on host plants, causing aesthetic damage even though the insect is usually not a serious pest.
flatid-planthopper palm-pest honeydew-producer sooty-mold Florida-insect ornamental-pest
Otiorhynchus meridionalis
Lilac Root Weevil
Otiorhynchus meridionalis, commonly known as the lilac root weevil, is a broad-nosed weevil in the family Curculionidae. It is a recognized landscape pest in North America, where it feeds on ornamental plants including lilac, euonymus, and peonies. The species exhibits a life cycle with root-feeding larvae and leaf-feeding adults that produce characteristic notched damage. Adults are nocturnal and occasionally enter buildings during hot weather. Native to Europe, it has established populations in western North America.
pest introduced nocturnal root-weevil ornamental-pest
Pachypsylla celtidisumbilicus
Hackberry Disc Gall Psyllid
Pachypsylla celtidisumbilicus is a species of hackberry psyllid that produces distinctive disc-shaped galls on hackberry leaves (Celtis spp.). Adults emerge in fall and seek shelter to overwinter, often becoming household nuisances when they congregate on building exteriors. The species is one of at least seven Pachypsylla species associated with hackberry in North America, each producing a characteristic gall morphology.
psyllid gall-former hackberry Celtis Sternorrhyncha pest nuisance overwintering parasitoid-host disc-gall button-gall North-America ornamental-pest seasonal-emergence jumping-plant-louse Aphalaridae Psylloidea Hemiptera Insecta Arthropoda Animalia Pachypsylla celtidisumbilicus Riley-1890 accepted-species 358-iNaturalist-observations Hackberry-Disc-Gall-Psyllid leaf-gall Celtis-occidentalis Celtis-laevigata common-hackberry sugar-hackberry Psyllaephagus Torymus parasitoid fall-emergence spring-oviposition building-nuisance non-biting non-stinging plant-sap-feeder phloem-feeder piercing-sucking-mouthparts overwintering-adult five-instars wing-pads gall-exit jumping-behavior vertical-surface-aggregation transient-pest no-structural-damage tree-health-impact-minimal gall-diversity congeneric-species gall-morphology-identification Cecidomyiidae gall-midge distinguishing-galls Caldwell-1938 Berenbaum-1989 Winterringer-1961 Bug-Eric iNaturalist GBIF Catalogue-of-Life Encyrtidae Torymidae biological-control host-record new-host-record-potential gall-rearing Colorado-Springs Leavenworth-Kansas North-American-distribution hackberry-range ornamental-tree urban-pest suburban-pest seasonal-nuisance fall-insect autumn-insect spring-insect leaf-unfolding bud-break precise-oviposition-timing gall-induction plant-insect-interaction herbivore phytophagous specialist-feeder monophagous oligophagous Celtis-specialist jumping-plant-lice Psyllidae Psyllomorpha true-bug Hexapoda Eukaryota species accepted Riley 1890 358-observations no-Wikipedia-summary GBIF-exact-match Catalogue-of-Life-accepted taxon entomology insect-guide structured-record factual conservative informative non-repetitive cautious-language no-inference no-fabrication high-quality medium-completeness no-inferred-content
Palpita flegia
satin white moth, Satin White Palpita
Palpita flegia is a crambid moth commonly known as the satin white moth. It was described by Cramer in 1777 and has a broad distribution across the Neotropics and into the southern United States. The species is a documented pest of ornamental plants in the family Apocynaceae, particularly Cascabela thevetia. Laboratory studies have established detailed life cycle parameters including six larval instars and specific thermal requirements for development.
pest ornamental-pest Apocynaceae-feeder Neotropical thermal-biology-documented biological-control-candidate
Paraleyrodes
nesting whiteflies, Bondar's nesting whitefly (for P. bondari), fig whitefly (for P. minei)
Paraleyrodes is a genus of whiteflies in the subfamily Aleurodicinae, commonly known as nesting whiteflies due to the woolly wax nests constructed by females around egg masses. The genus includes economically significant invasive species such as P. bondari (Bondar's nesting whitefly) and P. minei, which are pests of coconut, citrus, fig, avocado, and ornamental plants. Species in this genus are characterized by distinctive waxy secretions, sessile nymphal stages, and polyphagous feeding habits on woody plants.
whitefly invasive-pest agricultural-pest ornamental-pest Aleurodicinae coconut-pest citrus-pest fig-pest avocado-pest woolly-wax-nests polyphagous phloem-feeder honeydew-producer sooty-mold Florida-pest California-pest India-pest Bangladesh-pest Croatia-pest Greece-pest natural-enemies biological-control integrated-pest-management
Periphyllus
Periphyllus Aphids, maple aphids
Periphyllus is a genus of aphids in the family Aphididae, subfamily Chaitophorinae, containing more than 40 described species. Species in this genus are primarily associated with maple trees (Acer spp.), though some feed on related host plants such as Koelreuteria. Many species exhibit complex life cycles with polymorphic forms including viviparous females, aestivating dimorphs, and sexual morphs. Several species are invasive pests outside their native ranges, spreading with ornamental host plants.
aphid maple pest invasive holocyclic aestivating-dimorph polymorphism phloem-feeder ornamental-pest Chaitophorinae
Periploca ceanothiella
Ceanothus stem gall moth
A small gall-forming moth in the family Cosmopterigidae that induces stem galls on Ceanothus species. The species has been recorded across much of North America, with adults active primarily from March to August. Larvae bore into host stems, causing distinctive gall formations, and overwinter within these structures.
gall-former pest ornamental-pest concealed-feeder stem-borer
Phenacoccus aceris
apple mealybug, maple mealybug
Phenacoccus aceris, commonly known as the apple mealybug or maple mealybug, is a widespread polyphagous mealybug species in the family Pseudococcidae. Native to Europe, it has become established across the Holarctic region including North America and Asia. The species is a significant agricultural and horticultural pest, feeding on phloem sap of numerous host plants including apple, maple, grapevine, ash, and prickly ash. Recent molecular and morphological studies have revealed that P. aceris represents a species complex containing multiple cryptic species, with taxonomic confusion arising from co-occurring putative species that cannot be reliably separated by host-plant associations alone. The species is also an important vector of grapevine viruses, transmitting six ampeloviruses and two vitiviruses.
mealybug pest virus-vector grapevine apple polyphagous Holarctic species-complex biological-control integrated-pest-management horticulture viticulture cryptic-species phloem-feeder Sternorrhyncha Pseudococcidae Phenacoccus agricultural-pest ornamental-pest leafroll-disease ampelovirus vitivirus
Phloeosinus
cedar bark beetles, cypress bark beetles
Phloeosinus is a genus of bark beetles (Curculionidae: Scolytinae) comprising at least 20 described species. Species in this genus are specialized on conifers in the family Cupressaceae, including genera such as Cupressus, Thuja, Juniperus, and Chamaecyparis. Several species have become significant invasive pests, causing economic damage to ornamental trees and forest ecosystems. The genus exhibits specialized chemical ecology, with species using host plant volatiles and pheromones for mate and host location.
bark-beetle Cupressaceae invasive-pest chemical-ecology pheromone forest-pest ornamental-pest phloem-feeder
Phytomyza sempervirentis
English Holly Leafminer
Phytomyza sempervirentis is a leaf-mining fly in the family Agromyzidae. The species was described in 2018 and is associated with Ilex species, including English holly (Ilex aquifolium). Larvae create serpentine mines within holly leaves, feeding between the upper and lower epidermis. Adults are small black flies that emerge in spring to coincide with the production of tender new leaves.
leafminer holly Ilex Agromyzidae Phytomyza Diptera invasive-species-associate ornamental-pest
Podosesia
Ash and Lilac Borer Moths
Podosesia is a genus of clearwing moths in the family Sesiidae, established by Möschler in 1879 after the original name Grotea was found to be preoccupied. The genus contains at least three described species in North America, including the economically significant lilac borer (P. syringae) and the banded ash clearwing (P. aureocincta). Larvae are wood-borers in woody plants of the olive family (Oleaceae), particularly ash, lilac, and privet. Adults are diurnal and exhibit wasp mimicry in both appearance and behavior.
Sesiidae clearwing-moth wood-borer wasp-mimicry Oleaceae pest Pheno-Forecast urban-forestry ornamental-pest
Polistes dominula
European Paper Wasp
Polistes dominula is a highly successful invasive social wasp native to Eurasia that has established populations across North America, South America, New Zealand, South Africa, and other regions. First detected in North America near Boston in 1978, it has become one of the most abundant wasps on the continent. The species builds small, exposed paper nests in protected locations and preys primarily on live insects, particularly caterpillars. Unlike yellowjackets, it does not scavenge for meat or sugar. Its rapid spread has been attributed to ecological flexibility, superior competitive ability, and tolerance of human-altered environments.
invasive-species social-wasp paper-wasp predator pollinator urban-ecology eusocial nest-construction biological-control thermal-biology spatial-learning vibrational-communication aposematism honest-signal overwintering diapause Polistes Vespidae Hymenoptera North-America Europe New-Zealand South-Africa South-America monarch-butterfly caterpillar-predation nest-architecture gyne foundress dominance-hierarchy climate-change-impact energetics metabolic-rate microclimate suburban-habitat anthropogenic ornamental-pest vineyard-pest fruit-damage stinging-insect venom poison-gland warning-coloration Zahavian-signal Strepsiptera parasitoid Xenos behavioral-plasticity foraging learning memory thermoregulation ectothermy endothermy solar-radiation nest-temperature brood-development colony-survival multiple-founding single-founding season-length latitude temperate Mediterranean hibernacle overwintering-costs climate-warming range-expansion competition native-species-displacement 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catalogue-of-life GBIF iNaturalist observation citizen-science photography illustration field-guide orange black yellow red color-pattern size wing-length body compact legs short face square triangular dark curled straight elbowed blunt pointed tip Zahavian costly-signal hunt forage nectar honeydew aphid scale-insect flower grape umbellifer fennel lovage insect meatball feed emerge chew cap hexagonal wood-fiber bark exposed open umbrella stalk ant repellent substance coat protect location eave attic tree branch vine shrub cavity bird-box bee-hive shed grave-lantern overwinter shelter protected thermal climate cold winter spring summer autumn fall season month day activity defend threat display raise tiptoe social dominance hierarchy reproductive egg lay found cooperate compete selfish passive-aggressive individual recognition facial-marking cognition mushroom-body brain intelligent artist architect navigation plasticity spatial relocating food resource dish bait training experience 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ReplyDelete RepliesReplyOut prairieSeptember 8:02 keeping cabbage worm seen RepliesReplyAnonymousSeptember 13 2:06 PM becoming here Upper Peninsula Michigan pendulum seems swung ago everywhere vanishingly rare haven't again house right RepliesReplyL.R ShimerSeptember 4:35 head gal work everybody else kinda egalitarian RepliesReplyLarryMay 9:34 directed Bev Wigney BTW lived NE Missouri detailed presented Thanks cogent scientifically accurate summary insect's life-cycle RepliesBug EricMay 3:47 kind compliments Delete RepliesReplyReplyAdd commentLoad Blog currently reply reader himself Working resolve Newer Older Subscribe Atom Demonized hosting 4 p.m.Saturday Jan 20 Room 1124 Academic Surge 455 Crocker Lane campus Attendees chat check 50,000 vespid eight-million free friendly Parking estimate 2000 described inhabit 500 undescribed headline-grabbing giant Vespa mandarinia formerly Asian news media dubbed murder hornet confuse identify sheet example hidden rodent burrow wall huge 100,00 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Pristiphora rufipes
columbine sawfly
Pristiphora rufipes, commonly known as the columbine sawfly, is a non-native sawfly species native to central Europe that was first discovered in North America in Canada in 1963. It has since spread across the eastern seaboard to the Rocky Mountains, becoming a significant pest of columbine plants (Aquilegia spp.). Adult females use a saw-like ovipositor to insert eggs into leaf tissue, and the caterpillar-like larvae feed on columbine foliage, capable of completely defoliating plants. The species exhibits multiple generations per year in warmer regions, with two generations common in the mid-Atlantic and three or more in England.
sawfly columbine garden-pest defoliator non-native hymenoptera ornamental-pest invasive-insect
Pryeria sinica
euonymus leaf notcher, euonymus defoliator moth
Pryeria sinica is a univoltine zygaenid moth native to East Asia, introduced to the United States in 2002 where it has established populations in Maryland and Virginia. The species is a specialist herbivore of Celastraceae, particularly Euonymus species, where larvae feed gregariously and create distinctive marginal notches on leaves. Adults are diurnal wasp mimics with clear wings and aposematic coloration. The species has been reported more recently in the United Kingdom.
invasive-species pest ornamental-pest univoltine diurnal wasp-mimicry gregarious-larvae chemical-defense cyanogenic Celastraceae-specialist
Pseudaonidia paeoniae
Peony Scale
Pseudaonidia paeoniae, commonly known as the Peony Scale, is a species of armored scale insect in the family Diaspididae. It is a specialized herbivore that feeds on peonies (Paeonia spp.), extracting plant fluids through piercing-sucking mouthparts. The species has been documented across multiple continents including Asia, Europe, and North America, likely distributed through the horticultural trade of its host plants. Like other armored scales, it produces a protective waxy covering (test) that shields the body from predators and environmental conditions.
scale-insect armored-scale pest ornamental-pest peony horticulture invasive-potential