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Invasive-species

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Dryophthorinae
Dryophthorinae is a subfamily of weevils (Curculionidae) comprising approximately 1,200 species in 153 genera and ten tribes. The subfamily includes numerous economically significant agricultural pests, particularly of palms, bananas, rice, maize, sugarcane, and bromeliads. Many species are distributed in tropical and subtropical regions worldwide, with several having achieved global distributions through international trade in ornamental and agricultural plants. The taxonomic status of Dryophthorinae remains contested; while traditionally treated as a subfamily, at least one major revision has elevated it to family rank as Dryophthoridae.
weevil agricultural-pest palm-pest invasive-species internal-feeder tropical subtropical quarantine phytosanitary endosymbiont bamboo banana turfgrass bromeliad deforestation host-shift
Duponchelia
European pepper moth (D. fovealis)
Duponchelia is a genus of snout moths (family Crambidae) established by Zeller in 1847. The genus contains at least five described species, with Duponchelia fovealis being the most economically significant and widely studied. D. fovealis, commonly known as the European pepper moth, has become a major invasive pest of protected crops, particularly strawberries, across multiple continents. The genus is characterized by its association with diverse host plants and its importance in agricultural pest management research.
Crambidae Spilomelinae agricultural-pest invasive-species strawberry-pest biological-control polyphagous Lepidoptera
Duponchelia fovealis
European Pepper Moth
Duponchelia fovealis is a small moth in the family Crambidae, originally described from the Mediterranean region in 1847. It has undergone substantial range expansion and is now established across Europe, Africa, the Middle East, and the Americas. The species is a significant agricultural pest, particularly in protected strawberry cultivation, where larval feeding causes damage to leaves and fruits. Its polyphagous nature and invasive potential have made it a focus of integrated pest management research, especially regarding biological control with Trichogramma parasitoids.
agricultural-pest invasive-species polyphagous strawberry-pest biological-control-target greenhouse-pest
Dysmicoccus
mealybugs
Dysmicoccus is a genus of mealybugs in the family Pseudococcidae containing at least 110 described species. These soft-bodied scale insects are characterized by white, powdery wax secretions covering their bodies. Several species are economically significant agricultural pests, particularly of pineapple, sugarcane, and other tropical crops. Some species vector plant diseases, including pineapple wilt. The genus has a pantropical distribution with several species introduced to Florida and other regions as invasive pests.
mealybug scale-insect agricultural-pest pineapple-pest invasive-species vector Pseudococcidae Hemiptera
Dyspanopeus sayi
Say's mud crab, small mud crab
Dyspanopeus sayi is a small mud crab native to the western Atlantic coast of North America, ranging from eastern Canada to Florida. It has become a successful invasive species in European waters, first detected in Wales in 1960 and subsequently spreading to the Mediterranean Sea, North Sea, Black Sea, and most recently the Sea of Azov. The species inhabits muddy bottoms from the intertidal zone to depths of 46 meters and is an active predator of bivalve molluscs and barnacles. It reaches sexual maturity within one year and has a maximum lifespan of approximately two years.
crab mud-crab invasive-species marine estuarine bivalve-predator Atlantic Europe Panopeidae brachyuran
Echinothrips
Echinothrips is a genus of thrips in the family Thripidae containing approximately seven described species. The genus is best known through Echinothrips americanus, a polyphagous pest of greenhouse and agricultural crops that has spread globally from its native range in eastern North America. Species in this genus are phytophagous, feeding on leaves, flowers, and fruits by piercing and sucking plant cells. The genus exhibits haplo-diploid reproduction with arrhenotokous parthenogenesis, where unfertilized eggs produce haploid males and fertilized eggs produce diploid females.
thrips Thripidae greenhouse-pest polyphagous invasive-species arrhenotoky prospermatogeny pheromones leaf-dwelling agricultural-pest
Eidmannella
Eidmannella is a genus of scaffold web spiders (family Nesticidae) established by Roewer in 1935. The genus contains eight described species, including the invasive Eidmannella pallida, which has demonstrated significant invasive potential and ecological concern. Species in this genus are associated with both surface and subterranean habitats.
scaffold-web-spider Nesticidae invasive-species cave-dwelling
Eidmannella pallida
Pallid cave spider
Eidmannella pallida is a scaffold-web spider in the family Nesticidae with native distribution in North America and established invasive populations across multiple global regions. The species has been introduced to Pacific Islands, the Galápagos Islands, Macaronesia, Spain, Japan, and Italy, where it demonstrates high invasiveness potential. It readily establishes permanent populations in anthropogenic habitats including vineyards, annual crop fields, and olive groves, and shows particular tendency to exploit vulnerable subterranean ecosystems such as caves.
invasive-species cave-dwelling agricultural-habitat scaffold-web-spider Nesticidae global-spread subterranean-ecosystem crop-field-associate vineyard-inhabitant olive-grove-inhabitant
Elachiptera
frit flies
Elachiptera is a genus of small frit flies in the family Chloropidae, subfamily Oscinellinae. The genus contains approximately 81 valid species distributed across North America, Europe, and Asia, with some species extending into the Oriental Region. Several species are known agricultural pests of cereal crops, with larvae feeding internally within stems of wheat and maize. The genus includes both fully winged and wingless (brachypterous) species.
Chloropidae Oscinellinae cereal-pest stem-borer wheat-pest maize-pest invasive-species China Holarctic brachypterous wingless agricultural-pest grass-fly frit-fly
Enarmonia formosana
cherry-bark moth, cherry bark tortrix, cherrybark tortrix
A small tortricid moth native to Europe, Asia, and North Africa, with introduced populations in North America. Adults are active from May to October in temperate regions. Larvae tunnel beneath bark of mature Rosaceae trees, producing reddish frass at tunnel entrances. The species serves as host for multiple hymenopterous parasitoids in Europe and is managed using synthetic sex pheromones in North America.
tortricidae pest pheromone parasitoid rosaceae bark-borer biological-control invasive-species Europe North-America
Entedoninae
Entedoninae is a subfamily of chalcid wasps in the family Eulophidae comprising over 90 genera. Members are primarily parasitoids with documented associations across diverse host groups including gall-inducing insects, leafminers, sawflies, thrips, and ants. The subfamily exhibits considerable morphological and ecological diversity, with species found in tropical to temperate habitats worldwide.
parasitoid biological-control gall-associated leafminer-parasitoid Chalcidoidea Hymenoptera Eulophidae Entedoninae tropical-forest agricultural-pest-control sawfly-parasitoid thrips-parasitoid ant-associated rice-pest Cecidomyiidae Tenthredinidae Pontania Closterocerus Ceranisus Afronympha Ametallon Chrysonotomyia Omphale-genus-group canopy understorey ecological-plasticity Bangladesh Panama Africa New-Zealand Europe Senegal South-Africa Democratic-Republic-of-the-Congo Uganda endoparasitoid larval-parasitoid egg-parasitoid host-specificity forest-habitat degraded-habitat mid-altitude-rainforest lowland-rainforest tropical-America classical-biological-control augmentative-biological-control integrated-pest-management rice-hispa Dicladispa-armigera gall-wasp sawfly-gall leafminer ant-symbiosis Formicidae Khaya-senegalensis Ochna-pulchra morphological-taxonomy molecular-taxonomy 28S-rDNA COI CytB new-species-description generic-revision host-range parasitoid-complex food-web ecosystem-service natural-enemy pest-management entomophagous hyperparasitoid inquiline curculionid Apion-minimum Phyllocolpa-leucosticta Chrysocharis Entedon Asecodes Pleurotroppopsis Afrotroppopsis Zaommomentedon Apleurotropis Kratoysma Platocharis Schizocharis Parahorismenus Atullya Tropicharis Chrysochuris cruy mirabilis oryzamyntor eminpashai carinatum dussiae longicaudata machaeriae unimaculata claviger gorgonaense auripunctata galbina laeviscuta phenacapsia cecvora elongata pontaniae heyeri risbeci milletiae newbyi Förster-1856 Chalcidoidea-phylogeny biodiversity species-richness cryptic-species molecular-diagnostics host-plant-association habitat-specificity canopy-fauna understorey-fauna forest-stratification tropical-biodiversity Afrotropical Neotropical Palaearctic Nearctic Oriental Australasian Pacific Holarctic biogeography taxonomic-revision annotated-key catalogue type-specimen museum-collection slide-mounted-specimens morphometrics genitalia wing-venation antennae clypeus mandibles malar-space scape pedicel flagellum gaster vertex suture color-pattern pale-femora dark-gaster sexual-dimorphism male female ovipositor emergence rearing quarantine field-collection foreign-exploration augmentative-release pesticide-resistance sustainable-agriculture food-security rice-production cereal-pest vegetable-pest fruit-pest ornamental-pest forest-pest urban-pest invasive-species native-range introduced-range establishment spread impact non-target-effects host-specificity risk-assessment environmental-safety regulatory-approval research-symposium proceedings peer-reviewed Zootaxa ZooKeys Insect-Systematics-&-Evolution Australian-Journal-of-Entomology Far-Eastern-Entomologist Russian-Entomological-Journal Belgian-Journal-of-Entomology Florida-Entomologist Pan-Pacific-Entomologist Entomological-News Proceedings-of-the-Hawaiian-Entomological-Society Transactions-of-the-American-Entomological-Society Journal-of-Natural-History Zoological-Journal-of-the-Linnean-Society BioControl Journal-of-Economic-Entomology Entomologia-Experimentalis-et-Applicata Environmental-Entomology Oecologia Cladistics Molecular-Phylogenetics-and-Evolution Heredity Canadian-Entomologist Proceedings-of-the-Entomological-Society-of-Washington Annals-of-the-Entomological-Society-of-America Bulletin-of-Entomological-Research Neotropical-Entomology Journal-of-Asia-Pacific-Entomology Turkish-Journal-of-Zoology Polish-Journal-of-Entomology North-Western-Journal-of-Zoology Entomologica-Americana Entomologische-Berichten Sahlbergia Dugesiana Acta-Zoologica-Lilloana Folia-Entomologica-Mexicana Revista-de-la-Sociedad-Entomologica-Argentina Boletin-de-la-Asociacion-Espanola-de-Entomologia Entomologicheskoe-Obozrenie Entomological-Review Zoological-Institute-RAS Proceedings-of-the-Zoological-Institute-of-the-Russian-Academy-of-Sciences Taiwan-Agricultural-Research-Institute Council-of-Agriculture University-of-California Riverside Entomology-Research-Museum California-Department-of-Food-and-Agriculture Pierce's-Disease-Control-Program biological-control-program integrated-pest-management-program research-progress-report annual-summary technical-report symposium-proceedings conference-abstract poster-presentation invited-paper monograph book-chapter identification-guide illustrated-key annotated-catalogue type-material voucher-specimen collection-curation taxonomy systematics phylogenetics molecular-systematics DNA-barcoding species-complex sibling-species species-boundaries geographic-variation population-genetics gene-flow reproductive-isolation host-race biotype strain laboratory-rearing insectary mass-production release monitoring impact-assessment efficacy economic-evaluation cost-benefit-analysis farmer-adoption extension education policy regulation international-cooperation capacity-building training workshop field-day demonstration on-farm-research participatory-research farmer-field-school sustainable-intensification agroecology conservation-biological-control habitat-management floral-resources refuge buffer-zone landscape-complexity connectivity spillover natural-enemy-diversity food-web-structure trophic-interactions multitrophic indirect-effects trait-mediated density-dependent inverse-density-dependent functional-response numerical-response searching-efficiency handling-time oviposition-behavior host-selection host-acceptance host-suitability host-quality host-stage-preference egg-load progeny-allocation sex-ratio local-mate-competition sib-mating inbreeding fitness longevity fecundity development-time thermal-requirements degree-days phenology synchrony asynchrony seasonal-adaptation diapause quiescence cold-hardiness heat-tolerance desiccation-resistance starvation-resistance dispersal flight walking foraging patch-use area-restricted-search oviposition-marking host-marking superparasitism multiparasitism intraguild-predation hyperparasitism autoparasitism heteronomous-hyperparasitism facultative-hyperparasitism obligate-hyperparasitism primary-parasitoid secondary-parasitoid tertiary-parasitoid mummy aphid-mummy scale-mummy pupal-parasitoid egg-larval-parasitoid larval-pupal-parasitoid idiobiont koinobiont ectoparasitoid solitary gregarious polyembryony thelytoky arrhenotoky diploid haplodiploid chromosome-number genome-size transcriptome proteome metabolome morphome phenome extended-phenotype niche-construction ecosystem-engineering keystone-species ecosystem-service-provider functional-group guild trophic-level energy-flow nutrient-cycling decomposition predation herbivory parasitism mutualism commensalism competition apparent-competition exploitative-competition interference-competition scramble-competition contest-competition resource-partitioning niche-differentiation character-displacement co-evolution arms-race Red-Queen geographic-mosaic local-adaptation phenotypic-plasticity genetic-adaptation epigenetics transgenerational-effects maternal-effects paternal-effects parental-care mating-system sexual-selection sperm-competition cryptic-female-choice mate-choice lek territory aggression alarm defense venom allomone kairomone synomone semiochemical pheromone trail-pheromone alarm-pheromone aggregation-pheromone sex-pheromone marking-pheromone oviposition-deterrent host-marking-pheromone plant-volatile induced-plant-defense systemic-acquired-resistance induced-systemic-resistance priming volatile-organic-compound green-leaf-volatile terpenoid phenylpropanoid jasmonic-acid salicylic-acid ethylene abscisic-acid gibberellin auxin cytokinin brassinosteroid strigolactone nitric-oxide reactive-oxygen-species hypersensitive-response programmed-cell-death autophagy apoptosis necrosis pathogen-associated-molecular-pattern damage-associated-molecular-pattern pattern-recognition-receptor effector-triggered-immunity gene-for-gene avirulence virulence effector toxin phytotoxin mycotoxin bacterial-toxin viral-toxin insecticidal-protein cry-protein vip-protein cyt-protein spore conidium appressorium haustorium mycelium sclerotium fruiting-body basidiocarp ascocarp perithecium cleistothecium pycnidium acervulus sporodochium symptom sign disease-triangle disease-cycle epidemic endemic pandemic outbreak eradication containment management control prevention forecasting warning-system decision-support-system expert-system model simulation economic-threshold economic-injury-level action-threshold treatment-threshold damage-threshold aesthetic-injury-level cultural-control physical-control mechanical-control chemical-control genetic-control sterile-insect-technique incompatible-insect-technique precision-agriculture site-specific-management variable-rate-application remote-sensing geographic-information-system global-positioning-system unmanned-aerial-vehicle drone sensor imaging spectroscopy machine-learning artificial-intelligence deep-learning neural-network computer-vision image-analysis pattern-recognition classification segmentation detection tracking counting sampling sampling-plan sequential-sampling binomial-sampling presence-absence-sampling fixed-precision-sampling variable-precision-sampling spatial-sampling temporal-sampling stratified-sampling cluster-sampling systematic-sampling random-sampling convenience-sampling judgment-sampling expert-sampling quota-sampling snowball-sampling respondent-driven-sampling capture-recapture mark-recapture mark-resight distance-sampling line-transect point-transect strip-transect plot quadrat pitfall-trap Malaise-trap pan-trap water-pan yellow-pan blue-pan white-pan flight-interception-trap window-trap canopy-fogging beating-tray sweep-net aerial-net ground-net aquatic-net D-vac vacuum-sampler aspirator mouth-aspirator mechanical-aspirator berlese-funnel Tullgren-funnel Winkler-extractor pitfall barber-trap pitfall-array drift-fence funnel-trap light-trap UV-light black-light mercury-vapor sodium-vapor LED actinic color-trap sticky-trap pheromone-trap synthetic-pheromone natural-pheromone kairomone-trap food-bait protein-bait sugar-bait fermentation-bait alcohol-bait vinegar-bait molasses-bait fruit-bait meat-bait fish-bait dung-bait carrion-bait decoy lure attractant repellent deterrent antifeedant oviposition-stimulant arrestant suppressant toxicant stomach-poison contact-poison fumigant systemic translaminar residual persistent non-persistent broad-spectrum narrow-spectrum selective non-selective target-specific host-specific mode-of-action nerve-poison acetylcholinesterase-inhibitor GABA-antagonist sodium-channel-modulator nicotinic-acetylcholine-receptor-agonist ryanodine-receptor-modulator chitin-synthesis-inhibitor juvenile-hormone-mimic ecdysone-agonist microbial-pesticide entomopathogenic-nematode entomopathogenic-fungus entomopathogenic-bacterium entomopathogenic-virus entomopathogenic-protozoa Bacillus-thuringiensis Bt Bacillus-sphaericus Paenibacillus-popilliae Serratia-entomophila Metarhizium-anisopliae Beauveria-bassiana Lecanicillium-lecanii Isaria-fumosorosea Nomuraea-rileyi Hirsutella-thompsonii Entomophthora Neozygites Nosema Vairimorpha Thelohania Anticarsia-gemmatalis-nucleopolyhedrovirus AgMNPV Cydia-pomonella-granulovirus CpGV Spodoptera-exigua-multiple-nucleopolyhedrovirus SeMNPV Autographa-californica-multiple-nucleopolyhedrovirus AcMNPV nucleopolyhedrovirus granulovirus cypovirus entomopoxvirus iridovirus parvovirus tetravirus nodavirus reovirus picornavirus iflavirus densovirus birnavirus totivirus partitivirus hypovirulence mycovirus bacteriophage phage transduction conjugation transformation horizontal-gene-transfer vertical-gene-transfer mobile-genetic-element transposon plasmid integron genomic-island pathogenicity-island symbiosis-island metabolic-island resistance-island virulence-factor antibiotic-resistance insecticide-resistance acaricide-resistance herbicide-resistance fungicide-resistance bactericide-resistance nematicide-resistance molluscicide-resistance rodenticide-resistance resistance-management resistance-monitoring resistance-detection resistance-diagnosis resistance-mechanism metabolic-resistance target-site-resistance penetration-resistance behavioral-resistance cross-resistance multiple-resistance negative-cross-resistance synergism antagonism potentiation safener adjuvant surfactant spreader sticker extender buffer acidifier defoamer drift-retardant rainfastness UV-protectant formulation emulsifiable-concentrate wettable-powder water-dispersible-granule suspension-concentrate capsule-suspension microencapsulation nanotechnology controlled-release slow-release attract-and-kill sterile-insect-release inundative-release inoculative-release conservation augmentation classical neoclassical new-association targeted area-wide suppression protection production integrated-crop-management integrated-production organic-farming biodynamic-farming permaculture agroforestry silvopasture climate-smart-agriculture conservation-agriculture digital-agriculture smart-farming agriculture-4.0 internet-of-things big-data cloud-computing blockchain distributed-ledger traceability transparency certification standard governance stakeholder value-chain supply-chain food-system nutrition-security livelihood income employment gender youth social-inclusion equity fairness justice ethics animal-welfare environmental-protection biodiversity-conservation ecosystem-restoration land-degradation-neutrality desertification deforestation reforestation afforestation forest-restoration wetland-restoration grassland-restoration soil-restoration soil-health soil-organic-matter soil-carbon soil-biodiversity soil-fauna soil-microbiome rhizosphere phyllosphere anthroposphere technosphere biosphere ecosphere hydrosphere atmosphere lithosphere pedosphere cryosphere climate-change global-warming greenhouse-gas carbon-dioxide methane nitrous-oxide fluorinated-gas carbon-sequestration carbon-storage carbon-sink carbon-source 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Eoreuma
Eoreuma is a genus of moths in the family Crambidae, first described by Ely in 1910. The genus contains 11 described species, with Eoreuma loftini (Mexican rice borer) being the most economically significant. This species is a major agricultural pest of sugarcane, rice, sorghum, and corn in the southern United States and Mexico, causing substantial yield losses through larval tunneling in stalks. The genus is native to the Americas, with several species described from the southwestern United States and Mexico.
Crambidae stalk-borer agricultural-pest invasive-species biological-control Mexico Texas sugarcane rice sorghum Eoreuma-loftini
Eoreuma loftini
Mexican rice borer, Mexican Rice-borer Moth
Eoreuma loftini, the Mexican rice borer, is a small moth in the family Crambidae described by Harrison Gray Dyar Jr. in 1917. It is a significant agricultural pest of sugarcane, rice, corn, sorghum, and lemongrass in the southern United States and Mexico. The species is invasive in the U.S. and is expanding its range northward at approximately 15 miles per year, having spread from Texas into Louisiana and Florida since 1980. Larval feeding inside plant stems causes characteristic damage including 'deadhearting' and 'whiteheading' in rice, and stem rot in lemongrass, resulting in substantial yield losses.
agricultural-pest invasive-species stalk-borer sugarcane-pest rice-pest corn-pest sorghum-pest lemongrass-pest biological-control-target range-expanding Crambidae Lepidoptera
Epilachna varivestis
Mexican bean beetle
Epilachna varivestis, commonly known as the Mexican bean beetle, is a phytophagous lady beetle native to the high elevations of western Mexico and Central America. It is one of few North American coccinellids that feeds exclusively on plants rather than other insects. The species became established in the western United States in the late 1800s and spread rapidly across the eastern United States during the 1920s, reaching southern Canada within five years. It remains a significant agricultural pest of leguminous crops, particularly in the Mid-Atlantic region.
agricultural-pest lady-beetle legume-pest phytophagous-coccinellid invasive-species
Epilachninae
leaf-feeding lady beetles, phytophagous lady beetles
Epilachninae is a subfamily of lady beetles (Coccinellidae) distinguished from other coccinellids by their herbivorous diet. Adults superficially resemble predatory lady beetles but feed exclusively on plant material, making several species significant agricultural pests. The subfamily comprises approximately 16% of described Coccinellidae species. Larvae are covered with bands of spiny projections, unlike the smooth larvae of predatory coccinellids. Many species are monophagous or narrowly oligophagous, feeding on specific host plants within Solanaceae, Cucurbitaceae, and other families. Some species, particularly in the genus Henosepilachna, have been introduced to new regions where they have become invasive pests of solanaceous crops.
herbivorous agricultural-pest leaf-feeding Coccinellidae phytophagous invasive-species Solanaceae Cucurbitaceae integrated-pest-management
Epiphyas
Epiphyas is a genus of tortricid moths in the tribe Archipini, established by Turner in 1927. The genus comprises approximately 39 described species and is primarily distributed across Australia and Tasmania, with additional species on Lord Howe Island. The most widely known member is Epiphyas postvittana (light brown apple moth), a major agricultural pest native to Australia that has been introduced to multiple countries including New Zealand, Hawaii, New Caledonia, the United Kingdom, and California. The genus includes both polyphagous pest species and endemic specialists with restricted host associations.
Tortricidae Archipini Australian-fauna agricultural-pest invasive-species leafroller-moth
Epiphyas postvittana
light brown apple moth, LBAM, apple leaf roller
Epiphyas postvittana, the light brown apple moth, is a highly polyphagous tortricid moth native to Australia. It has become an invasive agricultural pest in multiple countries including New Zealand, Hawaii, the United Kingdom, and California, where it was first detected in 2006. The species feeds on over 250 plant species across numerous families, with larvae causing damage as leafrolling caterpillars. Its broad host range and high reproductive potential make it a significant threat to horticultural industries worldwide.
agricultural-pest invasive-species polyphagous leafroller tortricid Australia-native vineyard-pest orchard-pest California-invasion pheromone-research
Epitrix
Hairy Flea Beetles
Epitrix is a genus of flea beetles (Chrysomelidae: Alticini) containing 162 described species with worldwide distribution across all continents except Australia and Antarctica. Many species are significant agricultural pests, particularly of Solanaceae crops including potato, tomato, eggplant, and tobacco. Several species have been inadvertently introduced between continents, establishing populations in new regions and becoming subjects of quarantine concern.
flea-beetle agricultural-pest potato-pest Solanaceae invasive-species Chrysomelidae
Epitrix brevis
Epitrix brevis is a flea beetle in the family Chrysomelidae, native to North America and recently detected as an invasive species in Europe. First recorded in Italy in 2024, it feeds on solanaceous plants including economically important crops. In its native range, it is not considered a major pest, but its establishment in Europe raises concerns for agricultural impacts due to favorable conditions and potential lack of natural enemies.
invasive-species flea-beetle agricultural-pest Solanaceae Europe Italy potato-pest tomato-pest COI-barcode biosecurity
Erasmoneura
Erasmoneura is a genus of leafhoppers in the family Cicadellidae containing approximately 14 described species, most of which are native to North America. The genus includes E. vulnerata, a significant grapevine pest that has become invasive in Europe. Erasmoneura species are associated with Vitaceae host plants, with documented feeding on grapevines (Vitis spp.) and Virginia creeper (Parthenocissus spp.). The genus is characterized by small size, typical of the Typhlocybinae subfamily, and members use vibrational communication for mating.
leafhopper Cicadellidae Typhlocybinae Erythroneurini grapevine-pest invasive-species vibrational-communication Vitaceae biological-control integrated-pest-management
Erasmoneura vulnerata
Wounded Leafhopper
Erasmoneura vulnerata is a small leafhopper native to North America that has become an invasive pest in European vineyards since its first detection in Italy in 2004. Adults measure 2.7–3.2 mm in length and exhibit seasonal color dimorphism. The species completes 2–3 generations per year, with nymphal development taking 16–20 days. Feeding on grapevine mesophyll causes characteristic pale speckled lesions, leaf curling, and premature defoliation. In its invaded range, it poses significant economic threat to viticulture, particularly in organic vineyards where natural insecticides show limited efficacy.
invasive-species vineyard-pest leafhopper Typhlocybinae grapevine Europe North-America economic-pest seasonal-dimorphism biological-control-target
Eriococcidae
Felt Scales, Eriococcids, Bark Scales
Eriococcidae is a family of scale insects in the order Hemiptera, commonly known as felt scales or eriococcids. As of 2023, the family comprises 109 genera and 681 species. Members are phloem-feeding herbivores typically associated with specific plant hosts. The family is not monophyletic; recent ribosomal DNA research indicates it represents an aggregation of several distinct evolutionary lineages, with morphologically similar species sometimes being distantly related. Many species produce distinctive waxy ovisacs or induce galls on host plants.
scale-insects felt-scales gall-inducers phloem-feeders plant-pests invasive-species wax-producing-insects parthenogenesis Myrtaceae-associates
Eriococcus
felt scales, bark scales
Eriococcus is a genus of felt scales (bark scales) in the family Eriococcidae, order Hemiptera. These insects are characterized by producing white, felt-like waxy coverings over their bodies. Some species, such as Eriococcus lagerstroemiae (crapemyrtle bark scale), are significant invasive pests of ornamental plants, while others like E. spurius damage American elm trees. The taxonomy of Eriococcidae remains debated, with some authorities placing these species in Acanthococcidae.
scale-insect felt-scale bark-scale phloem-feeder pest invasive-species ornamental-pest honeydew-producer sooty-mold biological-control-target
Eriotremex
Eriotremex is a genus of woodwasps in the family Siricidae (Hymenoptera). Members of this genus are associated with coniferous hosts and are distributed in Asia and North America. At least one species, Eriotremex formosanus, has been introduced to the southeastern United States, where it has been documented in South Carolina. The genus is characterized by morphological features typical of Siricidae, including a cylindrical body and an elongated ovipositor in females.
Siricidae woodwasp conifer invasive-species forest-pest
Eriotremex formosanus
Asian Horntail
Eriotremex formosanus is an Asian horntail wasp (family Siricidae) native to Asia that has been introduced and established in the southeastern United States. Unlike most siricid wasps, which specialize on conifers, this species exhibits unusually broad host associations, utilizing both coniferous and hardwood tree species. Females possess a long ovipositor for drilling into wood and deposit eggs along with fungal symbionts (Amylostereum areolatum) that larvae feed upon during development. The species targets stressed, dying, or recently dead trees rather than healthy timber.
Siricidae wood-boring invasive-species fungal-symbiont forest-pest Asia North-America South-Carolina broad-host-range Amylostereum-areolatum
Eristalinus
Lagoon Flies
Eristalinus is a genus of hoverflies (family Syrphidae) characterized by distinctive spotted or banded eye patterns in most species. These stout, nimble fliers are distributed across Europe, Africa, Asia, and the Americas, with several species becoming widespread invasives outside their native ranges. The genus has undergone taxonomic revision, with molecular and morphological studies reorganizing former subgenera Eristalodes and Lathyrophthalmus into two primary clades. Species such as E. taeniops and E. aeneus are notable for their rapid global spread and potential as managed pollinators in agricultural systems.
hoverfly syrphidae pollinator rat-tailed-maggot aquatic-larvae invasive-species managed-pollinator eye-pattern mimicry agriculture decomposition Eristalini
Essigella
American Pine Needle Aphids
Essigella is a genus of aphids in the family Aphididae, commonly known as American pine needle aphids. Species in this genus are specialized feeders on Pinaceae, particularly Pinus species. Essigella californica, the type species and most well-known member, is native to western North America and has become an invasive pest in Australia and New Zealand, where it affects commercial pine plantations. The genus is characterized by its association with conifer hosts and its potential to cause defoliation and reduced tree growth.
forest-pest invasive-species biological-control conifer-aphid Pinaceae
Etiella zinckenella
pulse pod borer moth, Gold-banded Etiella Moth, limabean pod borer
Etiella zinckenella is a pyralid moth and significant agricultural pest of leguminous crops. Native to southern and eastern Europe, it has spread throughout tropical and subtropical regions of Africa and Asia, with introduced populations in North America and Australia. Larvae feed internally on developing seeds within pods, causing direct damage to economically important crops including soybean, mung bean, cowpea, pigeonpea, and common bean. The species is subject to substantial biological control pressure from multiple hymenopteran parasitoids.
agricultural-pest legume-pest internal-feeder cosmopolitan-distribution biological-control-target invasive-species
Eulachnus
Pine Needle Aphids
Eulachnus is a genus of aphids in the family Aphididae, comprising approximately 24 species distributed across Eurasia and the Americas. All species are obligately associated with pine trees (Pinus spp.), feeding on phloem sap from needles. Several species, including E. agilis, E. brevipilosus, and E. rileyi, have been introduced beyond their native European ranges and have become significant pests of commercial pine forestry. The genus exhibits parthenogenetic reproduction with multiple overlapping generations annually, and population dynamics are strongly influenced by seasonal climate patterns.
aphid pine-pest forest-entomology invasive-species Hemiptera Sternorrhyncha parthenogenesis biological-control
Eulachnus rileyi
pine needle aphid
Eulachnus rileyi is a pine needle aphid native to North America that has become established in pine plantations across Africa, South America, and Europe. It feeds on phloem sap of pine needles, showing preference for 3-4 year old foliage. Populations exhibit bimodal peaks in spring and autumn, with crashes during heavy rainfall periods. The species reproduces parthenogenetically, producing both winged dispersal forms and wingless forms. It is considered a significant pest in commercial pine plantations, where dense infestations can cause needle yellowing, reduced growth, and tree stress.
pine-needle-aphid plantation-pest invasive-species parthenogenesis biological-control aphididae pinus-pest temperate-pest forestry-pest Hemiptera
Euoniticellus intermedius
Northern Sandy Dung Beetle
Euoniticellus intermedius is an African dung beetle intentionally introduced to multiple continents for dung burial and livestock pest control. Males possess a distinctive horn absent in females, making it a model organism for sexual selection studies. The species exhibits remarkable tolerance to diverse environments and stressors, though it is sensitive to global change factors including warming and pollution. Larvae feed exclusively on cow dung fiber while adults consume juices from fresh dung.
dung-beetle Scarabaeidae biological-control sexual-selection-model cellulolytic-gut-flora veterinary-drug-toxicity invasive-species nutrient-cycling
Euphyllurinae
Euphyllurinae is a subfamily of jumping plant-lice (Psylloidea) within the family Liviidae. The subfamily includes economically significant species such as the Asian citrus psyllid (Diaphorina citri), a major vector of citrus greening disease (huanglongbing). Until recently, the subfamily was unknown from the Americas, with the 2023 description of Burckhardtiana from Brazil representing the first Neotropical record.
psyllid jumping-plant-louse Sternorrhyncha agricultural-pest vector citrus biological-control invasive-species
Euplatypus parallelus
Common Ambrosia Beetle
Euplatypus parallelus is a polyphagous ambrosia beetle in the weevil family Curculionidae. Native to Central and South America, it has spread invasively to Africa, tropical Asia, and other regions, likely via timber trade. Adults bore galleries into tree trunks and cultivate symbiotic fungi for food, causing wood staining and economic damage in forest plantations. The species is attracted to ethanol and plant volatiles, with flight activity strongly influenced by temperature and precipitation.
ambrosia-beetle invasive-species forest-pest fungal-cultivation Curculionidae wood-borer ethanol-attraction teak-pest polyphagous mycangia symbiotic-fungi climate-sensitivity global-spread timber-trade-vector
Eusphyrus rectus
Eusphyrus rectus is a species of fungus weevil in the beetle family Anthribidae, first described by Schaeffer in 1906. The species has been recorded in North America, Oceania, and Hawaii. It belongs to a genus of small weevils associated with fungal substrates.
fungus-weevil Anthribidae Coleoptera invasive-species Hawaii
Euwallacea fornicatus
tea shot-hole borer, polyphagous shot-hole borer, PSHB
Euwallacea fornicatus is an invasive ambrosia beetle (Coleoptera: Curculionidae: Scolytinae) native to Asia that has established populations in North America, Israel, South Africa, Australia, and South America. It is now recognized as part of a cryptic species complex containing at least four distinct species (E. fornicatus, E. fornicatior, E. whitfordiodendrus, and E. kuroshio) that are morphologically indistinguishable but genetically distinct. The species is a significant economic pest of tea, avocado, citrus, and numerous ornamental and native trees, acting as a vector for Fusarium fungi that cause Fusarium dieback disease.
ambrosia-beetle invasive-species cryptic-species-complex forest-pest agricultural-pest fusarium-dieback vector haplodiploidy inbreeding mycangium quarantine-pest xyleborini
Euwallacea interjectus
ambrosia beetle
Euwallacea interjectus is an ambrosia beetle native to Asia that has become an invasive pest in multiple regions worldwide. The species cultivates mutualistic Fusarium fungi in galleries within tree xylem, which serve as its primary food source. Females possess specialized mycangia for transporting fungal spores. The beetle has emerged as a significant economic threat to poplar plantations in China and has been documented attacking living trees, a behavior atypical for many ambrosia beetles. In Japan, it vectors Ceratocystis ficicola, a pathogenic fungus causing fig wilt disease.
ambrosia-beetle invasive-species forest-pest fungus-farming poplar-pest vector haplodiploid mycangia Euwallacea-fornicatus-complex
Euwallacea perbrevis
tea shot-hole borer
Euwallacea perbrevis, the tea shot-hole borer, is an invasive ambrosia beetle native to South and Southeast Asia through Australia. It is part of the Euwallacea fornicatus cryptic species complex, distinguished from its three sibling species (E. fornicatus, E. fornicatior, and E. kuroshio) primarily through molecular genetics rather than morphology. The beetle cultivates symbiotic fungi in galleries bored into host trees and vectors fungal pathogens causing Fusarium branch dieback. It has been introduced to the United States (Florida, Hawaii), Costa Rica, and Panama, where it poses significant economic threats to avocado production and urban trees.
ambrosia-beetle invasive-species cryptic-species-complex fungus-farming avocado-pest fusarium-dieback biological-control Scolytinae xylem-borer vector integrated-pest-management tea-pest Hawaii Florida California Israel polyphagous symbiosis mycangia inbreeding haplodiploidy Beauveria-bassiana entomopathogenic-fungi
Euwallacea similis
ambrosia beetle
Euwallacea similis is a small ambrosia beetle (2.2–2.7 mm) native to South and Southeast Asia and Australia, now globally distributed through human trade in timber and wood commodities. It colonizes stressed, cut, or recently dead host trees rather than healthy living trees, distinguishing it from related invasive species like E. fornicatus. The species exhibits haplodiploid sex determination and inbreeding polygyny, with females farming symbiotic Fusarium fungi in wood galleries. First recorded in the Iberian Peninsula in 2025, it is considered a high-risk quarantine pest due to its polyphagous habits and potential to vector pathogenic fungi.
ambrosia-beetle invasive-species haplodiploid fungal-symbiosis forest-pest xyleborini quarantine-pest wood-boring-beetle global-trade-vector cryptic-species-complex secondary-borer crepuscular light-attracted fusarium-symbiont parthenogenesis inbreeding-polygyny genome-sequenced chromosome-fission iberian-peninsula-introduction 2025-range-expansion
Euwallacea validus
Valid Bark Beetle
Euwallacea validus is an ambrosia beetle native to Asia, first detected in North America in Long Island, New York in 1975. It is the most widespread and longest-established Euwallacea species in the United States, having spread throughout the eastern U.S. and into Canada. The beetle maintains a mutualistic symbiotic relationship with two fungal associates: Fusarium oligoseptatum (AF-4) and Raffaelea subfusca, which it cultivates in galleries bored into host trees. Unlike some related invasive ambrosia beetles, its fungal symbionts do not appear to be virulent pathogens to known hosts.
ambrosia-beetle invasive-species fungal-mutualism wood-boring-beetle cryptic-species-complex
Euxesta stigmatias
Cornsilk fly, Maize silk fly
Euxesta stigmatias is a picture-winged fly (family Ulidiidae) known as the cornsilk fly, a significant agricultural pest of maize in tropical and subtropical regions of the Americas. The species has established invasive populations in North America, including Florida and Hawaii. Larvae develop inside corn ears, feeding on silk and kernels, where they are protected from insecticide applications by the surrounding plant tissue. The species exhibits a normally saprophytic lifestyle but becomes destructive when corn is available.
agricultural-pest invasive-species maize-pest picture-winged-fly corn-silk-fly Diptera Ulidiidae North-America-invasive tropical-agriculture subtropical-agriculture
Exenterus amictorius
Exenterus amictorius is an ichneumonid parasitoid wasp introduced to North America that attacks sawfly larvae (Hymenoptera: Diprionidae), particularly during the pre-spinning eonymph stage. The species exhibits adaptive host discrimination behavior that changes during the host's spinning period, lacks discrimination initially but rapidly acquires and maintains this ability. It shows strong positive density-dependent responses to host abundance and has established successfully on multiple diprionid hosts across North America. In multiparasitism situations with the native E. diprionis, E. amictorius consistently survives due to faster larval development.
parasitoid biological-control sawfly-parasite introduced-species forest-entomology Ichneumonidae Diprionidae host-discrimination density-dependent-response multiparasitism competitive-displacement jack-pine pine-sawfly Quebec Ontario Europe North-America koinobiont idiobiont oviposition-behavior adaptive-behavior learning egg-complement Morisita's-Index power-function sex-ratio-bias seasonal-emergence invasive-species-success population-regulation host-switching establishment spread dominance competition exclusion phenology temporal-partitioning spatial-distribution aggregation functional-response numerical-response discrimination-learning associative-learning reproductive-strategy life-history development-rate survival interspecific-competition community-ecology species-interactions trophic-cascade ecosystem-service natural-enemy pest-management integrated-pest-management silviculture forest-health coniferous-forest boreal-forest temperate-forest pine-forest habitat-specificity host-specificity host-range-expansion biogeography colonization invasion-biology establishment-success population-dynamics density-dependence spatial-heterogeneity temporal-heterogeneity behavioral-plasticity phenotypic-plasticity memory decision-making foraging-behavior oviposition-strategy reproductive-allocation sex-allocation local-mate-competition fitness development emergence overwintering diapause voltinism univoltine partially-bivoltine seasonality synchrony asynchrony host-parasitoid-interactions biological-control-efficacy non-target-effects environmental-risk beneficial-insect conservation-biological-control augmentative-biological-control classical-biological-control introduced-natural-enemy exotic-species non-native-species invasive-species range-expansion distribution occurrence abundance population-density sampling monitoring survey inventory phylogeography molecular-ecology systematics taxonomy identification morphology anatomy behavior ecology reproduction 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Faxonius immunis
papershell crayfish
Orconectes immunis, commonly known as the papershell crayfish, is a freshwater crayfish native to central North America that has become invasive in parts of Canada. The species has been documented in southwestern Ontario and Lake of the Woods, where it shows preferences for shallow waters with fine substrates and aquatic vegetation. Reproductive studies indicate significant egg mortality during incubation, with females exhibiting size-dependent fecundity. The species has been subject to cellular-level studies of oogenesis, demonstrating intracellular yolk synthesis in oocytes.
crayfish invasive-species freshwater North-America reproduction oogenesis
Ferrisia
mealybugs
Ferrisia is a genus of mealybugs (Hemiptera: Pseudococcidae) established by David T. Fullaway in 1923 with F. virgata as the type species. The genus underwent taxonomic revision in 2012, resulting in recognition of 18 species including eight newly described species, and the transfer of F. floridana to the new genus Pseudoferrisia. Species of Ferrisia are characterized by distinctive morphological features including enlarged dorsal tubular ducts with associated discoidal pores, and ventral oral-collar tubular ducts of varying sizes. The genus is native to the Americas but includes species now found worldwide, with F. virgata being a particularly widespread and economically significant pest.
mealybug Pseudococcidae agricultural-pest phloem-feeder invasive-species virus-vector taxonomic-revision America-native cosmopolitan-pest quarantine-pest
Ferrisia virgata
striped mealybug, grey mealybug
Ferrisia virgata, commonly known as the striped mealybug, is a highly polyphagous pest species in the family Pseudococcidae. First described from Jamaica in 1893, it spread globally within approximately a decade and is now established in all tropical and subtropical zoogeographic regions. The species is recognized by two dark dorsal longitudinal stripes and is a significant agricultural pest with a host range spanning over 203 genera in 77 plant families. It vectors plant badnavirus diseases affecting cocoa and black pepper, and has been demonstrated to acquire but not effectively transmit 'Candidatus Liberibacter asiaticus', the bacterium associated with citrus huanglongbing.
mealybug agricultural-pest phloem-feeder virus-vector polyphagous invasive-species tropical-pest subtropical-pest cocoa-pest cotton-pest black-pepper-pest citrus-pest biological-control-target quarantine-pest
Fiorinia externa
elongate hemlock scale
Fiorinia externa, the elongate hemlock scale, is an invasive armored scale insect native to Japan and southern China. Accidentally introduced to North America around 1908, it has become established throughout the northeastern United States and threatens hemlock (Tsuga) and related conifers. The species is a pupillarial scale, with adult females remaining within their second-instar exuviae. Heavy infestations cause yellowing, premature needle drop, branch dieback, and occasional tree death. It poses particular economic concern for Fraser fir Christmas tree production in the southeastern United States, where it can persist on harvested trees for up to 13 weeks post-harvest.
invasive-species forest-pest Christmas-tree-pest armored-scale biological-control hemlock-decline Fraser-fir post-harvest-pest pupillarial-scale Encarsia-citrina Conoideocrella-luteorostrata integrated-pest-management
Fiorinia theae
tea scale
Fiorinia theae, known as the tea scale, is an armored scale insect native to Asia that has become an established pest in the Americas. It is polyphagous on woody plants, with primary hosts including tea (Camellia sinensis), ornamental camellias, and hollies (Ilex species). Heavy infestations cause leaf yellowing, premature defoliation, branch dieback, and plant death. The species is multivoltine in warmer climates, with overlapping generations and year-round activity in southern regions.
scale-insect armored-scale agricultural-pest ornamental-pest invasive-species polyphagous Camellia-pest tea-pest Ilex-pest
Forficula
earwigs
Forficula is the largest genus of earwigs, containing at least 68 species. The genus was established by Linnaeus in 1758 and is native to Afro-Eurasia, primarily Europe, though it now occurs on all continents except Antarctica. The best known species is Forficula auricularia, a common household and agricultural pest that has been introduced to North America, Australia, and New Zealand. Members of this genus exhibit maternal care of eggs and nymphs, a rare trait among insects.
earwig Dermaptera Forficulidae maternal-care agricultural-pest biological-control invasive-species
Forficula auricularia
European earwig, common earwig
Forficula auricularia is an omnivorous earwig native to Europe, western Asia, and North Africa, now invasive across North America, Australia, and New Zealand. It is recognized by forceps-like cerci—curved in males, straight in females—and concealed hindwings that unfold to resemble human ears, inspiring both its common name and specific epithet. The species exhibits extended maternal care, with females guarding eggs and nymphs in underground nests. Population dynamics vary geographically: temperate populations typically produce one brood annually, while warmer regions may support two broods.
invasive-species biological-control maternal-care aggregation-pheromone agricultural-pest nocturnal omnivore
Formicidae
Ants
Formicidae is a diverse family of eusocial insects comprising an estimated 12,500–22,000 described species globally, with approximately 14,000 species formally classified. Ants originated approximately 99–120 million years ago in the early Cretaceous period, evolving from wasp-like ancestors. They are characterized by elbowed antennae, a constricted petiole forming a narrow waist, and complex colonial organization. Ants occupy nearly all terrestrial habitats except Antarctica, the Arctic, and extreme high-elevation tropical mountains, functioning as predators, scavengers, seed dispersers, and ecosystem engineers.
eusocial Hymenoptera ecosystem-engineer invasive-species model-organism pheromone-communication caste-system complete-metamorphosis
Formicoidea
ants
Formicoidea is a superfamily within the order Hymenoptera comprising ants and their extinct relatives. The group originated approximately 120 million years ago during the Early Cretaceous from vespoid wasp ancestors. Formicoidea is defined by the inclusion of the family Formicidae (living and fossil ants) and the extinct family †@@@idae, which includes the mid-Cretaceous genera †Camelomecia and †Camelosphecia. The superfamily is characterized by eusocial behavior, a distinct narrow petiole (waist), and metapleural glands. Ants within Formicoidea represent one of the most ecologically dominant and diverse groups of terrestrial insects, with more than 13,800 described species and estimates of up to 22,000 total species.
eusocial Hymenoptera Aculeata ecosystem-engineers invasive-species model-organisms