Mycangia
Guides
Asteromyia
Asteromyia is a genus of gall midges comprising approximately nine described species. Members of this genus induce distinct gall morphologies on host plants in the Asteraceae family, particularly Solidago (goldenrod) species. The genus is notable for complex ecological interactions, including obligate mutualisms with symbiotic fungi that form protective gall structures and mediate defense against parasitoids. Some species complexes exhibit incipient adaptive radiation with sympatric sibling species producing phenotypically distinct galls on the same host plant.
Asteromyia carbonifera
Carbonifera goldenrod gall midge
Asteromyia carbonifera is a gall midge that forms an obligate mutualistic relationship with the fungus Botryosphaeria dothidea. Females carry fungal conidia in specialized mycangia on the ovipositor and deposit them alongside eggs on Solidago host plants. The fungus induces gall formation, and larvae feed on fungal tissue within the gall. This insect-fungus mutualism is essential for successful development; neither partner can complete its life cycle independently under natural conditions.
Euplatypus parallelus
Common Ambrosia Beetle
Euplatypus parallelus is a polyphagous ambrosia beetle in the weevil family Curculionidae. Native to Central and South America, it has spread invasively to Africa, tropical Asia, and other regions, likely via timber trade. Adults bore galleries into tree trunks and cultivate symbiotic fungi for food, causing wood staining and economic damage in forest plantations. The species is attracted to ethanol and plant volatiles, with flight activity strongly influenced by temperature and precipitation.
Euwallacea interjectus
ambrosia beetle
Euwallacea interjectus is an ambrosia beetle native to Asia that has become an invasive pest in multiple regions worldwide. The species cultivates mutualistic Fusarium fungi in galleries within tree xylem, which serve as its primary food source. Females possess specialized mycangia for transporting fungal spores. The beetle has emerged as a significant economic threat to poplar plantations in China and has been documented attacking living trees, a behavior atypical for many ambrosia beetles. In Japan, it vectors Ceratocystis ficicola, a pathogenic fungus causing fig wilt disease.
Euwallacea perbrevis
tea shot-hole borer
Euwallacea perbrevis, the tea shot-hole borer, is an invasive ambrosia beetle native to South and Southeast Asia through Australia. It is part of the Euwallacea fornicatus cryptic species complex, distinguished from its three sibling species (E. fornicatus, E. fornicatior, and E. kuroshio) primarily through molecular genetics rather than morphology. The beetle cultivates symbiotic fungi in galleries bored into host trees and vectors fungal pathogens causing Fusarium branch dieback. It has been introduced to the United States (Florida, Hawaii), Costa Rica, and Panama, where it poses significant economic threats to avocado production and urban trees.
Urocerus cressoni
Black-and-red Horntail
Urocerus cressoni is a large wood-boring wasp in the family Siricidae, commonly known as the Black-and-red Horntail. Females range from 37–50 mm in length, males 18–37 mm. The species is native to North America and has been recorded across Canada from Nova Scotia to Manitoba. Like other horntails, females possess a prominent ovipositor for drilling into wood to deposit eggs, along with a shorter dorsal spine (cornus) that gives the family its common name. The species is non-venomous and poses no threat to humans.
Xyleborini
ambrosia beetles, xyleborine ambrosia beetles
Xyleborini is a tribe of ambrosia beetles within the subfamily Scolytinae (Curculionidae), comprising highly specialized weevils that cultivate symbiotic fungi for food. The tribe dominates ambrosia beetle faunas across Eurasia and the Americas, with the type genus Xyleborus containing over 500 species, though this genus represents an unnatural grouping of distantly related species. Many Xyleborini are economically significant invasive pests that attack healthy trees, while others are secondary colonizers of dead or dying wood. The tribe exhibits exceptional diversity with numerous genera, many of which are small or monotypic.
Xyleborus
ambrosia beetles, bark beetles
Xyleborus is a genus of ambrosia beetles in the family Curculionidae (subfamily Scolytinae). These small wood-boring beetles are characterized by their obligate symbiotic relationship with ambrosia fungi, which they cultivate in galleries carved into host trees and use as their primary food source. Several species have become significant invasive pests, including Xyleborus glabratus (redbay ambrosia beetle), which vectors the fungus causing laurel wilt disease. The genus includes both native and introduced species with varying degrees of host specificity.
Xyleborus ferrugineus
Xyleborus ferrugineus is an ambrosia beetle (subfamily Scolytinae) that cultivates symbiotic fungi for nutrition rather than feeding directly on wood. Females bore into host trees to create galleries where they farm ambrosia fungi, which serve as the sole food source for adults and larvae. The species exhibits strong sexual dimorphism, with females being larger, more numerous, and solely responsible for gallery formation and colony founding. It has been documented as a potential vector of Ceratocystis cacaofunesta, the causal agent of lethal wilt disease in cocoa.
Xyleborus glabratus
Redbay Ambrosia Beetle
Xyleborus glabratus is a small ambrosia beetle native to Asia that has become a destructive invasive pest in the southeastern United States since its detection in 2002. The species is the primary vector of Raffaelea lauricola, the fungal pathogen that causes laurel wilt disease, which has killed millions of native Lauraceae trees including redbay, sassafras, and avocado. Females are the dispersing sex and initiate galleries in host wood, where they cultivate fungal gardens as food for their offspring. The beetle's rapid spread threatens both forest ecosystems and commercial avocado production.
Xylosandrus
Xylosandrus is a genus of ambrosia beetles in the subfamily Scolytinae, family Curculionidae, containing approximately 54 species globally. The type species is Xyleborus morigerus (Blandford, 1894). Species in this genus are fungus-farming beetles that excavate gallery chambers in wood, introduce symbiotic fungi, and feed on fungal growth rather than wood tissue. Several species, including X. crassiusculus and X. germanus, have become invasive pests of ornamental and fruit trees in North America and elsewhere.
Xylosandrus compactus
black twig borer, black coffee borer, black coffee twig borer, tea stem borer
Xylosandrus compactus is a highly invasive ambrosia beetle (Scolytinae) native to subtropical Asia. Females tunnel into twigs of living woody plants, cultivate symbiotic fungi as larval food, and reproduce through arrhenotokous parthenogenesis. The species has spread to over 50 countries, becoming a significant agricultural pest of coffee, tea, cocoa, and avocado, and threatening native forest ecosystems in invaded regions.