Cryptic-species
Guides
Detonidae
woodlice
Detonidae is a family of terrestrial isopods (woodlice) in the order Isopoda, containing at least 3 genera and more than 30 described species. Members of this family are primarily coastal, inhabiting upper intertidal and supratidal zones of rocky shores. The genus Deto includes species with complex taxonomic histories and documented cases of cryptic diversity, such as Deto echinata in southern Africa.
Dichrorampha
Dichrorampha is a genus of small tortricid moths in the subfamily Olethreutinae, comprising over 100 described species distributed across Europe, Asia, and North America. The genus includes both diurnal and crepuscular species, many associated with mountain and alpine habitats. Several species have been investigated as biological control agents for invasive weeds, including D. aeratana for oxeye daisy and D. odorata for Chromolaena odorata. The genus has been subject to recent taxonomic revision, with cryptic species discovered through integrative approaches combining morphology and molecular data.
Dinacoma sanfelipe
Dinacoma sanfelipe is a melolonthine scarab beetle endemic to southern California, described in 2020 as the first new species in the genus in 90 years. The genus Dinacoma is restricted to California and contains cryptic species that are difficult to distinguish. One congener (D. caseyi) is federally listed under the U.S. Endangered Species Act, and all known species in the genus may be of conservation concern.
Diodontus
Diodontus is a genus of solitary aphid-hunting wasps in the family Crabronidae (subfamily Pemphredoninae). Species are distributed across Europe, Africa, and North America. The genus includes cryptic species that can be morphologically nearly identical but ecologically differentiated, as demonstrated by D. argillicola and D. tristis, which are separated primarily by nesting substrate preference (clay versus sand) rather than visible morphology. Females construct nest tunnels in soil or substrate and provision them with paralyzed aphids as larval food.
Disholcaspis prehensa
clasping twig gall wasp
Disholcaspis prehensa is a gall-inducing cynipid wasp native to California that induces distinctive galls on scrub oak and leather oak. The species exhibits heterogony, alternating between sexual and asexual generations that produce two morphologically different gall types. The asexual generation forms conspicuous mushroom-shaped twig galls with clasping bases, while the sexual generation produces small, hidden bud galls.
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Dissomphalus is the most species-rich genus in the family Bethylidae (Hymenoptera), with over 600 valid species globally. The genus exhibits exceptional diversity in the Neotropical region, where the majority of species occur, though it has a cosmopolitan distribution including the Oriental region and southwestern Pacific islands. Species delimitation relies heavily on male genitalia characters and metasomal tergal process structures, with species-groups defined primarily by these morphological features. Taxonomic revisions have revealed high levels of undescribed diversity, particularly in underexplored regions such as Panama, Thailand, and the Fiji-Solomon Islands archipelago.
Elachista
grass-miner moths, typical grass miner moths
Elachista is a large genus of very small gelechioid moths, the type genus of the family Elachistidae. These grass-miner moths are characterized by reduced, "feathery" hindwings and typically display one to three light transverse bands on the forewing uppersides. The genus has a near-cosmopolitan distribution, being absent only from very cold regions and some oceanic islands, with highest diversity in the Palearctic. Taxonomic complexity includes numerous undescribed species, cryptic species complexes, and disputed subgeneric classifications.
Elachistinae
grass-miner moths
Elachistinae is a subfamily of small moths in the family Elachistidae, superfamily Gelechioidea. The subfamily contains approximately 805 valid species placed in 10 genera, with the genus Elachista being the most diverse. Species are predominantly leaf miners, with larvae feeding on monocotyledonous plants, especially sedges (Cyperaceae) and grasses (Poaceae). The subfamily has a worldwide distribution with significant diversity in the Holarctic region. Many species show narrow host plant associations and some exhibit vicariant distribution patterns.
Elmohardyia atlantica-complex
Elmohardyia atlantica-complex is a species complex within the long-legged fly family Dolichopodidae. The group comprises morphologically similar species that have not been fully resolved taxonomically, likely due to cryptic diversity and insufficient sampling. Members are small to medium predatory flies associated with forested habitats. The complex is part of a Neotropical genus with limited published ecological or behavioral documentation.
Empoa gillettei
Empoa gillettei is a Nearctic leafhopper species in the family Cicadellidae, originally described by Van Duzee in 1917. It belongs to the gillettei complex—a group of 14 Nearctic species within Typhlocyba subgenus Empoa that share identical male genitalia. Two nominal species, Empoa acericola and E. caryata (both Christian, 1953), have been synonymized with E. gillettei based on color variation. The species is documented from multiple localities across western and northern North America.
Empria
Empria is a genus of sawflies in the family Tenthredinidae, distributed across the Palaearctic region with records from Europe, Japan, and other parts of Asia. Species within the genus are primarily herbivorous, with documented host plant associations predominantly in Rosaceae and a single record in Ericaceae. The genus includes several morphologically similar species groups that have required taxonomic revision using integrated approaches combining morphology, morphometrics, and molecular data.
Enargia decolor
Pale Enargia, Aspen Twoleaf Tier Moth
A noctuid moth with boreal-transcontinental distribution across North America, primarily associated with trembling aspen and related poplar species. Adults are active in late summer, with larvae capable of reaching outbreak densities that cause extensive defoliation of host stands. The species shows a disjunct western distribution pattern that may represent cryptic diversity.
Encarsia lanceolata
Encarsia lanceolata is a species of parasitoid wasp in the family Aphelinidae, described by Evans and Polaszek in 1997. The genus Encarsia comprises minute chalcidoid wasps that parasitize whiteflies and other hemipteran hosts. Species in this genus are extensively studied for their importance in biological control programs targeting agricultural pests. E. lanceolata belongs to a genus noted for its extensive cryptic species diversity, making accurate identification dependent on detailed morphological or molecular analysis.
Enchenopa binotata
Two-marked Treehopper Complex, twomarked treehopper
Enchenopa binotata is a species complex of treehoppers in the family Membracidae, currently recognized as comprising multiple cryptic species that have diverged through host plant specialization. The complex occurs primarily in eastern North America with some records in Central America. Members are morphologically similar but reproductively isolated through assortative mating based on host plant-associated vibrational communication signals. This system is a classic model for studying sympatric speciation via host shifts.
Endothenia nr-infuscata
Endothenia nr-infuscata is a small tortricid moth in the family Tortricidae, subfamily Olethreutinae. The species epithet 'nr-infuscata' indicates a near relationship to E. infuscata, suggesting morphological similarity that may represent an undescribed or cryptic species. Members of the genus Endothenia are generally associated with herbaceous plants and often have specialized larval feeding habits.
Enicocephalidae
unique-headed bugs, gnat bugs
Enicocephalidae, commonly known as unique-headed bugs or gnat bugs, is a family of approximately 300 species within the ancient and relictual infraorder Enicocephalomorpha. Members are small true bugs, typically around 4 mm in length, characterized by their elongated, constricted heads that give the family its common name. The family exhibits global distribution with greatest species richness in humid tropical and subtropical forests. Enicocephalidae are considered living fossils due to their evolutionarily conserved morphology. The family is classified into about 47 genera placed in five subfamilies, with notable polymorphism including winged males and wingless or short-winged females in many species.
Enigmina
Enigmina is a genus of armored harvestmen in the family Phalangodidae, established by Ubick & Briggs in 2008. The genus contains at least two described species: Enigmina granita and Enigmina warrenorum. Members of this genus belong to the suborder Laniatores, a diverse group of short-legged harvestmen characterized by robust bodies and often elaborate defensive armor. The genus name reflects the enigmatic nature of these cryptic arachnids, which are poorly known in terms of their biology and ecology.
Entedoninae
Entedoninae is a subfamily of chalcid wasps in the family Eulophidae comprising over 90 genera. Members are primarily parasitoids with documented associations across diverse host groups including gall-inducing insects, leafminers, sawflies, thrips, and ants. The subfamily exhibits considerable morphological and ecological diversity, with species found in tropical to temperate habitats worldwide.
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Epermenia is a genus of small moths in the family Epermeniidae, established by Jacob Hübner in 1825. Species in this genus are distributed across the Palearctic, Nearctic, and Oriental regions, with some extending into the Afrotropical and Neotropical regions. Several species are associated with host plants in the family Apiaceae, particularly Angelica species. The genus includes the subgenus Calotripis, originally described by Hübner in the same publication.
Epiblema brightonana
Brighton's Epiblema Moth
Epiblema brightonana is a small tortricid moth in the family Tortricidae, described by Kearfott in 1907. It belongs to the leafroller moth group, a diverse family known for caterpillars that roll or tie leaves for shelter. The species has been recorded in North America, with confirmed observations from Manitoba, Canada and Vermont, USA. As with many Epiblema species, detailed life history and host plant information remain poorly documented in accessible literature.
Epinotia cinereana
Grey Aspen Bell
Epinotia cinereana is a Holarctic tortricid moth in the tribe Eucosmini, resurrected from synonymy with E. nisella in 2012 based on DNA barcoding, genital morphology, and life history differences. It occurs across Europe and North America, with moderate intraspecific genetic variation between continents but no correlated morphological differences. The species was previously hidden under the name E. nisella, a highly variable species with which it had been conflated for decades.
Eremaeoidea
Eremaeoidea is a superfamily of oribatid mites (Acari: Oribatida) established by Oudemans in 1900. Recent integrative taxonomic studies have revealed substantial cryptic diversity within this group, particularly in the genus Caleremaeus. What was long considered a single widespread species, C. monilipes, has been demonstrated to comprise at least five distinct species based on combined genetic, ecological, and morphological evidence. These mites are small, soil-dwelling microarthropods with varied habitat specializations ranging from strict deadwood association to alpine moss communities.
Euceraphis papyrifericola
Paper Birch Aphid
A host-specific aphid species described in 2002, distinguished from close relatives by morphometric differences and karyotype. Restricted to paper birch (Betula papyrifera). Part of a cryptic species complex previously grouped under E. betulae.
Euctenizidae
Wafer-lid Trapdoor Spiders
Euctenizidae is a family of mygalomorph spiders elevated from subfamily status in 2012. The family contains approximately 79 described species across eight genera, including Aptostichus, Myrmekiaphila, and Promyrmekiaphila. Members are characterized by their fossorial lifestyle and construction of silk-lined burrows with distinctive trapdoors. The family is now considered more closely related to Idiopidae than to other trapdoor spider families.
Eupithecia
pug moths, pugs
Eupithecia is the largest genus of moths in the family Geometridae, with over 1,400 described species. Members are commonly known as pugs and are found on every continent except Australasia. Adults are small (typically 12–35 mm), with muted colors and a distinctive resting posture where forewings are held flat at right angles to the body. Species-level identification is difficult and often requires examination of dissected genitalia.
Eupitheciini
Pug Moths, Pugs
Eupitheciini is a tribe of geometer moths in the subfamily Larentiinae, commonly known as pugs. With approximately 1,900 described species distributed across 47 genera, it is the most species-rich tribe in the family Geometridae. The genus Eupithecia alone contains roughly 1,400 species and is one of the largest insect genera globally. Members are typically small moths with narrow wings and cryptic coloration. Identification relies heavily on genitalia dissection due to morphological homogeneity among species.
Eupsilia schweitzeri
Schweitzer's Sallow
Eupsilia schweitzeri is a moth species in the family Noctuidae, described in 2016 from the northeastern United States. It belongs to the sallow moth genus Eupsilia, which are medium-sized noctuid moths active during late fall, winter, and early spring. The species was named in honor of entomologist Dale F. Schweitzer. Like other members of its genus, it is likely a cold-season active adult with larvae feeding on woody plants.
Eurhopalothrix floridana
Eurhopalothrix floridana is a species of ant in the subfamily Myrmicinae, originally described by Brown & Kempf in 1960. It belongs to the genus Eurhopalothrix, a group of small, cryptic ants within the Attini tribe. The species is known from the Dominican Republic, with records from the Jardín Botánico Nacional in Santo Domingo de Guzmán. Very little biological information has been published about this species; it has been mentioned in taxonomic literature primarily in comparative studies of male morphology.
Eurytomidae
seed chalcids, eurytomid wasps
Eurytomidae is a family of chalcid wasps within the superfamily Chalcidoidea, comprising approximately 1,420 described species across 87 genera. Members exhibit diverse life histories: larvae are endophytic, developing within plant tissues (stems, seeds, or galls) as phytophages, parasitoids of other insects, or both. The family is cosmopolitan, occurring in virtually all terrestrial habitats. Some species are agricultural pests, notably the almond seed wasp (Eurytoma amygdali) and plum seed wasp (Eurytoma schreineri), while others serve as biocontrol agents. Phylogenomic studies have confirmed monophyly of the four recognized subfamilies (Buresiinae, Eurytominae, Heimbrinae, Rileyinae), though many large genera remain para- or polyphyletic.
Eusphalerum near-subangulatum
Eusphalerum near-subangulatum is a rove beetle in the subfamily Omalinae, recognized as a distinct but closely related entity to Eusphalerum subangulatum. It is a small beetle associated with riparian and wetland habitats. The species remains poorly documented in formal taxonomic literature.
Euteliidae
Euteliid Moths
Euteliidae is a small family of moths in the superfamily Noctuoidea, comprising approximately 540 species globally. The family is characterized by highly specialized larval hostplant relationships with lactiferous plants containing high levels of resin or latex. Molecular phylogenetic analyses recognize two subfamilies: Euteliinae and Stictopterinae, with Stictopterinae further divided into tribes Stictopterini and Odontodini. The family has been the focus of recent taxonomic revision due to the discovery of cryptic species complexes and polyphyletic genera requiring reclassification.
Euura
Euura is a genus of sawflies in the family Tenthredinidae, subfamily Nematinae, with exceptional species diversity concentrated in northern Holarctic regions rather than tropical zones. The genus contains both free-feeding larvae that consume plant foliage externally and gall-inducing species whose larvae develop entirely within plant galls. Gall-forming females inject a stimulant during oviposition that initiates gall growth before eggs hatch, a distinctive mechanism among gall insects. Taxonomic revision in 2014 consolidated several former genera (Eupontania, Phyllocolpa, Pontania, Tubpontania) into Euura based on phylogenetic evidence. The genus exhibits substantial genetic complexity, with frequent mito-nuclear discordance and multiple functional COI variants within individuals complicating species identification.
Euwallacea
Ambrosia beetles, Shot hole borers
Euwallacea is a genus of ambrosia beetles within the tribe Xyleborini. All species are obligate symbionts with ambrosia fungi, which they farm in galleries excavated within host trees. The genus has a pantropical distribution with origins in Asia and Wallacea, and several species have become globally significant invasive pests. Members of the Euwallacea fornicatus species complex, including the polyphagous shot hole borer, cause substantial economic damage to fruit and timber trees through mass colonization and associated fungal pathogens.
Euxoa maimes
Euxoa maimes is a species of noctuid moth described by Smith in 1903. It belongs to the genus Euxoa, a diverse group of owlet moths commonly known as dart moths or cutworms. The species is documented from prairie regions of western Canada, with records from Alberta, Manitoba, and Saskatchewan. Like other Euxoa species, it likely has a life cycle involving overwintering larvae and nocturnal adult activity, though specific biological details for this species remain poorly documented.
Euxoa simona
cutworm moth, dart moth
Euxoa simona is a species of cutworm or dart moth in the family Noctuidae, described by James Halliday McDunnough in 1932. It belongs to the large genus Euxoa, which contains numerous similar-looking species of owlet moths. The species is known from North America, with confirmed records from the Canadian provinces of Alberta and Saskatchewan. Like other Euxoa species, it is nocturnal and attracted to light sources.
Exosphaeroma diminutum
Exosphaeroma diminutum is a small marine isopod in the family Sphaeromatidae, described by Menzies and Frankenberg in 1966. The species epithet 'diminutum' reflects its notably small body size relative to congeners. Like other Exosphaeroma species, it belongs to a group of crustaceans commonly known as marine pillbugs or rolly pollies, which are relatives of terrestrial isopods. The species has been recorded from Saint Thomas in the Caribbean region.
Gargara
Gargara is a genus of treehoppers in the family Membracidae, subfamily Centrotinae, tribe Gargarini. The genus was established by Amyot and Audinet-Serville in 1843. Species in this genus are distributed across the Palearctic region, with documented occurrences in Russia, Kazakhstan, Central Asia, Denmark, and Norway. The Gargara genistae species group contains cryptic species that are morphologically nearly identical but distinguished by male calling signal patterns.
Gargara genistae
Gargara genistae is a small treehopper in the family Membracidae, subfamily Centrotinae, characterized by a prominent backward-projecting pronotum and dark brown coloration. The species is oligophagous, feeding exclusively on phloem sap of shrubby Fabaceae including brooms and related genera. It is widely distributed across the Holarctic, Ethiopian, and Oriental regions, with occasional records in North America as an apparent introduction. Males produce species-specific substrate-borne calling signals used for mate recognition; cryptic species within this group are distinguished by acoustic rather than morphological characters.
Garriscaphus amplus
Garriscaphus amplus is a species of soil centipede in the family Himantariidae, described by Chamberlin in 1941. As a member of the order Geophilomorpha, it is an elongate, multi-legged myriapod adapted for burrowing through soil and leaf litter. The genus Garriscaphus is poorly known, and this species has received limited study. Available records indicate it occurs in the United States, though specific ecological details remain largely undocumented.
Gelechiidae
twirler moths, gelechiid moths
Gelechiidae is a large family of small moths, commonly known as twirler moths, that serves as the namesake family for the superfamily Gelechioidea. The family contains over 4,500 described species across more than 900 genera, with particularly high diversity in North America. Members are characterized by their small size, narrow fringed wings, and larvae that typically feed internally on host plant tissues. Many species are significant agricultural pests, while others have been employed for biological control of invasive plants.
Glaresis california
Glaresis california is a species of scarab beetle in the family Glaresidae, described in 2014 from California. Members of this family are small, soil-dwelling beetles often called 'enigmatic scarab beetles' due to their cryptic habits and uncertain phylogenetic placement. The species is known only from its type locality in California and is among the more recently described species in this poorly studied family.
Gnorimoschema nr-septentrionella-two
Gnorimoschema nr-septentrionella-two is a species of gelechiid moth in the genus Gnorimoschema. The "nr" designation indicates this is a near-reference specimen that closely resembles but may not be definitively identified as Gnorimoschema septentrionella. The "two" suffix suggests it represents a second distinct morphotype or genetic cluster within this near-septentrionella group. Species in this genus are typically small moths associated with herbaceous host plants, often forming galls or feeding internally in stems or roots.
Gonatista
Grizzled Mantises
Gonatista is a genus of praying mantises in the family Epaphroditidae, distributed across North America and the Caribbean. The genus contains five recognized species that are morphologically similar and difficult to distinguish based on coloration and body size alone. Species-level identification relies primarily on the morphology of the external copulatory apparatus. The most well-known member is Gonatista grisea, commonly called the Grizzled Mantis.
Gonatocerus morrilli
Gonatocerus morrilli is a mymarid wasp and specialized egg parasitoid of sharpshooters in the genus Homalodisca, particularly the glassy-winged sharpshooter (Homalodisca vitripennis, formerly H. coagulata). It has been widely used in California as a biological control agent against this pest, which vectors Xylella fastidiosa, the bacterium causing Pierce's disease in grapevines. Molecular studies have revealed significant genetic differentiation between California and Texas populations, with evidence suggesting the California population may represent a cryptic species. The species exhibits high parasitism efficiency, with females capable of parasitizing virtually all eggs in egg masses within 24 hours.
Graemeloweus glimmei
A small scorpion species in the family Vaejovidae, originally described as Vaejovis glimmei in 1972 and later transferred to the genus Graemeloweus. The genus Graemeloweus was established in 2016 to accommodate several vaejovid species previously placed in Vaejovis. This species is known from a limited number of observations and specimens, primarily from western North America.
Gryllotalpa
mole crickets
Gryllotalpa is a genus of mole crickets in the family Gryllotalpidae, characterized by highly modified fossorial front legs adapted for digging. The genus contains numerous species distributed across Africa, Europe, Asia, Australia, and the Americas, with several cryptic species complexes distinguished primarily by song patterns. Species within this genus are found in diverse habitats ranging from steppe zones and wet grasslands to gardens and agricultural fields. Some species, particularly Gryllotalpa gryllotalpa, are of conservation concern in parts of Europe while also being agricultural pests in other regions.
Gryllotalpa gryllotalpa
European mole cricket
Gryllotalpa gryllotalpa, the European mole cricket, is a subterranean insect native to the Western Palaearctic and introduced to the eastern United States. It is characterized by fossorial forelegs adapted for digging, velvety body covering, and a subterranean lifestyle. The species is now understood to represent a cryptic species complex distinguishable by song patterns. In the United Kingdom, it is legally protected and considered endangered due to dramatic range contraction.
Gryllus rubens
Southeastern field cricket
Gryllus rubens, the southeastern field cricket, is a trilling field cricket native to the southeastern United States. It is a cryptic sister species to G. texensis, from which it originated via peripatric speciation. The species exhibits remarkable phenotypic plasticity in its communication system, with male calling songs and female preferences shifting reversibly between spring and fall generations in response to temperature. Wing polymorphism is environmentally and genetically influenced, with long-winged morphs appearing in spring/summer and short-winged morphs predominating in autumn/winter.
Gryllus texensis
Texas Trilling Cricket
Gryllus texensis is a field cricket species described by Cade and Otte in 2000, distinguished from the morphologically similar Gryllus rubens by differences in call song structure. The species was previously misidentified as Gryllus integer in collections from the southeastern United States. It belongs to a closely related complex of Gryllus field crickets and is phylogenetically most closely related to G. rubens based on mitochondrial DNA analysis.
Gymnochthebius fossatus
Gymnochthebius fossatus is a species of minute moss beetle in the family Hydraenidae, first described by LeConte in 1855. It is one of the smallest beetles in North America, with adults typically measuring less than 2 mm in length. The species has a broad geographic distribution spanning the Caribbean Sea, Central America, North America, and South America. As a member of Hydraenidae, it belongs to a family of beetles commonly known as moss beetles or minute moss beetles, which are primarily associated with moist microhabitats.