Biocontrol
Guides
Eupelmidae
Eupelmidae is a family of parasitic wasps in the superfamily Chalcidoidea, containing approximately 905-1100 described species across 39-45 genera. The family is currently defined to include three subfamilies: Eupelminae, Calosotinae, and Eusandalinae. Members exhibit remarkable sexual dimorphism, with females of many species possessing modified mesothoracic musculature that enables powerful jumping using the middle legs. The majority are parasitoids of other insects, with hosts including beetle larvae, spider eggs, mantid oothecae, and various other arthropods. Some species are facultative hyperparasitoids. The group has significant potential as biological control agents for agricultural and forest pests.
Eupeodes americanus
American hoverfly, Long-tailed Aphideater
Eupeodes americanus, the American hoverfly, is a Nearctic aphidophagous hoverfly commercialized in Canada as a biological control agent for aphid management. Adults feed on nectar and pollen, while larvae are voracious predators of aphids, with one larva consuming approximately 2,000 aphids during development. The species is notable for its ability to remain active at low temperatures (12–18°C) where other biocontrol agents become inactive. It has been evaluated as a biocontrol agent for greenhouse and field crops including sweet pepper, cucumber, cotton, soybeans, melons, and potatoes.
Eurytomidae
seed chalcids, eurytomid wasps
Eurytomidae is a family of chalcid wasps within the superfamily Chalcidoidea, comprising approximately 1,420 described species across 87 genera. Members exhibit diverse life histories: larvae are endophytic, developing within plant tissues (stems, seeds, or galls) as phytophages, parasitoids of other insects, or both. The family is cosmopolitan, occurring in virtually all terrestrial habitats. Some species are agricultural pests, notably the almond seed wasp (Eurytoma amygdali) and plum seed wasp (Eurytoma schreineri), while others serve as biocontrol agents. Phylogenomic studies have confirmed monophyly of the four recognized subfamilies (Buresiinae, Eurytominae, Heimbrinae, Rileyinae), though many large genera remain para- or polyphyletic.
Eustala
humpbacked orb-weavers
Eustala is a genus of orb-weaver spiders in the family Araneidae, first described by Eugène Simon in 1895. As of November 2024, the genus contains 87 species distributed primarily across the Americas, from the United States through Central America to South America, with highest diversity in Brazil, Uruguay, and Argentina. Several species have been documented exhibiting specialized ecological relationships, including myrmecophily (living among ants) in Central American species that inhabit ant-defended acacia plants. The genus includes the well-known humpbacked orb-weaver, Eustala anastera, found across North and Central America.
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htperchroostnestdenburrowtunneldigexcavateminequarryextractremoveeliminateeradicateexterminatedestroydemolishruinwreckdamageharminjurehurtwoundmaimcripplelamedisableincapacitateparalyzestunshockastonishamazeastoundstaggerstupefydazebewilderconfusepuzzleperplexbafflemystifyenigmaEustenopus villosus
yellow starthistle hairy weevil
Eustenopus villosus is a small weevil in the family Curculionidae, widely known as the yellow starthistle hairy weevil. It is native to southern Europe and the Mediterranean region and has been introduced to the western United States as a biological control agent against the invasive weed yellow starthistle (Centaurea solstitialis). Both adults and larvae feed on the plant's reproductive structures, with larvae capable of destroying nearly all developing seeds within a single flower head. The species is considered one of the more effective biocontrol agents for this noxious weed.
Euteliidae
Euteliid Moths
Euteliidae is a small family of moths in the superfamily Noctuoidea, comprising approximately 540 species globally. The family is characterized by highly specialized larval hostplant relationships with lactiferous plants containing high levels of resin or latex. Molecular phylogenetic analyses recognize two subfamilies: Euteliinae and Stictopterinae, with Stictopterinae further divided into tribes Stictopterini and Odontodini. The family has been the focus of recent taxonomic revision due to the discovery of cryptic species complexes and polyphyletic genera requiring reclassification.
Exapion fuscirostre
Scotch broom seed weevil
Exapion fuscirostre is a small straight-snouted weevil introduced as a biological control agent for the invasive plant Scotch broom (Cytisus scoparius). Adults feed on new stem growth while larvae develop inside seed pods, consuming seeds. Native to Europe, it has been established in the western United States and New Zealand.
Exorista
uzi fly
Exorista is a genus of tachinid flies comprising approximately 70 described species distributed across multiple subgenera. The genus includes both beneficial biocontrol agents and economically significant pests of sericulture. Species such as E. sorbillans (uzi fly) are major constraints on silk production, causing up to 80% crop loss in outdoor silkworm operations through endoparasitic larval development. Other species have been evaluated for biological control of agricultural pests, including E. larvarum for in vitro rearing and E. deligata for control of tea loopers. The genus exhibits diverse host associations, primarily targeting Lepidoptera.
Gargaphia
Gargaphia is a genus of lace bugs (family Tingidae) containing more than 70 described species. Members are small, phytophagous true bugs characterized by intricate reticulated forewings. Several species are economically significant as agricultural pests, particularly on solanaceous crops and passion fruit. The genus is notable for exhibiting maternal care behaviors, including egg guarding and defensive responses to predators.
Harmonia dimidiata
Greater Asian Lady Beetle
Harmonia dimidiata is a medium to large predaceous ladybird beetle (7.4–10 mm) native to southern Asia with introduced populations in North America. Adults display reddish-yellow elytra with black markings in a thirteen-spot pattern. The species is a voracious aphid predator with documented prey including Rhopalosiphum maidis, Aphis gossypii, Myzus persicae, and Aphis pomi. Temperature strongly influences its development, fecundity, and predatory capacity, with optimal performance at 24°C. Males produce yellow reflex blood containing harmonine as both chemical defense and nuptial gift during courtship.
Hayhurstia
Hayhurstia is a monotypic genus of aphids in the family Aphididae, containing the single species H. atriplicis. The genus is notable for its role in biological control research, as populations serve as reservoirs for parasitoids that have been used in pest management programs. It is associated with Chenopodium (goosefoot) plants in agricultural and urban environments.
Hayhurstia atriplicis
Chenopodium aphid, Saltbush Leaf-Fold Aphid
Hayhurstia atriplicis is a holocyclic, monoecious aphid species associated with Chenopodium host plants. It is considered economically indifferent but serves as a perennial reservoir for parasitoid species, particularly Diaretiella rapae and Ephedrus nacheri. These parasitoids have been utilized in biological control programs targeting other pest aphids, including Diuraphis noxia and Schizaphis graminum.
Hyadaphis coriandri
coriander aphid
Hyadaphis coriandri is a species of aphid specialized on coriander (Coriandrum sativum), where it is considered a major pest. It has been documented as a prey species for the ladybird beetle Menochilus sexmaculatus in laboratory biocontrol studies, though it supports predator development with reduced growth metrics compared to alternative aphid hosts. The species has a wide geographic distribution including parts of Asia, Europe (Madeira), and North America (Hawaii, conterminous United States).
Hydrochara
water scavenger beetles
Hydrochara is a genus of water scavenger beetles (Hydrophilidae) containing 23 species distributed across North America, Europe, Asia, and Africa. The genus was revised taxonomically in 1980, with 21 species recognized at that time and multiple synonymies established. Species in this genus are aquatic and have been studied for their potential role in biological control of mosquito larvae. At least one species, Hydrochara rickseckeri, is endemic to California.
Hypera rumicis
Dock Hyper Weevil
Hypera rumicis is a small true weevil in the family Curculionidae, known as the Dock Hyper Weevil. The species is native to Europe and has been introduced to North America. It is a specialist herbivore associated with dock and sorrel plants (Rumex spp.), where both larvae and adults feed on host tissues. The species has been investigated as a potential biological control agent for invasive Rumex weeds, though it can also cause damage to crops. Adults show complex behavioral responses to plant volatile compounds, with attraction and repellency depending on concentration and mating status.
Hyperaspis cincta
Corona Sigil Lady Beetle
Hyperaspis cincta is a small lady beetle species in the family Coccinellidae, measuring approximately 2.0–2.6 mm in length. It is native to western North America, with confirmed records from California and Utah. Adults are distinguished from the similar Hyperaspis inflexa by their wider marginal vitta on the elytra. The genus Hyperaspis includes species known as predatory biocontrol agents of scale insects.
Inocelliidae
Square-headed Snakeflies, inocelliid snakeflies
Inocelliidae is a small family of snakeflies (order Raphidioptera) containing eight extant genera plus fossil taxa. Members are commonly known as square-headed snakeflies due to their distinctive head shape. The family is distributed across the Northern Hemisphere, with extant species found in temperate forests of Eurasia and North America. Inocelliidae are holometabolous predators with larvae that inhabit bark and wood of coniferous and deciduous trees, where they prey on other insects. The family includes notable fossil representatives such as Fibla carpenteri from Baltic amber, the largest known snakefly species.
Isturgia
Isturgia is a genus of geometrid moths established by Jacob Hübner in 1823. The genus belongs to the subfamily Ennominae and contains species distributed across Europe and parts of Asia. Several species have been studied for their biology and ecological roles, including one species (I. disputaria) investigated as a potential biological control agent for invasive acacia species. Taxonomic relationships within the genus remain incompletely resolved, particularly for species complexes such as I. limbaria/roraria.
Lasiochilidae
Lasiochilid Pirate Bugs
Lasiochilidae is a family of minute predatory true bugs in the order Hemiptera, commonly referred to as lasiochilid pirate bugs. These insects are closely related to Anthocoridae (flower bugs) and share similar ecological roles as predators of small arthropods and pests. The family has been documented across the Oriental and Palaearctic regions, with recent taxonomic work in India identifying multiple species and new regional records.
Lasiochilus pallidulus
Pale Pirate Bug
Lasiochilus pallidulus, commonly known as the Pale Pirate Bug, is a predatory true bug in the family Lasiochilidae. It is distributed across the Caribbean, Central America, and North America. The species is known to prey on eggs and early instars of Blissus insularis, a chinch bug species. Like other members of Lasiochilidae, it likely functions as a generalist predator in various terrestrial habitats.
Lasioptera
Lasioptera is a genus of gall midges in the family Cecidomyiidae containing at least 140 described species. Species in this genus are primarily known for inducing galls on host plants, with larvae developing inside plant tissues. Some species have established complex relationships with fungi, including ambrosia fungus feeding. Several species are significant agricultural pests, including the Mediterranean tomato gall midge (L. tomaticola) and a stem-boring pest on tomatoes reported from Greece and Turkey. The genus has a wide geographic distribution spanning Europe, Asia, North America, and other regions, with host associations including Rubus (raspberry/blackberry), tomato, Arundo donax (giant reed), Leea indica, and various other plants.
Leptostales
Leptostales is a genus of geometrid moths in the subfamily Sterrhinae, established by Möschler in 1890. The genus contains multiple species, with Leptostales ignifera being the most studied due to its potential as a biocontrol agent for the invasive weed Lantana camara. Members are leaf-feeding moths found in subtropical regions of the Americas.
Leucopis americana
silver fly
Leucopis americana is a species of silver fly in the family Chamaemyiidae, described by Malloch in 1921. Members of this genus are predatory insects that have been investigated as potential biological control agents for hemlock woolly adelgid (Adelges tsugae). The species is part of a group of Leucopis flies under evaluation for release in eastern North America to control this invasive forest pest.
Linyphia triangularis
Common sheetweb spider
Linyphia triangularis is a European sheet-web spider in the family Linyphiidae, first described by Carl Alexander Clerck in 1758. It is abundant throughout Europe and has been introduced to Maine, USA, where it is now established in coastal areas. The species constructs horizontal sheet-webs with 'barrage lines' above to intercept flying prey. Adults are active in late summer and autumn.
Malacocoris chlorizans
delicate apple capsid
Malacocoris chlorizans is a small green plant bug in the family Miridae, commonly known as the delicate apple capsid. The species is predatory, feeding on mites and aphids in deciduous tree canopies, particularly hazel. Adults are active from spring through autumn. It is native to Europe and extends eastward to the Caspian Sea region.
Mantis religiosa
European mantis, praying mantis
Mantis religiosa, the European mantis, is a large hemimetabolic insect native to Europe that has established widespread populations in North America since its introduction in 1899. It is distinguished by a diagnostic black spot with white center on the inner surface of each foreleg. The species exhibits remarkable color polymorphism, appearing in shades of green, brown, and bluish-green. Females deposit brown, Styrofoam-like egg cases (oothecae) containing over 100 eggs that overwinter and hatch in spring. Sexual cannibalism occurs, with females sometimes consuming males during or after mating.
Medetera
Woodpecker Flies
Medetera is a large genus of predatory long-legged flies (family Dolichopodidae) comprising approximately 350 species worldwide. Adults are commonly found resting on vertical surfaces, particularly tree trunks, where they adopt a characteristic upright stance that has earned them the common name "woodpecker flies." Adults prey on soft-bodied arthropods, while larvae develop as predators within bark beetle galleries, feeding on eggs, larvae, pupae, and newly emerged adults. Several species have been studied as potential biocontrol agents for forest pest management.
Medeterinae
Medeterinae is a subfamily of long-legged flies (Dolichopodidae) comprising approximately 40 genera arranged in four tribes: Medeterini, Systenini, Thrypticini, and Udzungwomyiini. The group includes both extant and fossil genera, with several described from amber deposits. The subfamily is most diverse in tropical and subtropical regions, with particular richness in the Afrotropical and Oriental realms.
Melangyna
Halfbands
Melangyna is a genus of hoverflies (Syrphidae) distributed across the Holarctic region, with subgenera in the Nearctic, Palearctic, and Australasian regions. Adults are frequent flower visitors, while larvae are predatory on aphids. The genus exhibits sexual dimorphism in behavior and morphology, with males typically larger than females and showing distinct foraging and habitat exploration patterns.
Meteorus
Meteorus is a large genus of parasitoid wasps in the family Braconidae, comprising over 330 described species worldwide. These wasps are distinguished from other braconids by two key morphological features: a second submarginal cell in the forewing and a petiolate (stalked) first tergite. The genus includes species with remarkable behavioral adaptations, such as Meteorus stellatus, which constructs star-shaped communal cocoon masses suspended by threads up to 1 meter long. Meteorus species function as biological control agents, parasitizing diverse host insects including caterpillars of moths and beetles.
Microvelia pulchella
smaller water strider
Microvelia pulchella is a semiaquatic predatory bug in the family Veliidae, distributed throughout the American continents from Canada to Argentina. It is the type species of the genus Microvelia and one of the most common species in the Neotropical Region. Laboratory studies demonstrate its potential as a biological control agent against disease-vector mosquitoes and agricultural pests in rice fields. The species serves as a model organism for behavioral and ecological research due to its convenient rearing characteristics and measurable traits.
Mirax
miracine wasps
Mirax is a genus of tiny parasitoid wasps in the subfamily Miracinae (Braconidae). Species are typically 1–2 mm in length and are koinobiont endoparasitoids of leaf-mining caterpillars. The genus has been documented in Australia, Puerto Rico, and other regions, with several species described through citizen science initiatives. Some species, such as Mirax insularis, have been studied for their potential as biological control agents of agricultural pests.
Mogulones
minute seed weevils
Mogulones is a genus of minute seed weevils in the family Curculionidae, containing at least 30 described species. Several species have been investigated as biocontrol agents for invasive plants, particularly houndstongue (Cynoglossum officinale). The genus is characterized by small body size typical of seed-feeding weevils.
Mozena
leaf-footed bugs
Mozena is a genus of leaf-footed bugs in the family Coreidae, containing more than 30 described species. The genus was established by Amyot & Serville in 1843 and is placed in the tribe Nematopodini. Some species, such as M. obtusa, have been investigated as potential biocontrol agents for invasive Prosopis (mesquite) species. Members of this genus are found primarily in the Americas, with records from the southwestern United States through Mexico.
Mozena obtusa
leaf-footed bug
Mozena obtusa is a species of leaf-footed bug (family Coreidae) native to North America. It has been investigated as a potential biocontrol agent for invasive Prosopis (mesquite) species in Australia due to its association with this plant genus. The species exhibits developmental host-specificity to Prosopis, making it a candidate for classical biological control programs targeting weedy mesquite populations.
Nabis capsiformis
Pale Damsel Bug
Nabis capsiformis, commonly known as the pale damsel bug, is one of the most widespread species in the genus Nabis. It is a generalist predator found across six continents including Africa, Europe, the Americas, and Oceania. The species is particularly abundant in agricultural ecosystems where it serves as a biocontrol agent. Both nymphs and adults are effective predators using raptorial front legs to capture prey.
Nabis propinquus
Marsh Damsel Bug
Nabis propinquus, commonly known as the Marsh Damsel Bug, is a small predatory true bug in the family Nabidae. Like other damsel bugs, it is a nocturnal predator that subdues other insects, including prey larger than itself. The species has been observed in marshy habitats and is attracted to lights at night, presumably to hunt other insects drawn to illumination. As with most nabids, it possesses a muscular, swollen front femur armed with teeth to secure prey, and a short rostrum compared to plant-feeding true bugs.
Nasonia
jewel wasp
Nasonia is a genus of small pteromalid parasitoid wasps comprising four described species: N. vitripennis, N. giraulti, N. longicornis, and N. oneida. These wasps are ectoparasitoids of fly pupae, primarily targeting blow flies and flesh flies. The genus has become a prominent model system in behavioral genetics and evolutionary biology due to its short generation time, ease of laboratory rearing, and the ability to hybridize species in controlled crosses. Nasonia wasps exhibit a distinctive emerald sheen on their exoskeleton, earning them the common name 'jewel wasps.'
Nealiolus
Nealiolus is a genus of parasitoid wasps in the family Braconidae, subfamily Brachistinae. The genus comprises nine described species, with three occurring in the Neotropical region. Members are specialized parasitoids of stem-boring and fruit-boring weevils (Curculionidae), including significant agricultural pests such as the sunflower stem weevil and cotton boll weevil. The genus has recognized potential as a biological control agent due to its host specificity and high parasitism rates.
Ophiomyia lantanae
Lantana Seed Fly
Ophiomyia lantanae is a small fly in the family Agromyzidae, native to the Americas from the southern United States to Brazil. It was introduced to Australia in 1914 as a biological control agent for the invasive weed Lantana camara. The species has since established populations in multiple countries across several continents. It is commonly known as the Lantana Seed Fly due to its association with its host plant.
Orius tristicolor
minute pirate bug, minute flower bug
Orius tristicolor is a minute pirate bug in the family Anthocoridae, known for its role as a generalist predator of small arthropod pests. The species exhibits behavioral plasticity in foraging, adjusting search effort based on leaf damage and prey type. It develops through five nymphal instars before reaching adulthood and has been documented in agricultural systems, particularly in association with thrips and spider mites. The species occurs across a broad geographic range including North, Central, and South America, the Caribbean, and Oceania.
Oxyopidae
Lynx Spiders
Oxyopidae, commonly known as lynx spiders, is a family of hunting spiders characterized by their spiny legs and active predatory behavior. Most species are diurnal hunters that rely on vision and agility rather than web-building to capture prey. The family includes notable genera such as Oxyopes, Peucetia, and Hamataliwa, with Peucetia viridans (green lynx spider) being among the largest North American species. While most lynx spiders are solitary, at least one species, Tapinillus sp., has been documented exhibiting social behavior including cooperative prey capture and communal web maintenance—the first such report for the family.
Pediobius
Pediobius is a genus of minute chalcid wasps in the family Eulophidae, containing over 200 described species. The larvae are parasitoids of various arthropods, primarily insects. Several species have been employed in biological pest control programs, including Pediobius foveolatus against Mexican bean beetles and Pediobius cajanus against the Asian fly in pigeon pea crops.
Pegomya
Pegomya is a genus of flies in the family Anthomyiidae comprising diverse species with varied larval feeding strategies. Larvae exhibit leaf mining, stem boring, gall induction, seed feeding, and mycophagy. Several species are economically significant agricultural pests, including those attacking sugar beet, spinach, and cruciferous crops. Some species have been investigated for biological control of invasive plants.
Pipunculinae
big-headed flies
Pipunculinae is a subfamily of big-headed flies within the family Pipunculidae. It comprises approximately 9 genera and at least 130 described species. Members are parasitoids, primarily attacking leafhoppers and planthoppers (Auchenorrhyncha). The subfamily is divided into five tribes: Cephalopsini, Microcephalopsini, Eudorylini, Tomosvaryellini, and Pipunculini.
Poecilanthrax willistonii
Williston's bee fly
Poecilanthrax willistonii is a North American bee fly in the family Bombyliidae. The genus Poecilanthrax is strictly North American in distribution, with the greatest abundance of species and individuals found in the Great Basin region. Like other members of the genus, the larvae develop as parasites inside caterpillars of cutworms and armyworms (family Noctuidae). Adults are typically found in arid and semi-arid habitats, often associated with flowering plants where they feed on nectar.
Polistes
Umbrella Paper Wasps, Umbrella Wasps, Paper Wasps
Polistes is a cosmopolitan genus of social paper wasps and the sole genus in the tribe Polistini. With over 200 recognized species, it is the largest genus in the family Vespidae. These wasps construct distinctive open, single-layered umbrella-shaped nests from wood fibers mixed with saliva, typically suspended by a narrow stalk from protected surfaces. They exhibit complex social behavior including dominance hierarchies, kin recognition through learned chemical cues, and cooperative colony founding. While generally less aggressive than yellowjackets, they will defend nests when provoked.
Polistes dominula
European Paper Wasp
Polistes dominula is a highly successful invasive social wasp native to Eurasia that has established populations across North America, South America, New Zealand, South Africa, and other regions. First detected in North America near Boston in 1978, it has become one of the most abundant wasps on the continent. The species builds small, exposed paper nests in protected locations and preys primarily on live insects, particularly caterpillars. Unlike yellowjackets, it does not scavenge for meat or sugar. Its rapid spread has been attributed to ecological flexibility, superior competitive ability, and tolerance of human-altered environments.
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flavus
Yellow Paper Wasp
Polistes flavus, commonly known as the yellow paper wasp, is a large, conspicuous paper wasp native to the southwestern United States and Mexico. It is strongly associated with the Sonoran Desert, where it occupies the same elevation belt as saguaro cacti. The species builds exposed paper nests from wood fibers mixed with saliva, typically placing them under building eaves or among tree branches. Adults forage for caterpillars and other insects to feed larvae, while consuming nectar and honeydew for their own energy needs. Despite its relative abundance in suitable habitats, detailed natural history information remains limited.
Pristomerus
Pristomerus is a genus of parasitoid wasps in the family Ichneumonidae, subfamily Cremastinae, distributed worldwide with greatest diversity in tropical regions. Members are solitary koinobiont endoparasitoids that attack concealed larvae of small Lepidoptera. Several species have been employed in biological control programs. The genus exhibits high regional endemism, with 49 species recognized from the Afrotropical region, 22 from Australia (19 newly described in 2016), and 15 from Madagascar (12 endemic).