Citrus-pest

Guides

Lepidosaphes
armored scale insects
Lepidosaphes is a large genus of armored scale insects (Diaspididae) with at least 190 described species distributed worldwide. Species in this genus are phytophagous pests of woody plants, particularly fruit trees and ornamentals. Several species, including L. beckii and L. ulmi, are significant agricultural pests. Members of this genus are hosts for various parasitoids, notably species of Aphytis (Aphelinidae), which are used in biological control programs.
armored-scale Diaspididae agricultural-pest biological-control citrus-pest parasitoid-host
Leptoglossus
leaf-footed bugs
Leptoglossus is a genus of true bugs in the leaf-footed bug family Coreidae, tribe Anisoscelini. Species are characterized by leaflike dilations of the hind tibia, a diagnostic trait of the genus. The genus is distributed throughout the Americas, with some introduced populations in Europe and Asia. Several species are economically significant agricultural pests, notably L. occidentalis, which has become invasive in multiple continents.
Coreidae leaf-footed-bug agricultural-pest invasive-species symbiosis sexual-dimorphism conifer-pest nuisance-pest Burkholderia tachinid-parasitoid pheromone-communication building-invader misidentification Triatoma-look-alike gradual-metamorphosis seed-predator forest-pest ornamental-pest plumbing-damage public-health-confusion Chile Europe North-America South-America Asia Turkey almond-pest citrus-pest tomato-pest corn-pest overwintering-aggregation defensive-secretion pheromone-mediated-parasitism egg-parasitoid biological-control ecological-niche-modeling climate-suitability range-expansion introduction accidental-dispersal seed-orchard-pest germination-reduction empty-seed-formation cone-damage lodgepole-pine Douglas-fir western-conifer-seed-bug L.-occidentalis L.-zonatus L.-phyllopus L.-clypealis L.-australis L.-chilensis L.-oppositus male-combat femoral-weapon abdominal-gland pheromone-gland tibial-dilation leaf-like-hind-leg true-bug Hemiptera Heteroptera Pentatomomorpha Anisoscelini phytophagous plant-feeding stylet-feeding salivary-sheath overwintering diapause building-entry structural-pest nuisance-odor flight-sound droning-flight aggregation-behavior male-pheromone female-choice parasitoid-host biological-control-agent integrated-pest-management monitoring early-detection prevention invasion-biology alien-species non-native-pest global-spread climate-change niche-modeling suitable-habitat establishment-risk quarantine phytosanitary forest-health seed-production reforestation afforestation conifer-forestry pine-seed seed-viability economic-impact crop-loss yield-reduction kernel-damage fruit-damage nut-damage vector-of-disease Chagas-disease kissing-bug public-alarm misinformation education identification-tools health-system-burden pesticide-overuse urban-ecology domiciliary peridomiciliary Andean-region Patagonia Mediterranean temperate subtropical tropical agroecosystem natural-enemy predator parasite pathogen symbiont gut-microbiome soil-ingestion fitness-benefit reproductive-success sperm-morphology testicular-morphology accessory-gland pheromone-blend species-specific-odor cherry-scent vanilla-scent cinnamon-scent rose-scent olfactory-communication mate-location mate-recognition sexual-selection male-male-competition weapon-morphology allometry developmental-plasticity nymphal-instar egg-barrel egg-arrangement linear-oviposition leaf-surface plant-tissue stylet-penetration enzymatic-digestion fluid-feeding phloem-feeding xylem-feeding seed-feeding cone-feeding fruit-feeding kernel-feeding nut-feeding crop-feeding host-plant-range polyphagy oligophagy specialist generalist pest-status damage-threshold economic-injury-level management-strategy cultural-control physical-control chemical-control resistance tolerance host-plant-resistance monitoring-trap light-trap population-density distribution-map spread-rate colonization establishment population-dynamics life-table developmental-rate thermal-requirements degree-days phenology voltinism univoltine bivoltine multivoltine generation-time overwintering-site hibernaculum shelter-seeking cold-hardiness freeze-tolerance supercooling water-relations desiccation-resistance starvation-resistance dispersal-ability flight-capacity walking-behavior climbing-behavior aggregation-pheromone alarm-pheromone stink-gland metathoracic-gland dorsoabdominal-gland ventral-abdominal-gland sexual-gland morphological-defense chemical-defense mechanical-defense autotomy leg-loss predation-risk parasitism-risk parasitoid-risk pathogen-risk competition intraspecific interspecific resource-competition mating-competition sperm-competition cryptic-female-choice reproductive-isolation speciation phylogeny systematics taxonomy morphology anatomy histology ultrastructure spermatogenesis spermatozoa sperm-length nuclear-length cyst-production follicle-number testis-structure reproductive-anatomy genitalia mating-behavior copulation insemination oviposition egg-production fecundity fertility hatching-success nymphal-survival adult-longevity sex-ratio operational-sex-ratio population-sex-ratio effective-population-size genetic-diversity gene-flow population-structure phylogeography biogeography historical-biogeography vicariance dispersal range-shift range-contraction altitudinal-distribution latitudinal-distribution longitudinal-distribution island-distribution continental-distribution endemism cosmopolitanism introduced-range native-range source-population founder-population invasion-front lag-phase establishment-phase spread-phase equilibrium-phase impact-assessment risk-assessment horizon-scanning early-warning rapid-response eradication containment control management adaptation mitigation restoration conservation biodiversity ecosystem-service ecosystem-function food-web trophic-level primary-consumer herbivore fruit-predator plant-animal-interaction mutualism antagonism predation parasitism commensalism amensalism facilitation indirect-interaction trait-mediated-interaction density-mediated-interaction behavioral-ecology evolutionary-ecology functional-ecology physiological-ecology population-ecology community-ecology ecosystem-ecology landscape-ecology macroecology global-ecology applied-ecology agricultural-ecology forest-ecology conservation-biology invasion-ecology restoration-ecology pest-management conservation-biological-control augmentative-biological-control classical-biological-control parasitoid nematode fungus bacterium virus entomopathogen microbial-control sterile-insect-technique genetic-control mating-disruption attract-and-kill push-pull trap-crop border-crop refuge habitat-management cultural-practice crop-rotation tillage irrigation fertilization pruning harvest-timing sanitation residue-management weed-management cover-crop companion-planting intercropping agroforestry silviculture forest-management seed-orchard-management cone-collection seed-extraction seed-testing germination-testing seed-quality seed-vigor empty-seed insect-damage insect-injury feeding-scar puncture stylet-sheath salivary-flange symptom sign diagnosis sampling economic-threshold decision-support expert-system modeling simulation forecasting climate-model niche-model species-distribution-model habitat-suitability risk-mapping invasion-pathway introduction-vector trade transport tourism migration natural-dispersal human-mediated-dispersal accidental-introduction intentional-introduction release escape cultivation ornamental forestry agriculture horticulture urbanization globalization land-use-change habitat-fragmentation habitat-loss degradation pollution pesticide fertilizer nutrient-enrichment eutrophication acidification warming drought extreme-event disturbance succession recovery resilience stability variability uncertainty sustainable-development green-economy circular-economy bioeconomy ecosystem-approach one-health planetary-health environmental-health public-health veterinary-health plant-health food-security nutrition livelihood poverty inequality gender indigenous-knowledge traditional-knowledge local-knowledge citizen-science stakeholder-engagement policy governance regulation legislation international-cooperation capacity-building awareness communication outreach extension advisory-service research innovation technology-transfer knowledge-exchange networking partnership collaboration evaluation evidence-based-policy science-policy-interface diplomacy advocacy activism social-movement environmental-justice environmental-ethics environmental-philosophy environmental-humanities environmental-history environmental-literature environmental-art environmental-education environmental-communication science-communication risk-communication crisis-communication health-communication behavior-change pro-environmental-behavior sustainable-behavior consumption production waste recycling reuse reduction efficiency renewable clean green blue natural organic agroecology permaculture regenerative restorative healing transformative transdisciplinary interdisciplinary multidisciplinary cross-disciplinary disciplinary disciplinary-integration knowledge-integration system-thinking complexity emergence non-linearity feedback tipping-point regime-shift alternative-stable-state resilience-thinking adaptive-management adaptive-governance learning reflection participation inclusion equity justice fairness ethics values norms culture identity place sense-of-place attachment well-being quality-of-life happiness flourishing thriving sustainability sustainable development growth degrowth post-growth steady-state circular doughnut-economics planetary-boundaries safe-operating-space tipping-elements critical-transition early-warning-signal resilience-indicator sustainability-indicator benchmark target goal commitment pledge agreement treaty convention protocol framework strategy plan program project initiative campaign movement coalition alliance network platform hub center institute organization association society union federation confederation foundation trust fund bank facility mechanism instrument tool method approach technique procedure process system structure institution administration operation service delivery provision supply demand market economy finance investment funding financing resource capital asset liability income expenditure revenue cost benefit profit loss return yield outcome output input throughput productivity effectiveness performance quality standard criterion indicator metric measure assessment appraisal review audit verification validation certification accreditation recognition award prize honor distinction excellence best-practice lesson-learned experience expertise competence capacity capability skill knowledge information data evidence science discovery invention creation design testing piloting scaling mainstreaming uptake adoption diffusion dissemination publication reporting documentation archiving preservation rehabilitation reconstruction replication duplication backup security safety protection safeguarding insurance risk-management emergency-preparedness response relief humanitarian peace prosperity health employment housing infrastructure energy water food fishery aquaculture mining manufacturing industry commerce recreation heritage nature ecosystem environment climate atmosphere ocean freshwater land soil mineral biota flora fauna microorganism fungi plant animal vertebrate invertebrate arthropod insect bug Hemipteran Heteropteran pentatomomorphan coreoid coreid squash-bug boxelder-bug stink-bug shield-bug plant-bug mirid lygaeid seed-bug broad-headed-bug alydid reduviid assassin-bug triatomine bed-bug cimicid water-bug nepomorphan gerromorphan leptopodomorphan enicocephalomorphan dipsocoromorphan ceratocomboid schizopteroid peloridioid coleorrhynchan moss-bug archaeorrhynchan fulgoromorphan cicadomorphan membracoid treehopper leafhopper planthopper psyllid jumping-plant-louse whitefly aleyrodid scale-insect coccoid mealybug aphid adelgid phylloxeran sternorrhynchan thysanopteran thrips psocopteran barklouse booklouse phthirapteran louse sucking-louse chewing-louse mallophagan anopluran dermapteran earwig blattodean cockroach termite isopteran mantodean mantid phasmid stick-insect leaf-insect orthopteran grasshopper locust katydid cricket mole-cricket pygmy-mole-cricket camel-cricket cave-cricket weta ensiferan caeliferan gryllotalpid myrmecophilid tettigoniid gryllid acridid pamphagid pneumorid lentulid tristirid eumastacid proscopiid tridactylid tetrigid grouse-locust pygmy-grasshopper plecopteran stonefly embiopteran webspinner zorapteran angel-insect dictyopteran
Leptoglossus zonatus
leaf-footed bug
Leptoglossus zonatus is a polyphagous leaf-footed bug native to the Americas, ranging from the southwestern United States through Central America to South America. Adults measure approximately 20 mm in length with gray coloration, a zigzagging whitish band across the dorsum, and two distinctive yellowish spots on the anterior pronotum that serve as the primary identifying characteristic. The species has emerged as a significant agricultural pest, particularly in California almonds and Gulf Coast satsuma mandarins, causing direct feeding damage and transmitting plant pathogens. Two genetically distinct strains occur in the Western Hemisphere, with low overall genetic diversity suggesting potential for further range expansion.
agricultural-pest invasive-potential polyphagous leaf-footed-bug almond-pest citrus-pest pathogen-vector egg-parasitoid-host pheromone-communication territorial-behavior
Leptomastidea
Leptomastidea is a genus of parasitoid wasps in the family Encyrtidae (Hymenoptera: Chalcidoidea), first described by Mercet in 1916. The genus contains approximately 24 described species worldwide, with members distributed across the Nearctic, Neotropical, Palaearctic, and Oriental regions. Species are primary parasitoids of mealybugs (Hemiptera: Pseudococcidae), with some species also attacking eriococcids (Eriococcidae). Several species, particularly L. abnormis and L. dactylopii, have been widely used in classical and augmentative biological control programs against citrus and vine mealybugs. The genus is classified in the tribe Anagyrini based on morphological and molecular phylogenetic evidence.
parasitoid mealybug biological-control Encyrtidae citrus-pest integrated-pest-management sex-allocation host-selection
Leptomastidea abnormis
Leptomastidea abnormis is an encyrtid parasitoid wasp native to the Nearctic and Neotropical regions, with introduced populations in Europe and elsewhere. It is a primary parasitoid of mealybugs (Pseudococcidae), particularly the citrus mealybug Planococcus citri, and is widely used in biological control programs. Females exhibit size-dependent host selection, preferring second-instar mealybugs but also attacking later instars and adults. The species shows temperature-dependent reproduction, with maximum progeny production at 24°C maintained up to 34°C. Sex allocation follows host quality models, with female-biased sex ratios on larger hosts.
parasitoid mealybug biological-control citrus-pest Encyrtidae sex-allocation host-selection
Litargus tetraspilotus
Four-spotted Hairy Fungus Beetle
Litargus tetraspilotus is a small hairy fungus beetle in the family Mycetophagidae, first described from North America in 1856 and more recently recorded in Brazil. It feeds on powdery mildew fungi (Oidium sp.) and has been studied as a potential biological control agent for fungal pathogens on fruit trees. Population dynamics show seasonal variation tied to humidity and rainfall patterns.
mycophagous powdery-mildew citrus-pest biocontrol-potential Neotropical-expansion
Lopholeucaspis
Lopholeucaspis is a genus of armored scale insects (Diaspididae) containing species that are invasive pests of woody plants. The most documented species, L. japonica, has established populations across Asia, Europe, Australia, and North America. These scales infest a broad range of hosts including citrus, pomegranate, crapemyrtle, privet, and euonymus. The genus is notable for its expanding global distribution and association with multiple parasitoid wasp species.
scale-insect armored-scale invasive-pest phloem-feeder woody-plant-pest citrus-pest ornamental-pest
Mamestra curialis
Scripted Arches Moth, Scripted Arches
Mamestra curialis is a noctuid moth native to North America, first described by John Bernhard Smith in 1887. The species has been documented across western Canada including Alberta, Manitoba, and Saskatchewan. It was historically referred to as the "citrus cutworm" in California agricultural literature, where early-season larvae feed on young citrus fruit, though this common name has also been applied to related species. The species has gained attention in integrated pest management research due to its potential economic impact on citrus production.
Noctuidae cutworm citrus-pest North-America California-agriculture integrated-pest-management early-season-pest fruit-scarring
Marmara gulosa
Citrus Peelminer
Marmara gulosa is a small gracillariid moth, approximately 4 mm in length, commonly known as the citrus peelminer. Native to the United States where it originally fed on willow (Salix lasiolepis), the species has undergone a documented host-shift to non-native plants including citrus varieties and ornamentals such as oleander. It is known from California, Arizona, Texas, and Florida, as well as Cuba. The species is of economic concern as a pest of citrus, particularly grapefruit in southwestern growing regions. A distinct species from the similar Marmara salictella, it was formally described in 2001.
leafminer citrus-pest agricultural-pest Gracillariidae host-shift invasive-behavior
Microcentrum rhombifolium
greater angle-wing katydid, broad-winged katydid, angular-winged katydid
A large, leaf-mimicking katydid native to North America, recognized by its rhombus-shaped green wings and distinctive 'ticking' call. Adults reach 50–65 mm in length and are most active in late summer and autumn. The species is among the most convincing leaf mimics in its family, with broad, reticulate-veined tegmina that provide excellent crypsis in deciduous vegetation. Males produce intermittent calling songs and courtship songs consisting of repeated ticks once a female is attracted.
leaf-mimic acoustic-communication nocturnal arboreal sphecid-wasp-prey citrus-pest
Microterys
Microterys is a large genus of parasitoid wasps in the family Encyrtidae (Chalcidoidea), with its center of distribution in the northern parts of the Northern Hemisphere. Species are important natural enemies of various scale insects (Coccoidea), including soft scales (Coccidae), wax scales (Ceroplastes), and mealybugs (Pseudococcidae). The genus has been extensively studied for biological control applications, particularly for managing pest scale insects on citrus and other crops. Several species have been introduced to new regions as biocontrol agents, including Microterys flavus in California.
parasitoid biological-control scale-insect Coccoidea Encyrtidae Chalcidoidea Hymenoptera ovipositor integrated-pest-management citrus-pest soft-scale mealybug hyperparasitoid host-specificity biotype classical-biological-control synchrotron-microtomography oviposition-behavior natural-enemy agricultural-entomology taxonomy Girault Australia China California Syria India Czech-Republic Europe Asia North-America Southern-Hemisphere Ceroplastes Coccus Phenacoccus Kermes Parthenolecanium Metaceronema Chaetococcus Kerria Laccifer Gascardia Sphaerolecanium Cerococcus Chloropulvinaria pest-management host-selection host-penetration envenomation egg-deposition host-feeding valvifer valvula musculoskeletal-system 3D-morphology SEM SR-µCT invasive-species biocontrol-introduction hyperparasitism efficiency-reduction natural-enemy-conservation citrus Camellia Magnolia fruit-crops ornamental-plants forestry lac-insect wax-scale plum-scale cottony-maple-scale European-fruit-lecanium brown-soft-scale citricola-scale citrus-snow-scale Japanese-wax-scale type-material species-description redescription identification-key intraspecific-variation diagnostic-characters Northern-Hemisphere center-of-distribution cosmopolitan new-species new-record faunistics museum-collection UCR Timberlake Compere Gordh Triapitsyn Noyes Xu Sugonjaev Prinsloo Rosen Ishii Mercet Dalman Howard Ashmead Walker Ferrière Kerrich Annecke Mynhardt Hayat Subba-Rao Trjapitzin Viggiani De-Santis Mayr Thomson Westwood Gahan Crawford Blanchard Perkins Risbec Brues Bennett Erdös Silvestri Domenichini Dozier Cockerell Coquillet Fonscolombe Ratzeburg Matsumura Vassiljev Yoshimoto Ghesquière Tachikawa Logvinovskaya Siscaro Agarwal Ahmad Ghani Burks Girault-species Australian-fauna Chinese-fauna Indian-fauna Syrian-fauna European-fauna Californian-fauna global-distribution economic-entomology applied-entomology systematics morphology behavioral-ecology functional-morphology evolutionary-success diverse-group understudied minute-wasps micro-Hymenoptera slide-collection voucher-specimens host-records undetermined-specimens valid-genera synonymy manuscript-names nomina-nuda unpublished-names re-organization curator historical-notes collection-growth collecting-activities identification sorting drawers points slides paratypes backbone biological-control-projects California-entomologists Citrus-Experiment-Station Hawaiian-collection bee-taxonomy systematists students staff world largest important value mere-number taxa-represented reported scientific-literature conducted augmented contributions countries rapidly growing complete list presented italicized miscellaneous undetermined full-drawers unsorted specimens-per-slide more-than-500-slides at-least-700-slides sometimes-more-than-one-specimen-per-slide synchrotron-X-ray-phase-contrast-microtomography scanning-electron-microscopy in-vivo-documentation 3D-model microstructures cuticular-elements motion-patterns phases parasitization working-mechanism key-trait adaptive-evolution reproductive-success mechanical-aspects functional-aspects fully-understood enormously-diverse specialized parasitising plant-pests ecological-role economic-role improve-understanding mechanics mode-of-function detailed analysis identified consisting recently-discovered described elucidated penetration assessment internal-organs developed underlying observed contributing highly-diverse-group chalcidoid-wasps Frontiers-in-Zoology Acta-Musei-Silesiae Arab-Journal-for-Plant-Protection Australian-Journal-of-Entomology Insect-Science Oriental-Insects Journal-of-Natural-History Phytoparasitica Annals-of-the-Entomological-Society-of-America Environmental-Entomology GBIF Catalogue-of-Life NCBI-Taxonomy iNaturalist Wikipedia DOI 10.1186/s12983-025-00575-1 10.2478/cszma-2021-0007 10.22268/AJPP-41.3.292305 10.1111/j.1440-6055.1975.tb02058.x 10.1111/j.1744-7917.2000.tb00345.x 10.1080/00305316.2011.646827 10.1111/j.1440-6055.1973.tb01670.x 10.1080/00222933.2020.1786608 10.1007/s12600-019-00727-0 10.1007/s12600-018-0691-5 10.1093/aesa/54.2.222 10.1093/ee/7.6.874 2025 2021 2023 1975 2000 2011 1973 2020 2019 2018 1950s 1960s 1876 1999 2016 2017 April June February North-Moravia NE-Czech-Republic Moravskoslezské-Beskydy-Mts Morávka Turkmenistan Central-Asia Lattakia-Province Al-Sanobar Dabba Syrian-coast Damascus-University Biological-Control-Studies-and-Research-Center BCSRC Faculty-of-Agriculture Queensland-Museum Zhejiang Fujian Yunnan Guangdong Northeast-China Western-Australia Jilin first-record new-to-science first-time added-to-fauna reviewed notes illustrations habitus redescriptions close-relationship discussed key-provided females males transferred extent validity evaluated correction behavior response honeydew taxonomy-match higher-rank accepted canonical-name rank status match-type classification Eukaryota Hexapoda Apocrita Terebrantes Encyrtinae observations-count none evidence paper-summary confidence-notes inferred explicitly-stated visible-text full-paper accessible provided-content cited-references detailed-biology type-locality native-range general limited-to based-on no-information-provided abstract genus-level-biology family-level-biology definitive full-text-required inferred-from Japoshvili-1999 Yasnosh-&-Japoshvili-1998 high-hyperparasitoids-levels reduced-efficiency naturally-occurring microscopic-images morphological morphometric-characteristics important-role economic-importance samples collected brought based-on-information-gathered may-play objective following provided illustrate keywords given cannot-be-properly-identified remaining-type-material recorded localities soft-scale-insect-hosts centre-of-distribution northern-parts morphological-diagnostic-characters critically-evaluated material-kept deals-with provinces host paper collected-from host-of-these-parasitoids on Homoptera Coccidoidea described-from as-parasitoid-of from-India associated-with comments host-relationship correction-to new-species-from parasitoid-of new-biological-race introduced-into control-of response-to and-its-honeydew NCBI Animalia Arthropoda Insecta Microterys Thomson-1876 genus higherrank 511-observations no-summary Metazoa wasps-ants-&-bees group matched distribution-records Encyrtid-Holdings Entomology-Research-Museum UCR-collection compiled-by Serguei-V.-Triapitsyn updated February-2000 one-of-the-largest most-important collections 779-valid-species around-1000 261-genera important-slide-collection largest-in-the-world about-8000-slides unpublished numerous-voucher-specimens started-by P.H.-Timberlake numerous-paratypes unfortunately lost-interest switched until-1970s H.-Compere notable-contributions P.-Timberlake J.-Maple P.-DeBach C.-Clausen 1970s-1980s G.-Gordh J.-LaSalle J.-Woolley J.-Pinto J.-Beardsley Mexico US other-countries presently growing-rapidly J.-Heraty majority V.-Trjapitzin J.-Noyes complete-re-organization S.-Triapitsyn current-curator species on-points on-slides undetermined-to-genus about-3-full-drawers Acerophagoides Acerophagus Achalcerinys Adelencyrtoides Adelencyrtus Aenasiella Aenasius Aeptencyrtus Aethognathus Agarwalencyrtus Ageniaspis Aglyptus Agromyzaphagus Alamella Aloencyrtus Ameromyzobia Ammonoencyrtus Anagyrus Ananusia Anasemion Anathrix Anicetus Anomalicornia Anthemus Anusia Anusioptera Aphidencyrtoides Aphycoides Aphycomorpha Aphycopsis Aphycus Apoanagyrus Apoleptomastix Apsilophrys Archinus Arhopoidiella Arrhenophagus Aschitus Astymachus Atropates Austrochoreia Avernes Avetianella Aztecencyrtus Baeoanusia Baeocharis Blastothrix Blepyrus Borrowella Bothriocraera Bothriophryne Bothriothorax Boucekiella Brethesiella Caenocercus Caenohomalopoda Callipteroma Carabunia Ceballosia Cerapterocerella Cerapteroceroides Cerapteroceroideus Cerapterocerus Cerchysiella Cerchysius Cercobelus Charitopus Cheiloneuromyia Cheiloneurus Cheilopsis Choreia Chrysoplatycerus Cicoencyrtus Cirrhencyrtus Clausenia Coccidencyrtus Coccidoctonus Coccidoxenoides Coelopencyrtus Comperia Comperiella Copidosoma Copidosomopsis Cowperia Cryptanusia Cyderius Deilio Diaphorencyrtus Dicarnosis Dinocarsiella Dinocarsis Discodes Diversinervus Dusmetia Echthrogonatopus Echthroplexiella Ectroma Ectromatopsis Encyrtus Eotopus Epanusia Epiblatticida Epiencyrtus Epistenoterys Epitetracnemus Erencyrtus Ericydnus Eucoccidophagus Eugahania Eupoecilopoda Eusemion Exoristobia Forcipestricis Formicencyrtus Fulgoridicida Gahaniella Ginsiana Gyranusoidea Habrolepis Habrolepopteryx Hambletonia Helegonatopus Hemencyrtus Hengata Hesperencyrtus Heterococcidoxenus Hexacladia Hexacnemus Hexencyrtus Holanusomyia Holcencyrtus Holcothorax Homalopoda Homalotyloidea Homalotylus Hoplopsis Hypergonatopus Iceromyia Isodromus Ixodiphagus Lakshaphagus Lamennaisia Leefmansia Leptomastidea Leptomastix Leurocerus Lirencyrtus Lyka Mahencyrtus Manicnemus Mariola Mayridia Meniscocephalus Meromyzobia Mesorhopella Metablastothrix Metanotalia Metaphaenodiscus Metaphycus Mira Monodiscodes Moorella Mucrencyrtus Neastymachus Neblatticida Negeniaspidius Neocharitopus Neocladia Neococcidencyrtus Neodusmetia Neoplatycerus Oesol Oobius Ooencyrtus Oriencyrtus Paksimmondsius Papaka Parablastothrix Parablatticida Paraclausenia Paraenasomyia Paraleurocerus Paralitomastix Paraphaenodiscus Parapyrus Paraschedius Paratetracnemoidea Paratetralophidea Parechthrodryinus Parectromoidella Parectromoides Pareusemion Pentelicus Perpolia Philosindia Plagiomerus Platencyrtus Prionomastix Prionomitus Prochiloneurus Proleurocerus Proleuroceroides Protyndarichoides Pseudaphycus Pseudectroma Pseudencyrtoides Pseudencyrtus Pseudhomalopoda Pseudleptomastix Pseudococcobius Pseudorhopus Psilophryoidea Psilophrys Psyllaephagus Psyllechthrus Raffaellia Rhopalencyrtoidea Rhopus Rhytidothorax Ruandella Saera Sauleia Schilleriella Sectiticlava Subprionomitus Synsemion Syrphophagus Szelenyiola Tachardiaephagus Tachardiobius Tachinaephagus Taftia Tanyencyrtus Tassonia Teleterebratus Tetarticlava Tetracnemoidea Tetracnemus Tetracyclos Thomsonisca Tineophoctonus Trechnites Tremblaya Trichomasthus Tricladia Trjapitzinellus Tropidophryne Tyndarichus Tyndaricopsis Vosleria Xenanusia Xenoencyrtus Yasumatsuiola Zaomma Zaommoencyrtus Zaplatycerus Zarhopaloides Zarhopalus spp.sp.sp(p).?nr.ms-name manuscript-name nomen-nudum including probably-mixed-with unsorted-miscellaneous University-of-California-Riverside collection museum specimens vouchers hosts parasitoids scale-insects mealybugs soft-scales wax-scales citrus-pests agricultural-pests natural-enemies hyperparasitoids efficiency oviposition anatomy 3D-reconstruction microtomography in-vivo-observation evolution adaptive-radiation diversity Africa South-America Middle-East Hawaii horticulture crop-protection augmentative-biological-control conservation-biological-control egg-laying population-dynamics ecosystem-services natural-pest-control sustainable-agriculture organic-farming biodiversity invasive-species-management quarantine phylogeny diagnosis description new-records biogeography museum-collections historical-collections type-specimens curatorial-practices collection-management data-quality scientific-infrastructure research-capacity collaboration knowledge-transfer education outreach public-engagement science-communication open-access data-sharing FAIR-principles digitalization imaging microscopy tomography 3D-modeling computational-biology biomechanics kinematics muscle-physiology neuroethology sensory-ecology chemical-ecology host-location host-recognition host-acceptance host-suitability foraging-behavior learning memory decision-making life-history development reproduction survival mortality population-ecology community-ecology trophic-interactions food-webs ecosystem-functioning ecosystem-stability resilience sustainability environmental-change climate-change land-use-change habitat-fragmentation pollution pesticide-use biological-invasions conservation 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Naupactus
White-fringed Weevils
Naupactus is a large genus of weevils in the family Curculionidae, commonly known as white-fringed weevils. The genus comprises exclusively Neotropical species, several of which have been introduced to other regions and become significant agricultural pests. Members of this genus are characterized by reduced humeri and lack of metathoracic wings, traits formerly used to separate them into the genus Graphognathus before taxonomic revision. Some species exhibit parthenogenetic reproduction, and flightless adults are common.
weevil pest agriculture parthenogenesis flightless invasive Neotropical white-fringed Curculionidae Entiminae Naupactini grape-pest citrus-pest quarantine-pest
Nipaecoccus
Nipaecoccus is a genus of mealybugs in the family Pseudococcidae. The genus includes several economically important pest species, most notably Nipaecoccus viridis (hibiscus mealybug or lebbeck mealybug) and Nipaecoccus nipae (coconut mealybug). Species in this genus are characterized by their white, powdery wax covering and polyphagous feeding habits on a wide range of host plants including citrus, palms, ornamentals, and various fruit and row crops. Nipaecoccus viridis has emerged as a significant invasive pest in Florida since 2009, with documented impacts on citrus, blueberries, hemp, and ornamental plants.
mealybug invasive-pest citrus-pest agricultural-pest Florida polyphagous phloem-feeder scale-insect biological-control-target diagnostic-testing
Novius cardinalis
Vedalia beetle, cardinal ladybird
Novius cardinalis is a small ladybird beetle native to Australia, notable as one of the earliest and most successful examples of classical biological control. The species was introduced to California in 1888 to combat cottony cushion scale (Icerya purchasi), a devastating citrus pest, and achieved rapid, dramatic suppression of the pest population. Adults measure 2.5–4 mm with variable red and black coloration. The species has undergone multiple taxonomic revisions, most recently transferred from Rodolia to Novius in 2020.
biological-control citrus-pest specialist-predator Australia ladybird-beetle
Oncometopia
sharpshooters
Oncometopia is a genus of sharpshooters (leafhoppers in the family Cicadellidae) found in North and South America, erected by Carl Stål in 1869. Species range from 9.4 to 15.0 mm in length and are characterized by distinctive morphological features including a moderately produced head, coriaceous forewings with apical membranes, and specialized hindleg tarsomere proportions. Several species are economically important as vectors of plant pathogens, particularly Xylella fastidiosa, which causes diseases such as phony peach disease and citrus variegated chlorosis.
sharpshooter leafhopper Cicadellidae Proconiini xylem-feeder vector Xylella-fastidiosa brochosomes honeydew pest North-America South-America citrus-pest disease-vector agricultural-pest
Pachnaeus
broad-nosed weevils, citrus root weevils
Pachnaeus is a genus of broad-nosed weevils comprising approximately seven described species. Several species, notably P. litus and P. opalus, are economically significant pests of citrus, commonly referred to as citrus root weevils. The genus has been targeted for biological control research, with studies evaluating parasitoid wasps for management of P. litus populations.
citrus-pest root-weevil biological-control agricultural-pest Entiminae Eustylini
Pachnaeus litus
blue-green citrus root weevil, blue-green citrus weevil
Pachnaeus litus is a broad-nosed weevil in the family Curculionidae, commonly known as the blue-green citrus root weevil or blue-green citrus weevil. It is a pest of citrus in North America, with eggs that serve as hosts for multiple parasitoid wasp species. The species has been studied extensively for biological control potential, particularly targeting its egg stage with parasitoids such as Brachyufens osborni (Trichogrammatidae), Pediobius irregularis, and Horismenus bennetti (Eulophidae).
citrus-pest biological-control egg-parasitoid-host Curculionidae Entiminae agricultural-entomology
Pachnaeus opalus
Northern Citrus Root Weevil
Pachnaeus opalus is a broad-nosed weevil in the family Curculionidae, commonly known as the northern citrus root weevil. It is native to the southeastern United States and has been documented in Alabama, Florida, Georgia, Mississippi, and New Jersey. The species is associated with citrus and related plants, where adults feed on foliage and larvae develop in roots. It was first described by Olivier in 1807 as Curculio opalus.
citrus-pest root-weevil agricultural-pest broad-nosed-weevil southeastern-US
Panonychus
spider mites
Panonychus is a genus of spider mites in the family Tetranychidae, established by Yokoyama in 1929. The genus comprises approximately 16 described species, including several economically significant agricultural pests. Notable species include Panonychus citri (citrus red mite), a major pest of citrus crops, and Panonychus ulmi (European red mite or fruit tree red spider mite), which attacks deciduous fruit and forest trees. These mites are characterized by their phytophagous feeding habits and have developed resistance to numerous pesticides.
agricultural-pest citrus-pest pesticide-resistance biological-control spider-mite
Papilio
Common Swallowtails
Papilio is a large genus of swallowtail butterflies containing approximately 200 species, representing the sole genus in the tribe Papilionini. The genus includes some of the most recognizable butterflies worldwide, such as the Old World swallowtail (P. machaon), western tiger swallowtail (P. rutulus), and eastern tiger swallowtail (P. glaucus). Modern taxonomic revisions have consolidated numerous former genera—including Pterourus (tiger swallowtails), Heraclides (giant swallowtails), and Achillides—into Papilio, making it one of the most diverse butterfly genera. The name derives from the Latin word for butterfly.
butterfly swallowtail Lepidoptera Papilionidae pollinator citrus-pest mimicry bird-dropping-mimicry snake-mimicry
Papilio anchisiades
ruby-spotted swallowtail, red-spotted swallowtail
Papilio anchisiades is a medium to large swallowtail butterfly with a wingspan of 70–100 mm, distributed from southern Texas through Central America to Argentina. The species exhibits considerable geographic variation across its range, with multiple recognized subspecies. Larvae feed on Rutaceae, particularly Citrus species, and adults are commonly observed drinking at moist ground near rivers rather than in forested habitats.
swallowtail Rutaceae-feeder riparian gregarious-larvae butterfly-farming citrus-pest
Papilio ornythion
ornythion swallowtail
Papilio ornythion is a swallowtail butterfly in the subfamily Papilioninae, native to Mexico and Guatemala with occasional vagrancy into the southern United States. The species exhibits sexual dimorphism with two female forms—one matching the male and a distinctive dark form. Adults fly from April to September with two generations per year. Larvae feed on Citrus leaves while adults nectar on flowers.
swallowtail Citrus-pest sexual-dimorphism vagrant Mexico Guatemala Papilio-thoas-group
Paraleyrodes
nesting whiteflies, Bondar's nesting whitefly (for P. bondari), fig whitefly (for P. minei)
Paraleyrodes is a genus of whiteflies in the subfamily Aleurodicinae, commonly known as nesting whiteflies due to the woolly wax nests constructed by females around egg masses. The genus includes economically significant invasive species such as P. bondari (Bondar's nesting whitefly) and P. minei, which are pests of coconut, citrus, fig, avocado, and ornamental plants. Species in this genus are characterized by distinctive waxy secretions, sessile nymphal stages, and polyphagous feeding habits on woody plants.
whitefly invasive-pest agricultural-pest ornamental-pest Aleurodicinae coconut-pest citrus-pest fig-pest avocado-pest woolly-wax-nests polyphagous phloem-feeder honeydew-producer sooty-mold Florida-pest California-pest India-pest Bangladesh-pest Croatia-pest Greece-pest natural-enemies biological-control integrated-pest-management
Phyllocnistis citrella
Citrus Leafminer, CLM
Phyllocnistis citrella, commonly known as the citrus leafminer or CLM, is a small moth in the family Gracillariidae. Native to tropical Asia, it has become a globally distributed agricultural pest of citrus since its discovery in Florida in 1993. The larvae create distinctive serpentine mines in citrus leaves, protected within the leaf epidermis from many topical insecticide treatments. The species is managed through integrated pest management strategies including biological control with introduced parasitoids, mating disruption using sex pheromones, and cultural practices.
agricultural-pest leaf-miner citrus-pest biological-control mating-disruption invasive-species integrated-pest-management
Planococcus
mealybugs
Planococcus is a genus of mealybugs in the family Pseudococcidae, comprising economically significant agricultural pests. The genus includes species such as Planococcus citri (citrus mealybug), Planococcus ficus (vine mealybug), and Planococcus minor, which infest a wide range of host plants including citrus, grapevines, and numerous horticultural crops. These insects cause direct damage through phloem feeding and indirect damage via honeydew secretion that promotes sooty mold growth; several species are also known vectors of plant viruses. The genus is characterized by small, soft-bodied insects covered in white, powdery wax secretions, with complex taxonomic histories involving multiple synonymies and cryptic species complexes.
mealybug agricultural-pest vineyard-pest citrus-pest phloem-feeder virus-vector biological-control-target
Planococcus citri
citrus mealybug
Planococcus citri, the citrus mealybug, is a highly polyphagous pest native to Asia that has established populations worldwide through human-mediated transport. Adult females are approximately 3 mm long, wingless, and covered in white wax with marginal filaments; males are slightly larger with functional wings. The species reproduces sexually with multiple generations per year, producing ovisacs containing up to 20 eggs. It feeds on phloem sap from over 200 host plant species across 82 families, causing direct damage through sap removal and indirect damage via honeydew secretion and sooty mold growth. It is also a known vector of Cacao swollen shoot virus. The species is difficult to distinguish morphologically from the vine mealybug (Planococcus ficus), requiring microscopic examination of pore and duct arrangements or molecular methods for accurate identification.
mealybug citrus-pest polyphagous invasive-species agricultural-pest phloem-feeder virus-vector greenhouse-pest
Platynota rostrana
Omnivorous Platynota Moth
Platynota rostrana is a small tortricid moth with a wingspan of 13–17 mm, distributed across the Americas from the United States through Central America to South America, with established populations in the West Indies and occasional records in Europe via accidental importation. The species is known to be a pest of citrus and other crops, with larvae that feed on a variety of plant materials.
tortricidae leafroller agricultural-pest citrus-pest neotropical nearctic introduced-europe small-moth
Prodiplosis
Prodiplosis is a genus of gall midges (Cecidomyiidae) comprising approximately 11 described species. Several species are economically significant agricultural pests, notably P. longifila, which infests citrus, solanaceous crops, and ornamental foliage. Members of this genus are characterized by complete metamorphosis and larval feeding on reproductive and vegetative tissues of host plants, causing necrosis and yield loss. Some species exhibit host specialization and cryptic genetic differentiation across geographic ranges.
agricultural-pest gall-midge complete-metamorphosis nocturnal cryptic-speciation biological-control citrus-pest solanaceous-pest foliage-crop-pest
Pseudococcus longispinus
long-tailed mealybug, longtailed mealybug
Pseudococcus longispinus is a cosmopolitan mealybug pest in the family Pseudococcidae. It is distinguished by two long, waxy filaments extending from the posterior abdomen of adult females. The species is a significant agricultural and horticultural pest, particularly damaging to grapevines, citrus, orchids, and tropical fruits. It is a known vector of Grapevine leafroll-associated virus 3 (GLRaV-3), transmitting the virus only during the first instar crawler stage. The species has been observed to develop three generations annually in temperate regions, with complex dispersal patterns shifting from exposed foliage in early instars to protected bark and fruit clusters for reproduction.
agricultural-pest virus-vector mealybug greenhouse-pest vineyard-pest citrus-pest biological-control-target
Resseliella
Resseliella is a genus of gall midges in the family Cecidomyiidae containing at least 50 described species. The genus includes economically significant agricultural pests, notably R. maxima (soybean gall midge) which threatens soybean production in the north-central United States, as well as R. citrifrugis, a major citrus pest in China, and R. theobaldi (raspberry cane midge), an important pest of cultivated red raspberry in Europe. Some species exhibit mutualistic relationships with plants, such as R. kadsurae which pollinates Kadsura longipedunculata in China.
gall-midge Cecidomyiidae agricultural-pest soybean-gall-midge citrus-pest raspberry-pest pollinator biological-control parasitoid-host fire-blight-vector genome-sequenced emerging-pest integrated-pest-management
Saissetia
soft scales, black scales
Saissetia is a genus of soft scale insects (Hemiptera: Coccidae) containing at least four described species. The genus includes economically important pests, notably Saissetia oleae (black scale) and Saissetia coffeae (hemispherical scale), which attack citrus, olive, coffee, and numerous ornamental plants. Saissetia species are phloem-feeding insects that produce honeydew, facilitating ant associations and sooty mold growth. Several species have been introduced globally through trade in infested nursery plants.
agricultural-pest biological-control scale-insect honeydew-producer ant-tended invasive-species citrus-pest olive-pest polyphagous
Saissetia coffeae
hemispherical scale, helmet scale, coffee brown scale
Saissetia coffeae is a polyphagous soft scale insect (Coccidae) with a cosmopolitan tropical and subtropical distribution. Adults are sessile, hemispherical, and helmet-shaped, ranging 2–4.5 mm in diameter depending on host plant. The species reproduces parthenogenetically; all individuals are female. It is a significant agricultural pest of coffee, citrus, avocado, fig, guava, and numerous ornamental plants, causing damage through phloem sap feeding and honeydew secretion that promotes sooty mold growth. Multiple parasitoid wasps and predatory insects are associated with natural population regulation.
scale-insect soft-scale agricultural-pest polyphagous parthenogenetic biological-control honeydew-producer sooty-mold ant-mutualism coffee-pest citrus-pest tropical-pest
Scirtothrips
leaf-feeding thrips, citrus thrips, chilli thrips
Scirtothrips is a genus of leaf-feeding thrips in the family Thripidae, established by Shull in 1909. The genus includes economically significant pest species such as the chilli thrips (S. dorsalis), citrus thrips (S. citri), and South African citrus thrips (S. aurantii), which damage agricultural crops through piercing-sucking feeding on tender leaves, buds, and fruit. Species exhibit diverse host associations ranging from highly host-specific forms on endemic Australian Acacia to broadly polyphagous invasive pests. The genus has undergone taxonomic revision, with Labiothrips synonymised under Scirtothrips.
thrips Thripidae agricultural-pest invasive-species leaf-feeding citrus-pest tea-pest strawberry-pest blueberry-pest IPM-target Australia Asia polyphagous host-specific
Scudderia
Scudder's Bush Katydids, bush katydids
Scudderia is a genus of bush katydids in the subfamily Phaneropterinae, established by Stål in 1873. Species in this genus are medium-sized insects, 30–40 mm in length, with a transcontinental distribution across North America. They are herbivorous, feeding on flowers as nymphs and woody deciduous plants as adults. Several species are economically significant as pests in citrus orchards and stone fruit production, particularly Scudderia furcata, the fork-tailed bush katydid. Males produce species-specific calling songs using file-and-scraper structures on their forewings, and species identification typically requires examination of the male supra-anal plate morphology.
katydid Tettigoniidae Phaneropterinae Orthoptera bush-katydid pest citrus-pest acoustic-communication North-America herbivore nocturnal agricultural-pest
Trioza
psyllids, jumping plant-lice
Trioza is the type genus of psyllids in the family Triozidae, comprising sap-sucking insects with worldwide distribution. Species in this genus exhibit diverse host associations, ranging from agricultural crops to native trees. Several species are economically significant: Trioza erytreae transmits citrus greening disease (Huanglongbing), T. apicalis is a major pest of carrot in northern Europe, and T. brevigenae is an invasive pest of ornamental Ficus in California. The genus is characterized by distinct morphological features and complex life cycles involving seasonal migration between summer hosts and overwintering sites.
psyllid sap-sucking phloem-feeder agricultural-pest disease-vector invasive-species quarantine-pest carrot-pest citrus-pest ornamental-pest seasonal-migration host-alternating Triozidae Hemiptera Sternorrhyncha
Unaspis
Unaspis is a genus of armored scale insects (family Diaspididae) containing economically significant agricultural and horticultural pests. The genus includes at least 19 described species, with several species infesting citrus crops (U. citri, U. yanonensis) and ornamental plants (U. euonymi). Species in this genus are characterized by their protective waxy coverings and phytophagous feeding habits on host plant tissues.
armored-scale Diaspididae agricultural-pest horticultural-pest citrus-pest biological-control invasive-species Asia global-distribution
Unaspis citri
Citrus Snow Scale, Orange Chionaspi, Orange Snow Scale, White Louse Scale, White Snow Scale
Unaspis citri is an armored scale insect (Hemiptera: Diaspididae) that infests citrus crops worldwide. The species forms dense colonies on bark, fruit, and leaves, causing direct damage through sap feeding and indirect harm by promoting sooty mold growth. It is economically significant in citrus-producing regions, particularly Florida, where management requires integrated approaches combining monitoring, biological control, and targeted chemical applications.
armored-scale citrus-pest economic-pest phloem-feeder cosmopolitan-distribution agricultural-pest