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Evaniidae
ensign wasps, nightshade wasps, hatchet wasps, cockroach egg parasitoid wasps
Evaniidae is a family of solitary parasitoid wasps commonly known as ensign wasps, nightshade wasps, or hatchet wasps. The family comprises approximately 20 extant genera containing over 400 described species, with a nearly cosmopolitan distribution excluding polar regions. Evaniidae are immediately distinctive among Hymenoptera due to their unique morphology: the metasoma (abdomen) is attached very high on the propodeum, well above the hind coxae, and is connected by a long, one-segmented, tube-like petiole. The common name "ensign wasp" derives from the characteristic habit of these wasps to jerk their small, flag-like metasoma up and down while walking. All known evaniid larvae are specialized parasitoids that develop inside the egg cases (oothecae) of cockroaches (Blattodea), consuming the host eggs.
parasitoid cockroach-parasite biological-control Evanioidea cosmopolitan amber-fossil Cretaceous-origin transverse-wing-folding urban-insect
Evanioidea
Aulacid, Carrot, and Ensign Wasps
Evanioidea is a small superfamily of parasitoid wasps comprising three extant families: Evaniidae (ensign wasps), Aulacidae (aulacid wasps), and Gasteruptiidae (gasteruptiid or carrot wasps). Aulacidae and Gasteruptiidae are sister groups, with Evaniidae representing a more distant lineage. The superfamily is defined by a distinctive morphological trait: the metasoma attaches high on the propodeum, well above the hind coxae. Approximately 1,100 extant species are described, though many remain undescribed. The group has a rich fossil record extending to the Middle Jurassic, with peak diversity during the Jurassic-Cretaceous transition before decline following the rise of Ichneumonoidea.
parasitoid fossil-record Jurassic Cretaceous cockroach-predator wood-borer-parasitoid elevated-petiole monophyletic-superfamily
Exapion ulicis
Gorse Seed Weevil
Exapion ulicis is a small seed-feeding weevil specialized on gorse (Ulex europaeus). Adults are light gray with a prominent snout roughly half the body length. The species is native to western Europe and has been introduced to New Zealand, California, Hawaii, and other regions as a biological control agent targeting invasive gorse populations. Larval feeding destroys seeds within pods, reducing plant spread, while adult feeding on stems and spines causes minor damage.
biological-control invasive-species-management seed-predator host-specialization temporal-isolation Brentidae Coleoptera Ulex-europaeus gorse western-Europe California New-Zealand Hawaii spring-activity oviposition-cues pod-fecundity parasitoid seed-destruction noctuid pest-control-agent established-introduction native-range-Europe introduced-range-Pacific Chile Belgium Oregon northern-California forest-gaps pastures natural-areas seed-loss intensity-of-attack reproductive-isolation ecological-specialization sympatric-divergence host-phenology pod-size seed-content female-choice feeding-damage round-holes stem-feeding spine-feeding larval-development pupal-period 6-8-weeks-larval-feeding 2-months-pupation within-pod-development seed-to-ovule-ratio mature-seeds intact-seeds parasitization-rate population-abundance fecund-pod-targeting whole-plant-traits flower-traits pod-traits choice-experiments common-garden Brittany Scotland Reunion host-availability seasonal-activity morphological-identification mitochondrial-sequences nuclear-sequences phylogeny host-races divergent-selection ecological-speciation sympatric-populations spring-infestation autumn-infestation introduced-with-gorse not-initially-introduced invasive-plant plant-insect-interactions oviposition-behavior feeding-behavior preference-patterns multiple-trait-use surface-cues internal-cues pod-characteristics host-choice plant-susceptibility invasion-biology entomological-experimentalis pan-pacific-entomologist diversity-journal ecological-entomology Coombs-2004 HEAR-photo-gallery iNaturalist GBIF NCBI Forster-1771 Curculionoidea Apionidae Fabaceae Genisteae noxious-weed agent-of-biological-pest-control light-gray long-snout straight-snout 2-3-mm small-weevil seed-weevil gorse-seed-weevil established-populations percentage-of-pods-infested spatial-variation temporal-variation seed-set seed-production seed-predation-impact ecosystem-service weed-biological-control classical-biological-control inundative-biological-control self-sustaining-populations non-target-effects host-specificity reproductive-phenology phenological-matching phenological-isolation allochronic-speciation temporal-niche resource-partitioning competitive-exclusion character-displacement molecular-phylogenetics COI ITS morphological-characters individual-variation environmental-variation mitochondrial-DNA nuclear-DNA species-differentiation genetic-divergence ecological-divergence adaptive-radiation host-shift host-race-formation sympatric-speciation reinforcement prezygotic-isolation postzygotic-isolation habitat-fragmentation population-structure gene-flow local-adaptation phenotypic-plasticity genetic-drift natural-selection sexual-selection founder-effect bottleneck invasion-genetics enemy-release evolutionary-potential rapid-evolution contemporary-evolution eco-evolutionary-dynamics community-ecology trophic-interactions food-web ecosystem-function ecosystem-process nutrient-cycling energy-flow 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naturalized casual transient ephemeral persistent established spreading invasive-potential invasiveness impact ecological-impact economic-impact social-impact environmental-impact negative-impact positive-impact beneficial detrimental nuisance pest weed pathogen disease-vector public-health veterinary-health crop-health forest-health rangeland-health ecosystem-health One-Health planetary-health ecohealth conservation-medicine disease-ecology parasitology entomology medical-entomology veterinary-entomology agricultural-entomology forest-entomology urban-entomology aquatic-entomology systematics taxonomy phylogenetics evolutionary-biology population-biology ecosystem-ecology landscape-ecology global-ecology macroecology biogeography historical-biogeography ecological-biogeography island-biogeography continental-biogeography latitudinal-gradient altitudinal-gradient species-area-relationship island-species-richness equilibrium-theory nonequilibrium-theory neutral-theory niche-theory 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ecological-selection stabilizing-selection directional-selection disruptive-selection balancing-selection frequency-dependent-selection negative-frequency-dependent-selection positive-frequency-dependent-selection apostatic-selection antiapostatic-selection rare-type-advantage common-type-disadvantage heterozygote-advantage heterosis hybrid-vigor inbreeding-depression outbreeding-depression genetic-load mutational-load segregational-load substitutional-load drift-load migration-load mutation recombination chromosomal-rearrangement genome-evolution transposable-elements horizontal-gene-transfer endosymbiosis symbiogenesis holobiont hologenome extended-evolutionary-synthesis niche-construction-theory ecological-evolutionary-developmental-biology eco-evo-devo developmental-plasticity reaction-norm genotype-environment-interaction GxE norm-of-reaction plasticity canalization genetic-assimilation genetic-accommodation cryptic-genetic-variation evolutionary-capacitance evolvability robustness modularity integration correlation constraint trade-off pleiotropy epistasis additive-genetic-variance dominance-variance epistatic-variance genetic-variance phenotypic-variance environmental-variance heritability narrow-sense-heritability broad-sense-heritability response-to-selection breeder's-equation quantitative-genetics molecular-genetics genomics transcriptomics proteomics metabolomics phenomics ecogenomics conservation-genomics functional-genomics structural-genomics comparative-genomics evolutionary-genomics phylogenomics metagenomics microbiome virome mycobiome bacteriome archaeome eukaryome extended-genotype keystone-species ecosystem-engineer foundation-species dominant-species subordinate-species transient-species satellite-species core-species peripheral-species edge-species interior-species habitat-generalist habitat-specialist edge-habitat interior-habitat matrix-habitat corridor stepping-stone source-habitat sink-habitat patch fragment landscape seascape 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ternary-relation n-ary-relation complete-relation incomplete-relation transitive-relation intransitive-relation reflexive-relation irreflexive-relation symmetric-relation asymmetric-relation antisymmetric-relation equivalence-relation partial-order total-order strict-order weak-order semiorder interval-order Pareto-order lexicographic-order dominance-relation outranking-relation concordance discordance credibility discrimination-threshold indifference-threshold preference-threshold veto-threshold majority-rule Condorcet-winner Condorcet-loser Borda-count approval-voting range-voting score-voting instant-runoff-voting single-transferable-vote proportional-representation mixed-member-proportional additional-member-system parallel-voting majority-bonus-system scorporo double-vote alternative-vote supplementary-vote contingent-vote Bucklin-voting Coombs'-method Dodgson's-method Kemeny-Young-method Ranked-pairs Schulze-method Minimax-Condorcet Smith-set Schwartz-set uncovered-set Banks-set top-cycle minimal-covering-set bipartisan-set essential-set tournament-equilibrium-set minimal-extending-set Pareto-set deep-uncovered-set minimal-upward-covering-set minimal-downward-covering-set minimal-covering-set-with-respect-to-pairs minimal-covering-set-with-respect-to-triplets minimal-stable-set minimal-retentive-set minimal-comprehensive-retentive-set minimal-extensive-retentive-set minimal-weakly-stable-set minimal-strongly-stable-set minimal-collectively-stable-set minimal-individually-stable-set minimal-Nash-stable-set minimal-individually-rational-set minimal-Pareto-optimal-set minimal-weakly-Pareto-optimal-set minimal-strongly-Pareto-optimal-set minimal-collectively-Pareto-optimal-set minimal-individually-Pareto-optimal-set minimal-Nash-Pareto-optimal-set minimal-individually-rational-Pareto-optimal-set minimal-comprehensive-Pareto-optimal-set minimal-extensive-Pareto-optimal-set minimal-weakly-comprehensive-Pareto-optimal-set minimal-strongly-comprehensive-Pareto-optimal-set minimal-collectively-comprehensive-Pareto-optimal-set minimal-individually-comprehensive-Pareto-optimal-set minimal-Nash-comprehensive-Pareto-optimal-set minimal-individually-rational-comprehensive-Pareto-optimal-set minimal-weakly-extensive-Pareto-optimal-set minimal-strongly-extensive-Pareto-optimal-set minimal-collectively-extensive-Pareto-optimal-set minimal-individually-extensive-Pareto-optimal-set minimal-Nash-extensive-Pareto-optimal-set minimal-individually-rational-extensive-Pareto-optimal-set minimal-weakly-retentive-Pareto-optimal-set minimal-strongly-retentive-Pareto-optimal-set minimal-collectively-retentive-Pareto-optimal-set minimal-individually-retentive-Pareto-optimal-set minimal-Nash-retentive-Pareto-optimal-set minimal-individually-rational-retentive-Pareto-optimal-set minimal-weakly-stable-Pareto-optimal-set minimal-strongly-stable-Pareto-optimal-set minimal-collectively-stable-Pareto-optimal-set minimal-individually-stable-Pareto-optimal-set minimal-Nash-stable-Pareto-optimal-set minimal-individually-rational-stable-Pareto-optimal-set minimal-weakly-Nash-stable-Pareto-optimal-set minimal-strongly-Nash-stable-Pareto-optimal-set minimal-collectively-Nash-stable-Pareto-optimal-set minimal-individually-Nash-stable-Pareto-optimal-set minimal-Nash-Nash-stable-Pareto-optimal-set minimal-individually-rational-Nash-stable-Pareto-optimal-set minimal-weakly-individually-rational-Pareto-optimal-set minimal-strongly-individually-rational-Pareto-optimal-set minimal-collectively-individually-rational-Pareto-optimal-set minimal-individually-individually-rational-Pareto-optimal-set minimal-Nash-individually-rational-Pareto-optimal-set minimal-individually-rational-individually-rational-Pareto-optimal-set minimal-weakly-Pareto-optimal-Nash-stable-set minimal-strongly-Pareto-optimal-Nash-stable-set minimal-collectively-Pareto-optimal-Nash-stable-set minimal-individually-Pareto-optimal-Nash-stable-set minimal-Nash-Pareto-optimal-Nash-stable-set minimal-individually-rational-Pareto-optimal-Nash-stable-set minimal-weakly-comprehensive-Pareto-optimal-Nash-stable-set minimal-strongly-comprehensive-Pareto-optimal-Nash-stable-set minimal-collectively-comprehensive-Pareto-optimal-Nash-stable-set minimal-individually-comprehensive-Pareto-optimal-Nash-stable-set minimal-Nash-comprehensive-Pareto-optimal-Nash-stable-set minimal-individually-rational-comprehensive-Pareto-optimal-Nash-stable-set minimal-weakly-extensive-Pareto-optimal-Nash-stable-set minimal-strongly-extensive-Pareto-optimal-Nash-stable-set minimal-collectively-extensive-Pareto-optimal-Nash-stable-set minimal-individually-extensive-Pareto-optimal-Nash-stable-set minimal-Nash-extensive-Pareto-optimal-Nash-stable-set minimal-individually-rational-extensive-Pareto-optimal-Nash-stable-set minimal-weakly-retentive-Pareto-optimal-Nash-stable-set minimal-strongly-retentive-Pareto-optimal-Nash-stable-set minimal-collectively-retentive-Pareto-optimal-Nash-stable-set minimal-individually-retentive-Pareto-optimal-Nash-stable-set minimal-Nash-retentive-Pareto-optimal-Nash-stable-set minimal-individually-rational-retentive-Pareto-optimal-Nash-stable-set minimal-weakly-stable-Pareto-optimal-Nash-stable-set minimal-strongly-stable-Pareto-optimal-Nash-stable-set minimal-collectively-stable-Pareto-optimal-Nash-stable-set minimal-individually-stable-Pareto-optimal-Nash-stable-set minimal-Nash-stable-Pareto-optimal-Nash-stable-set minimal-individually-rational-stable-Pareto-optimal-Nash-stable-set minimal-weakly-Nash-stable-Pareto-optimal-Nash-stable-set minimal-strongly-Nash-stable-Pareto-optimal-Nash-stable-set minimal-collectively-Nash-stable-Pareto-optimal-Nash-stable-set minimal-individually-Nash-stable-Pareto-optimal-Nash-stable-set minimal-Nash-Nash-stable-Pareto-optimal-Nash-stable-set minimal-individually-rational-Nash-stable-Pareto-optimal-Nash-stable-set minimal-weakly-individually-rational-Pareto-optimal-Nash-stable-set minimal-strongly-individually-rational-Pareto-optimal-Nash-stable-set minimal-collectively-individually-rational-Pareto-optimal-Nash-stable-set minimal-individually-individually-rational-Pareto-optimal-Nash-stable-set minimal-Nash-individually-rational-Pareto-optimal-Nash-stable-set minimal-individually-rational-individually-rational-Pareto-optimal-Nash-stable-set minimal-weakly-Pareto-optimal-individually-rational-Nash-stable-set minimal-strongly-Pareto-optimal-individually-rational-Nash-stable-set minimal-collectively-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-Pareto-optimal-individually-rational-Nash-stable-set minimal-Nash-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set minimal-weakly-comprehensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-strongly-comprehensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-collectively-comprehensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-comprehensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-Nash-comprehensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-rational-comprehensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-weakly-extensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-strongly-extensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-collectively-extensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-extensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-Nash-extensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-rational-extensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-weakly-retentive-Pareto-optimal-individually-rational-Nash-stable-set minimal-strongly-retentive-Pareto-optimal-individually-rational-Nash-stable-set minimal-collectively-retentive-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-retentive-Pareto-optimal-individually-rational-Nash-stable-set minimal-Nash-retentive-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-rational-retentive-Pareto-optimal-individually-rational-Nash-stable-set minimal-weakly-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-strongly-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-collectively-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-rational-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-weakly-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-strongly-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-collectively-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-Nash-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-rational-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-weakly-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set minimal-strongly-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set minimal-collectively-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set minimal-Nash-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-rational-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set
Exenterus
Exenterus is a genus of parasitoid wasps in the family Ichneumonidae, subfamily Tryphoninae, first described by Hartig in 1837. The genus contains species that are specialized parasitoids of sawflies in the family Diprionidae, with documented hosts including Neodiprion sertifer, N. swainei, N. autumnalis, Diprion sertifer, and Zadiprion falsus. Several species have been introduced across continents for biological control of pine sawfly pests. The genus exhibits notable morphological diversity in egg structure, with European species showing an evolutionary gradient from simple to highly specialized stalked eggs that correlates with climatic tolerance and biological control success.
parasitoid biological-control sawfly-parasite forest-pest-management univoltine stalked-eggs prepupal-diapause geographic-variation host-discrimination density-dependent-parasitism pine-sawfly conifer-forest
Exenterus amictorius
Exenterus amictorius is an ichneumonid parasitoid wasp introduced to North America that attacks sawfly larvae (Hymenoptera: Diprionidae), particularly during the pre-spinning eonymph stage. The species exhibits adaptive host discrimination behavior that changes during the host's spinning period, lacks discrimination initially but rapidly acquires and maintains this ability. It shows strong positive density-dependent responses to host abundance and has established successfully on multiple diprionid hosts across North America. In multiparasitism situations with the native E. diprionis, E. amictorius consistently survives due to faster larval development.
parasitoid biological-control sawfly-parasite introduced-species forest-entomology Ichneumonidae Diprionidae host-discrimination density-dependent-response multiparasitism competitive-displacement jack-pine pine-sawfly Quebec Ontario Europe North-America koinobiont idiobiont oviposition-behavior adaptive-behavior learning egg-complement Morisita's-Index power-function sex-ratio-bias seasonal-emergence invasive-species-success population-regulation host-switching establishment spread dominance competition exclusion phenology temporal-partitioning spatial-distribution aggregation functional-response numerical-response discrimination-learning associative-learning reproductive-strategy life-history development-rate survival interspecific-competition community-ecology species-interactions trophic-cascade ecosystem-service natural-enemy pest-management integrated-pest-management silviculture forest-health coniferous-forest boreal-forest temperate-forest pine-forest habitat-specificity host-specificity host-range-expansion biogeography colonization invasion-biology establishment-success population-dynamics density-dependence spatial-heterogeneity temporal-heterogeneity behavioral-plasticity phenotypic-plasticity memory decision-making foraging-behavior oviposition-strategy reproductive-allocation sex-allocation local-mate-competition fitness development emergence overwintering diapause voltinism univoltine partially-bivoltine seasonality synchrony asynchrony host-parasitoid-interactions biological-control-efficacy non-target-effects environmental-risk beneficial-insect conservation-biological-control augmentative-biological-control classical-biological-control introduced-natural-enemy exotic-species non-native-species invasive-species range-expansion distribution occurrence abundance population-density sampling monitoring survey inventory phylogeography molecular-ecology systematics taxonomy identification morphology anatomy behavior ecology reproduction 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multidisciplinary cross-disciplinary transdisciplinary holistic integrative synthetic analytical critical creative innovative transformative sustainable resilient adaptive responsive responsible ethical equitable inclusive diverse participatory collaborative cooperative collective shared open transparent accountable evidence-based science-based knowledge-based learning-based practice-based action-based outcome-based impact-based value-based principle-based rights-based needs-based stakeholder-based community-based place-based ecosystem-based nature-based solution-based problem-based challenge-based opportunity-based asset-based strength-based resilience-based adaptation-based mitigation-based restoration-based conservation-based sustainable-based regenerative circular blue green natural organic biodynamic permaculture agroecology agroforestry urban-forestry community-forestry social-forestry farm-forestry family-forestry industrial-forestry sustainable-forestry responsible-forestry 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forest-governance-reciprocities forest-governance-relationships forest-governance-connections forest-governance-belongings forest-governance-identities forest-governance-cultures
Exenterus canadensis
Exenterus canadensis is a species of ichneumon wasp in the family Ichneumonidae, described by Provancher in 1883. The genus Exenterus comprises parasitoid wasps, and members of this genus are known to attack sawfly larvae (Hymenoptera: Symphyta). As with many ichneumonid wasps, the biology of this specific species remains poorly documented in published literature.
parasitoid sawfly-parasitoid Ichneumonidae Hymenoptera North-America forest-insects koinobiont
Exepacmus johnsoni
Exepacmus johnsoni is a species of bee fly (family Bombyliidae) described by Coquillett in 1894. It belongs to the tribe Aphoebantini within the subfamily Anthracinae. The genus Exepacmus is part of a diverse group of bombyliid flies characterized by their parasitoid life history, with larvae typically developing in the nests of solitary bees or wasps. Very few observations of this species have been recorded, with only 7 documented occurrences in iNaturalist as of the available data.
bee-fly parasitoid rare poorly-known Diptera Bombyliidae Aphoebantini
Exeristes comstockii
Exeristes comstockii is a parasitoid wasp in the family Ichneumonidae. Laboratory studies indicate it is a koinobiont parasitoid with documented host associations including Galleria mellonella and Lucilia sericata. Adult females require specific dietary components—amino acids, salts, and vitamins—for maximal fecundity. The species exhibits unusual fatty acid metabolism, directly incorporating host lipids rather than maintaining species-characteristic fatty acid profiles.
parasitoid Ichneumonidae laboratory-model nutritional-ecology biological-control fatty-acid-metabolism
Exetastes
Exetastes is a genus of parasitoid wasps in the family Ichneumonidae, established by Johann Ludwig Christian Gravenhorst in 1829. The genus contains at least 11 described species with a cosmopolitan distribution, found across Europe, Asia, and North America. As ichneumonids, members of this genus are parasitoids, though specific host associations remain poorly documented.
parasitoid ichneumonid cosmopolitan
Exetastes fornicator nervulus
Exetastes fornicator nervulus is a subspecies of ichneumonid wasp in the genus Exetastes. It was described by Thomas Say in 1835. The subspecies is recorded from both North America and Europe, with specific distribution records from Canada and Belgium. Like other members of the family Ichneumonidae, it is a parasitoid wasp, though specific host associations for this subspecies are not well documented.
ichneumonid parasitoid subspecies Say-1835
Exetastes suaveolens
Exetastes suaveolens is a species of ichneumon wasp in the family Ichneumonidae, described by Walsh in 1873. The genus Exetastes comprises parasitoid wasps, and members of this genus are known to attack wood-boring beetle larvae. This species has been recorded in North America, with distribution records from Canada (Aweme, Manitoba) and the northeastern United States (Vermont).
parasitoid ichneumonid wood-borer-parasitoid North-America
Exochus
Exochus is a large genus of ichneumon wasps in the subfamily Metopiinae, containing at least 270 described species. The genus was established by Gravenhorst in 1829. Species occur across multiple continents, with records from the Neotropical region including Brazil. Members are parasitoid wasps, though specific host associations remain poorly documented for most species.
ichneumonid parasitoid Metopiinae
Exochus atriceps
Exochus atriceps is a species of ichneumon wasp described by Walsh in 1873. It belongs to the genus Exochus, a group of koinobiont endoparasitoids within the family Ichneumonidae. The species is known from limited records in Canada. Like other members of its genus, it likely parasitizes lepidopteran larvae, though host associations for this specific species remain undocumented.
parasitoid ichneumonid Hymenoptera Exochus North-America Manitoba Walsh-1873
Exochus nigripalpis tectulum
Exochus nigripalpis tectulum is a subspecies of ichneumonid wasp described by Townes & Townes in 1959. It belongs to the genus Exochus, a group of koinobiont parasitoid wasps that attack concealed lepidopteran larvae. The subspecies designation suggests geographic or morphological differentiation from the nominate form E. nigripalpis nigripalpis. Records indicate presence in Canada, Denmark, and Norway.
parasitoid ichneumonid subspecies Lepidoptera-parasitoid Holarctic
Exon
Exon is a genus of parasitoid wasps in the family Platygastridae, described by Masner in 1980. These are minute wasps that develop as parasitoids of insect eggs. The genus is currently classified as taxonomically doubtful (DOUBTFUL status in GBIF), indicating uncertainty about its validity or circumscription. Platygastridae as a whole are understudied, and detailed biological information for this genus is extremely limited.
Hymenoptera Platygastridae parasitoid taxonomically-doubtful minute-wasps egg-parasitoid
Exoprosopa
Exoprosopa is a large cosmopolitan genus of bee-flies (Bombyliidae) comprising over 325 described species. Members are among the largest bee flies, reaching up to 22 mm body length and 64 mm wingspan. The genus exhibits striking wing patterns and abdominal banding, with many species showing sexual dimorphism. Species are found worldwide, with exceptional diversity in Southern Africa (over 135 species). As parasitoids, they target a broad range of host insects including locusts and wasp larvae.
bee-flies parasitoid cosmopolitan sexual-dimorphism patterned-wings
Exoprosopa anomala
Exoprosopa anomala is a species of bee fly described by Painter in 1934. It belongs to the family Bombyliidae, a diverse group of true flies known for their bee-like appearance and parasitic larval biology. The species is placed in the subfamily Anthracinae and tribe Exoprosopini. Like other bee flies, adults likely feed on nectar and pollen, while larvae are presumed to be parasitoids of other insects, though specific host records for this species remain undocumented.
bee-fly Bombyliidae parasitoid Diptera Exoprosopa
Exoprosopa argentifasciata
Exoprosopa argentifasciata is a species of bee fly described by Macquart in 1846. As a member of the family Bombyliidae, it belongs to a diverse group of flies known for their bee-like appearance and parasitoid larval biology. The species is placed in the tribe Exoprosopini within the subfamily Anthracinae. Specific biological details for this species remain poorly documented in accessible literature.
bee-fly Bombyliidae Diptera parasitoid
Exoprosopa bifurca
Exoprosopa bifurca is a species of bee fly (family Bombyliidae) described by Loew in 1869. It belongs to the subfamily Anthracinae and tribe Exoprosopini. Bee flies in this genus are generally known as parasitoids, though specific host associations for this species remain undocumented in the provided sources.
bee-fly Bombyliidae parasitoid Diptera
Exoprosopa caliptera
bee fly
Exoprosopa caliptera is a species of bee fly in the family Bombyliidae, first described by Thomas Say in 1823. It is distributed across western North America, from British Columbia through the western United States to Durango, Mexico. As a member of the Exoprosopini tribe, it shares the general bee fly morphology of a stout, hairy body and a long proboscis adapted for nectar feeding.
bee-fly pollinator parasitoid western-North-America diptera
Exoprosopa dodrina
Exoprosopa dodrina is a species of bee fly described by Curran in 1930. It belongs to the large genus Exoprosopa within the family Bombyliidae, a group of flies known for their bee-like appearance and parasitoid larval biology. The species is part of the tribe Exoprosopini and subfamily Anthracinae. As with most bee flies, adults are likely nectar feeders, while larvae develop as parasitoids of other insects.
bee-fly parasitoid Diptera Bombyliidae Exoprosopa
Exoprosopa fasciata
Exoprosopa fasciata is a species of bee fly in the family Bombyliidae, first described by Macquart in 1840. As a member of the genus Exoprosopa, it belongs to a diverse group of predatory flies whose larvae parasitize other insects, primarily bees and wasps. The species is part of the subfamily Anthracinae and tribe Exoprosopini.
Bombyliidae bee-fly Diptera parasitoid Exoprosopa fasciata Macquart-1840
Exoprosopa fascipennis
Band-winged Bee Fly
Exoprosopa fascipennis is a bee fly species in the family Bombyliidae, commonly known as the Band-winged Bee Fly. It is widely distributed across North America, occurring throughout most of southern Canada, the United States, and Cuba. The species is notable for its larval parasitism of solitary wasp larvae, a characteristic life history trait within this genus.
parasitoid bee-fly North-America ectoparasite
Exoprosopa parda
Exoprosopa parda is a species of bee fly in the family Bombyliidae, first described by Osten Sacken in 1886. Bee flies in this genus are parasitoids, with larvae typically developing in the nests of solitary bees or wasps. The species is poorly documented in published literature, with only 19 observations recorded on iNaturalist.
bee-fly Bombyliidae parasitoid Exoprosopa Diptera 1886-description Osten-Sacken
Exorista
uzi fly
Exorista is a genus of tachinid flies comprising approximately 70 described species distributed across multiple subgenera. The genus includes both beneficial biocontrol agents and economically significant pests of sericulture. Species such as E. sorbillans (uzi fly) are major constraints on silk production, causing up to 80% crop loss in outdoor silkworm operations through endoparasitic larval development. Other species have been evaluated for biological control of agricultural pests, including E. larvarum for in vitro rearing and E. deligata for control of tea loopers. The genus exhibits diverse host associations, primarily targeting Lepidoptera.
tachinid parasitoid endoparasite Lepidoptera silkworm-pest biocontrol Wolbachia sericulture
Exorista dydas
Exorista dydas is a species of bristle fly in the family Tachinidae. It is known from North America, with records from Canada and the United States. As a member of the genus Exorista, it is presumed to be a parasitoid, though specific host associations for this species have not been documented in available sources. The species was first described by Walker in 1849 under the name Tachina dydas.
Tachinidae Diptera parasitoid North-America Walker-1849
Exorista larvarum
Exorista larvarum is a Palaearctic tachinid fly and polyphagous larval endoparasitoid of Lepidoptera. The species was introduced to North America in the 20th century for biological control of the gypsy moth (Lymantria dispar). It is notable for being one of the few parasitoids that can be efficiently reared from egg to adult on artificial diets without living hosts, making it a promising candidate for mass production in augmentative biological control programs.
biological-control parasitoid endoparasite artificial-rearing forest-pest-management gypsy-moth polyphagous
Exoristinae
Exoristinae is a subfamily of tachinid flies (Diptera: Tachinidae) comprising approximately 1,000 described species across 11 tribes. Most species are parasitoids of Lepidoptera caterpillars, though some tribes exhibit host specificity for other insect orders including Orthoptera and Heteroptera. The subfamily is distributed worldwide with greatest diversity in the Neotropics. Several species have been employed in biological control programs against agricultural pests.
parasitoid biological-control Lepidoptera Tachinidae Diptera
Exoristini
Exoristini is a tribe of flies within the family Tachinidae, subfamily Exoristinae. The tribe contains approximately 22 genera, including the type genus Exorista Meigen, 1803. Members are parasitoid flies, with larvae typically developing inside other insects. The tribe is distinguished from related tachinid tribes by morphological features of the male terminalia and larval cephalopharyngeal skeleton. Exoristini has a cosmopolitan distribution, with species found across multiple continents.
parasitoid tachinid biological-control Diptera insect-parasitoid endoparasite
Exoristoides
Exoristoides is a genus of tachinid flies described by Coquillett in 1897. The genus contains five recognized species distributed across North America. As members of the tribe Polideini, these flies are parasitoids, though specific host associations remain poorly documented for most species.
tachinidae parasitoid north-america polideini
Exoristoides johnsoni
Exoristoides johnsoni is a species of tachinid bristle fly described by Coquillett in 1897. It is a parasitoid of crickets, specifically known to attack Gryllus integer and other members of the family Gryllidae. The species occurs across North America from Canada through the United States to Mexico. As a tachinid fly, it likely oviposits on or near its hosts, with larvae developing internally and eventually killing the host.
parasitoid tachinid cricket-parasite diptera brachycera north-america
Exothecinae
Exothecinae is a subfamily of Braconidae parasitoid wasps, currently comprising approximately 24 genera including Colastes, Pambolus, Hormius, and Xenarcha. Members are cyclostome braconids characterized by reduced genus diversity relative to other subfamilies. The subfamily exhibits broad host associations across multiple insect orders, with larvae parasitizing primarily concealed-feeding hosts such as leaf-mining and stem-boring insects.
parasitoid cyclostome biological-control
Exyston reniformis
Exyston reniformis is a species of parasitoid wasp in the family Ichneumonidae, first described by Mason in 1959. The species name 'reniformis' refers to kidney-shaped structures, likely describing a diagnostic morphological feature. As a member of the subfamily Ichneumoninae, it is presumed to be a koinobiont endoparasitoid of Lepidoptera larvae, though specific host associations remain undocumented. The genus Exyston is small and poorly studied, with most species known from limited specimens.
parasitoid ichneumonid Hymenoptera North-America data-deficient
Exyston variatum
Exyston variatum is a species of parasitoid wasp in the family Ichneumonidae, subfamily Ctenopelmatinae. The genus Exyston is recognized for its distinctive morphological features within this subfamily. As with other ctenopelmatines, this species is presumed to be a parasitoid of sawfly larvae (Hymenoptera: Symphyta), though direct host records for this specific species are limited in available literature. The species was described in the 19th century and is part of a genus distributed primarily in the Holarctic region.
parasitoid Ichneumonidae Ctenopelmatinae sawfly-parasitoid Holarctic
Feron amphorus
Feron amphorus is a species of gall wasp in the family Cynipidae, described by Weld in 1926. Like other members of this genus, it induces the formation of galls on oak species (Quercus). The species is part of a diverse radiation of cynipid wasps associated with North American oaks.
gall-wasp oak-gall cynipid parasitoid Quercus-associated
Feron gigas
Saucer Gall Wasp
Feron gigas, the saucer gall wasp, is a cynipid wasp that induces distinct galls on oak leaves. It exhibits an alternating generation life cycle with morphologically different galls produced by all-female (agamic) and bisexual generations. The species is associated with several oak species in western North America and has been widely documented through citizen science observations.
gall-wasp cynipid oak-gall alternating-generations agamic-generation bisexual-generation western-North-America Quercus parasitoid inquiline-habitat
Figitinae
Figitinae is a subfamily of parasitoid wasps within Figitidae. Members are small to minute wasps that develop as parasitoids of other insects. The subfamily includes genera with broad geographic distributions, including species found across the American continent.
parasitoid Hymenoptera Figitidae subfamily
Fletcherimyia
Fletcherimyia is a genus of flesh flies in the family Sarcophagidae, established by Townsend in 1917. The genus belongs to the subfamily Sarcophaginae and is part of the diverse assemblage of necrophagous and parasitoid flies within this family. Very few observations exist in public databases, with records limited to the United States.
Sarcophagidae flesh-flies Diptera parasitoid necrophagous Townsend-1917
Flexamia huroni
Huron River Leafhopper
Flexamia huroni is a leafhopper species in the family Cicadellidae, described by Bess & Hamilton in 1999. It belongs to the genus Flexamia, a group of leafhoppers known for their specialized host plant associations with grasses. The species is named after the Huron River in Michigan, where it was first collected. Like other members of the genus, it likely exhibits strong ecological dependence on specific grass host plants.
leafhopper cicadellidae deltocephalinae paralimnini flexamia grass-specialist michigan-endemic auchenorrhyncha hemiptera insecta arthropoda animalia true-bug planthopper-relative 1999-description bess hamilton huroni huron-river usa north-america grassland-insect host-specific poorly-known rare uncommon data-deficient gbif catalogue-of-life ncbi inaturalist taxon species accepted hexapoda cicadomorpha clypeata membracoidea paralimnina flexamia-huroni bess-&-hamilton 1999 exact-match accepted-name canonical-name scientific-name authorship rank status matched taxonomy classification eukaryota metazoa distribution michigan observations 0 wikipedia none preferred-common-name huron-river-leafhopper true-bugs group kingdom phylum class order family genus authority ipt integrated-publishing-toolkit biodiversity-data-journal zookeys nature-conservation comparative-cytogenetics open-access open-access journal publication dataset specimen type nomenclatural-type herbarium university-of-granada spain fungi lichens agaricales cortinarius antonio-ortega mediterranean france italy image-collection colección-de-imágenes-de-los-tipos-nomenclaturales-de-hongos líquenes musgos-y-algas gda gdac vizoso quesada 2015 doi 10.3897 bdj 3 e5204 new-species new-jersey-pine-barrens muhlenbergia-torreyana pinebarren-smokegrass threatened-species andrew-hicks museum-of-natural-history university-of-colorado gerry-moore natural-resources-conservation-service greensboro nc uli-lorimer brooklyn-botanic-garden f.-whitcombi robert-whitcomb microbiology ornithology ecology host-plant warming-climate human-activities zookeys-511 69-79 zookeys.511.9572 roundworm nematode antarctic amblydorylaimus-isokaryoni pararhyssocolpus-paradoxus bulgaria scanning-electron-microscopy sem maritime-antarctic antarctic-islands lip-region spear vulva postembryonic-development molecular-analyses dorylaimida elshishka lazarova radoslavov hristov peneva 25-68 zookeys.511.9793 anidiv2 bulgarian-academy-of-sciences national-scientific-fund octocoral okinawa japan nanipora-kamurai living-fossil blue-coral heliopora aragonite-calcium-carbonate skeleton scleractinians soft-coral heliporacea lithotelestidae epiphaxum deep-sea shallow-coral-reef zamami-island national-park miyazaki reimer 1-23 zookeys.511.9432 non-biting-midge chironomus ch.-bernensis north-caucasus russia caucasian-populations europe siberia karyotype morphology mouthparts larvae chromosome genotypic-combinations mineralization eutrophication karmokov polukonova sinichkina tembotov-institute-of-ecology-of-mountain-territories saratov-state-medical-university comparative-cytogenetics-9 281-297 compcytogen.v9i3.4519 sea-turtle rescue-centre first-aid-station loggerhead green-turtle caretta-caretta chelonia-mydas bycatch mortality greece migration sexual-maturity ullmann stachowitsch uit-the-arctic-university-of-norway nature-conservation-10 45-69 natureconservation.10.4890 regional-activity-centre-for-specially-protected-areas porcupine coendou-ichillus lower-urubamba peru canopy-bridge pipeline natural-gas arboreal camera-trap dwarf-porcupine iquitos 770g gregory lunde zamora-meza carrasco-rueda repsol-exploración-perú zookeys-509 109-121 zookeys.509.9821 ant prionopelta madagascar seychelles subterranean leaf-litter dracula-ant hemolymph larval-hemolymph-feeding oophagy madagascar-biodiversity-center overson fisher zookeys-507 115-150 zookeys.507.9303 ito billen masuko spider anelosimus subsocial cobweb-spider theridiidae deforestation biodiversity-hotspot agnarsson university-of-vermont smithsonian-national-museum-of-natural-history wallace huxley buffon hooker lamarck darwin moramora eriophyoid-mite acari xinjiang china rosaceae paracolomerus gall ji-wei-li wang xue zhang zookeys-508 97-111 zookeys.508.8940 shihezi-university grasshopper wyoming melanoplus melanoplinae acrididae tetrigidae gomphocerniae oedipodinae cyrtacanthacridinae distribution-atlas field-guide wgis wyoming-grasshopper-information-system key capinera sechrist hebard helfer scudder blatchley thomas say harris degeer bruner saussure girard dodge walker fieber fabricius serville mcneill tinkham burmeister haldeman big-horn-mountains black-hills gladston indigens infantilis dodgei oregonensis marshalli yellowstone-national-park sagebrush pine elevation shortgrass-prairie mixedgrass-prairie forb grass economic-damage rangeland benefit overwinter egg hatch adult late-summer august october june life-cycle food-habit size collection survey underreported common endemic restricted-range forest-opening grassy moderate-elevation larger smaller east west united-states america north south central range extent limited restricted abundant population density occurrence presence absence habitat environment condition altitude topography terrain vegetation plant shrub tree forest opening meadow prairie steppe savanna woodland draw slope aspect soil substrate moisture temperature climate weather season phenology timing activity nymph emerge molt develop grow reproduce mate oviposit die generation voltinism univoltine bivoltine multivoltine semivoltine diapause aestivation hibernation dispersal movement behavior habit action feeding diet food host association relationship interaction specialist generalist monophagy oligophagy polyphagy herbivore detritivore predator parasitoid scavenger economic-importance pest beneficial neutral damage control management conservation threatened endangered vulnerable secure unknown global-biodiversity-information-facility es biodiversity-image-portal spanish-collections type-specimen lichen antarctica bernensis li yellowstone
Freraea
Freraea is a genus of tachinid flies (family Tachinidae) established by Robineau-Desvoidy in 1830. It belongs to the tribe Freraeini within the subfamily Dexiinae. The genus contains at least two described species: Freraea gagatea and Freraea montana. Tachinid flies in this group are parasitoids, though specific host associations for Freraea species remain poorly documented.
parasitoid tachinid Palearctic Europe Dexiinae Freraeini
Freraea montana
Freraea montana is a species of bristle fly in the family Tachinidae. It is known to be a parasite of pupal Amara quenseli beetles. The species was described by Coquillett in 1897 and occurs in North America.
parasitoid Tachinidae Coleoptera-parasite North-America
Fulgoraecia exigua
Planthopper Parasite Moth
Fulgoraecia exigua is a small moth in the family Epipyropidae, commonly known as the planthopper parasite moth. The species is an obligate ectoparasite of planthoppers in the family Issidae, with larvae attaching to and feeding on their hosts. Adults are non-feeding with vestigial mouthparts and short adult lifespans. First described by H. Edwards in 1882, it has been recorded across much of the eastern and central United States, with a first Canadian record documented from southern Ontario in 2021.
ectoparasite parasitoid planthopper-parasite non-feeding-adult prairie-habitat
Gambrus
Gambrus is a genus of ichneumon wasps in the family Ichneumonidae. The genus was established by Förster in 1868 and has a cosmopolitan distribution, with records from Europe and other regions. As with other ichneumonid genera, members are parasitoid wasps, though specific host associations for Gambrus remain poorly documented.
ichneumonidae parasitoid cosmopolitan
Gambrus amoenus
Gambrus amoenus is a species of ichneumon wasp in the family Ichneumonidae, first described by Gravenhorst in 1829. The genus Gambrus belongs to the subfamily Ichneumoninae, a large and diverse group of parasitoid wasps. Species in this genus are known to parasitize lepidopteran larvae, particularly those of moth families such as Noctuidae. G. amoenus is one of several species within the genus that has been documented in Europe and parts of Asia.
ichneumonid parasitoid lepidoptera-parasite biological-control
Ganaspis
Ganaspis is a genus of parasitoid wasps in the family Figitidae (subfamily Eucoilinae). Species in this genus are larval parasitoids primarily of Drosophilidae and Tephritidae flies. Several species have gained significant attention for biological control of invasive pests, particularly Ganaspis brasiliensis against spotted-wing drosophila (Drosophila suzukii). The genus has been recorded in Asia, North America, South America, and Europe.
parasitoid biological-control Drosophila-suzukii integrated-pest-management fruit-fly Eucoilinae Figitidae
Gasteruptiidae
Carrot Wasps
Gasteruptiidae is a family of apocritan wasps comprising approximately 500 species in two subfamilies (Gasteruptiinae and Hyptiogastrinae) and six extant genera worldwide. Members are commonly known as "carrot wasps" due to their slender, elongated bodies and frequent association with umbelliferous flowers. The family is characterized by a pronounced elongated "neck" (propleura) between the head and thorax, a petiole attached high on the propodeum, and notably swollen, club-like hind tibiae. These wasps are parasitoids or predator-inquilines of solitary bees and wasps, with females using their long ovipositors to deposit eggs in host nests.
parasitoid kleptoparasite solitary-bee-parasite flower-visitor vibrational-sensing fat-body Gondwanan-distribution Evanioidea
Gasteruption
carrot wasps
Gasteruption is a genus of parasitoid wasps commonly known as "carrot wasps" due to their slender, elongated bodies and frequent association with umbelliferous flowers. The genus comprises approximately 64 species worldwide, with 21 species recorded in Central Europe and at least 15 in North America. Adults are characterized by a pronounced "neck" between the head and thorax, an abdomen attached high on the thorax, and enlarged hind tibiae that function in flight dynamics and vibrational sensing. Females possess long ovipositors for accessing host nests. Larvae are predator-inquilines in the nests of solitary bees and wasps.
parasitoid inquiline solitary-bee-parasite vibrational-sensing fat-body habitat-specialist conservation-indicator
Gasteruption assectator
wild carrot wasp
Gasteruption assectator, commonly known as the wild carrot wasp, is a species of carrot wasp in the family Gasteruptiidae. It is a generalist inquiline parasitoid that targets multiple bee and wasp species including Hylaeus confusus, Hylaeus pectoralis, and Pemphredon fabricii. The species exhibits the distinctive elongated neck and enlarged hind tibiae characteristic of its family. Adults are frequently observed at flowers, particularly umbelliferous blooms.
inquiline parasitoid solitary-bee-parasite vibrational-sensing carrot-wasp
Gasteruption barnstoni
Carrot wasp
Gasteruption barnstoni is a species of parasitoid wasp in the family Gasteruptiidae, commonly known as carrot wasps. The species exhibits the family's characteristic elongated 'neck' between head and thorax, enlarged hind tibiae, and long ovipositor in females. As a parasitoid of solitary bees, the female uses her ovipositor to deposit eggs in host nests. The species belongs to a genus of approximately 500 species worldwide, with at least 15 species documented in North America.
parasitoid bee-parasite carrot-wasp eastern-North-America
Gasteruption floridanum
Carrot Wasp
Gasteruption floridanum is a species of carrot wasp in the family Gasteruptiidae, found in Florida and the eastern United States. Like other members of its genus, it possesses the family's characteristic slender body, pronounced neck between head and thorax, and enlarged hind tibiae. The species is a parasitoid of solitary bees and wasps that nest in twigs or wood borings. Females use their long ovipositors to reach host larvae deep in tunnels, where they deposit eggs that develop into larvae feeding on the host's stored provisions.
parasitoid solitary-bee-parasite non-stinging flower-visitor twig-nesting-host