Invasive
Guides
Creugas gulosus
Creugas gulosus is a species of true spider in the family Corinnidae, first described by Thorell in 1878. Native to South America, it has established introduced populations across Africa, Myanmar, Australia, and Pacific islands. The species belongs to a genus of ground-dwelling spiders that are part of the diverse corinnid fauna.
Crossopriza lyoni
Tailed Cellar Spider, Tailed Daddy Longlegs Spider, Box Spider
Crossopriza lyoni is a cosmopolitan cellar spider (Pholcidae) characterized by extremely long, fragile legs and a distinctive box-shaped abdomen. Females reach 3–7 mm in body length, males 2.5–6 mm, with leg spans up to 6 cm. The species is synanthropic, thriving in human dwellings worldwide, particularly in tropical to subtropical regions. It constructs extensive, irregular cobweb-like webs and is often regarded as both a household pest and beneficial predator of mosquitoes and other arthropods. Native range unknown; now pantropical through human-mediated introduction.
Cryptorhynchus
hidden snout weevils
Cryptorhynchus is a genus of hidden snout weevils in the family Curculionidae, containing at least 480 described species. The genus name derives from Greek roots meaning 'hidden' and 'snout,' referring to the concealed rostrum characteristic of these beetles. The genus includes economically significant species such as Cryptorhynchus lapathi, the poplar-and-willow borer, which is a major pest of Salix and Populus species in North America and elsewhere.
Cryptorhynchus lapathi
Poplar-and-Willow Borer, Poplar and Willow Borer, Willow Borer, Osier Weevil, Willow Weevil
Cryptorhynchus lapathi is a Eurasian-origin weevil (Curculionidae) introduced to North America, now established as a significant pest of willow (Salix) and poplar (Populus) species. Larvae bore into stems, creating infection courts for pathogens and causing structural weakening that leads to stem breakage. The species has expanded its range considerably in British Columbia since its first detection in 1923, with prevalence linked to thermal regimes—areas with more months above 10°C show higher attack rates. Climate modeling predicts continued northward range expansion and increased suitable habitat area through 2100.
Ctenolepisma longicaudatum
gray silverfish, long-tailed silverfish, paper silverfish
Ctenolepisma longicaudatum is a synanthropic silverfish species first described from South Africa in 1905, now distributed worldwide in human dwellings. It has become a significant indoor pest in Europe since the early 2000s, particularly in newly constructed buildings with stable climates. The species damages cellulose-based materials including books, documents, and textiles, and can survive extended periods without food.
Ctenophora
comb jellies, sea gooseberries, sea walnuts, Venus's girdles
Ctenophora is a phylum of marine invertebrates distinguished by eight rows of fused cilia used for locomotion, called comb rows. They are the largest animals to swim using cilia. Unlike cnidarians, they lack stinging cells and instead capture prey using colloblasts—sticky adhesive cells. The phylum contains approximately 150–186 recognized species exhibiting diverse body plans, from egg-shaped cydippids with retractable tentacles to flattened benthic platyctenids and large-mouthed beroids. Ctenophores possess a decentralized nerve net without a brain, and their phylogenetic position as either the earliest or second-earliest branching animal lineage remains actively debated.
Cydia pomonella
Codling Moth
Cydia pomonella is a tortricid moth and major global pest of apple and pear orchards. The caterpillar, commonly known as the 'worm in the apple,' bores into fruit and prevents normal development, causing premature ripening and crop loss. Adults are attracted to pheromone traps with limited plume reach, requiring multiple traps for reliable monitoring. The species has been detected via environmental DNA in yard waste shipments, highlighting its potential for human-mediated dispersal.
Desidae
intertidal spiders, desid spiders
Desidae is a family of spiders with remarkable ecological diversity, ranging from truly marine intertidal species to inland web-builders. The family was historically defined by intertidal members of the genus Desis, which construct air-filled silk retreats in barnacle shells or kelp holdfasts to survive tidal submersion. Modern circumscription includes inland genera such as Badumna and Phryganoporus. The family contains 63 genera and over 320 species, with the majority native to Australia, New Zealand, and New Caledonia. Some species have become invasive synanthropes, establishing populations in North and South America, Europe, and Asia.
Dieuches armatipes
Dieuches armatipes is a seed bug in the family Rhyparochromidae. Native to Africa, it has established populations in the Western Hemisphere including the Caribbean and Florida. The species was first documented in the Western Hemisphere from Grand Cayman and St. Kitts in 1993, with subsequent records from intercepted specimens in the Dominican Republic and Jamaica. It has since been confirmed on the Florida mainland and in Belgium.
Dinoderus minutus
bamboo borer, bamboo powderpost beetle
Dinoderus minutus is a small wood-boring beetle in the family Bostrichidae, commonly known as the bamboo borer or bamboo powderpost beetle. Native to Asia, it has become a globally distributed pest of bamboo products through international trade. The species is attracted to the internal starch content of bamboo and is considered one of the most damaging pests of the bamboo industry, capable of causing significant economic losses in storage and finished products. It has established populations across multiple continents including Europe, Africa, North America, South America, and Oceania.
Drepanaphis acerifoliae
painted maple aphid
Drepanaphis acerifoliae, commonly known as the painted maple aphid, is a species of aphid in the family Aphididae. It is one of 16 species in the genus Drepanaphis, all native to North America. This species has the broadest geographic range among its congeners and is the only member of the genus known to have been introduced to Europe, where it has established populations in the Mediterranean region and beyond. Climate modeling suggests potential for further northward expansion in both North America and Europe over the coming decades.
Dryadaula terpsichorella
dancing moth, Hawaiian Dancing Moth
Dryadaula terpsichorella is a small moth known for its distinctive dancelike movements upon landing. Native to south-eastern Polynesia, Samoa, and Fiji, it has established populations in Hawaii, Florida, and California. The species was first described by August Busck in 1910. It is commonly called the "dancing moth" or "Hawaiian Dancing Moth" due to its characteristic behavioral display.
Echidnophaga gallinacea
hen flea, sticktight flea, stickfast flea
Echidnophaga gallinacea is a small, dark brown flea that parasitizes a broad range of bird and mammal hosts including poultry, dogs, cats, and occasionally humans. Adult females embed permanently into host skin—typically on bare areas of the head and face—using a serrated mouthpart to feed on blood, while males remain mobile. Heavy infestations cause ulceration, dermatitis, anemia, and can be fatal to young or small hosts. The species has spread globally through poultry farming and is now established across tropical, subtropical, and temperate regions.
fleaectoparasitepoultry-pestvectorinvasivecosmopolitanintegrated-pest-managementcage-free-poultrybackyard-chickenshematophagypermanent-attachmentsessile-femaleresilinjumpingctenidia-absentanaemiadermatitisulcerationRickettsiaBartonellaplaguediatomaceous-earthmaldisonpesticide-resistanceorganic-farminganimal-welfareeconomic-entomologyveterinary-entomologyhost-generalisturban-wildlife-interfacesynanthropicEchinothrips americanus
Poinsettia thrips, Impatiens thrips, American thrips
Echinothrips americanus is a small thrips species in the family Thripidae, native to eastern North America and first described from Florida in 1913. It has become an invasive pest of global concern, spreading to Europe, Asia, Australia, and other regions since the 1990s. The species is highly polyphagous, feeding on foliage of at least 48 plant families, with particular preference for Araceae and Balsaminaceae. It is primarily a greenhouse and ornamental pest, causing chlorotic damage to leaves through piercing-sucking feeding.
Ectobius
wood cockroaches, field cockroaches
Ectobius is a genus of small, cool-adapted cockroaches in the family Ectobiidae. Adults measure 6–12 mm in length with brown to yellowish coloration and pale margins. The genus has a complex biogeographic history: fossil evidence from the 49-million-year-old Green River Formation in Colorado indicates Ectobius originated in North America, despite its long absence from the continent until recent reintroductions. Species are primarily distributed across Europe, Africa, the eastern Palearctic, and the Near East. Several species have been introduced to northeastern North America within the last 65 years, where Ectobius lapponicus has become synanthropic.
Eriosoma lanigerum
woolly apple aphid, woolly aphid, American blight
Eriosoma lanigerum is a small phloem-feeding aphid native to North America that has become a major worldwide pest of apple cultivation. Adults are concealed beneath dense white, cotton-like wax secretions produced from specialized abdominal glands, giving the species its common name. The aphid induces gall formation on both aerial and root tissues of host plants, disrupting vascular transport and potentially killing trees. Populations are primarily parthenogenetic in most regions, though sexual reproduction occurs where American elm (Ulmus americana) is present.
Euphyllura olivina
olive psyllid
Euphyllura olivina is a psyllid species in the family Liviidae that feeds exclusively on olive (Olea europaea). Native to the Mediterranean region, it has become invasive in California where it threatens olive production. The species develops two generations annually, with spring and autumn reproductive activity and summer reproductive diapause induced by high temperatures. Nymphal infestations concentrate near fruits and on shaded, lower canopy portions. Temperature sensitivity shapes its distribution and seasonal activity patterns.
Eupteryx
Eupteryx is a genus of leafhoppers in the family Cicadellidae, subfamily Typhlocybinae. Species within this genus are small, plant-feeding insects with documented associations primarily with plants in the mint family (Lamiaceae) and stinging nettles (Urtica dioica). Some species, notably E. decemnotata, have undergone rapid range expansion in recent decades. The genus exhibits host-specific oviposition behavior and color polymorphism in nymphal stages.
Eurhinus magnificus
Jewel Weevil, Iridescent Weevil
Eurhinus magnificus, commonly known as the jewel weevil, is a brilliantly colored weevil native to Central America. Adults are small (5–6 mm long, 3–4 mm wide) with striking metallic blue-green bodies accented by metallic red-copper on the humeri, elytra apex, pronotum, rostrum, and legs. The species was first described by Leonard Gyllenhaal in 1836 and has been introduced to southern Florida, where it was first detected in 2002. It is a gall-forming herbivore whose larvae develop within stem galls on its primary host plant, Cissus verticillata.
Euura lipovskyi
Azalea Sawfly
Euura lipovskyi, commonly known as the Azalea Sawfly, is a sawfly species in the family Tenthredinidae. The species was first described from the eastern United States and has undergone recent taxonomic revision, with the combination Euura lipovskyi newly proposed. Larvae are specialized feeders on Rhododendron species in section Pentanthera. The species has shown notable range expansion in recent decades, now occurring in the Pacific Northwest, Europe, and Canada.
Exapion ulicis
Gorse Seed Weevil
Exapion ulicis is a small seed-feeding weevil specialized on gorse (Ulex europaeus). Adults are light gray with a prominent snout roughly half the body length. The species is native to western Europe and has been introduced to New Zealand, California, Hawaii, and other regions as a biological control agent targeting invasive gorse populations. Larval feeding destroys seeds within pods, reducing plant spread, while adult feeding on stems and spines causes minor damage.
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-collectively-retentive-Pareto-optimal-setminimal-individually-retentive-Pareto-optimal-setminimal-Nash-retentive-Pareto-optimal-setminimal-individually-rational-retentive-Pareto-optimal-setminimal-weakly-stable-Pareto-optimal-setminimal-strongly-stable-Pareto-optimal-setminimal-collectively-stable-Pareto-optimal-setminimal-individually-stable-Pareto-optimal-setminimal-Nash-stable-Pareto-optimal-setminimal-individually-rational-stable-Pareto-optimal-setminimal-weakly-Nash-stable-Pareto-optimal-setminimal-strongly-Nash-stable-Pareto-optimal-setminimal-collectively-Nash-stable-Pareto-optimal-setminimal-individually-Nash-stable-Pareto-optimal-setminimal-Nash-Nash-stable-Pareto-optimal-setminimal-individually-rational-Nash-stable-Pareto-optimal-setminimal-weakly-individually-rational-Pareto-optimal-setminimal-strongly-individually-rational-Pareto-optimal-setminimal-collectively-individually-rational-Pareto-optimal-setminimal-individually-individually-rational-Pareto-optimal-setminimal-Nash-individually-rational-Pareto-optimal-setminimal-individually-rational-individually-rational-Pareto-optimal-setminimal-weakly-Pareto-optimal-Nash-stable-setminimal-strongly-Pareto-optimal-Nash-stable-setminimal-collectively-Pareto-optimal-Nash-stable-setminimal-individually-Pareto-optimal-Nash-stable-setminimal-Nash-Pareto-optimal-Nash-stable-setminimal-individually-rational-Pareto-optimal-Nash-stable-setminimal-weakly-comprehensive-Pareto-optimal-Nash-stable-setminimal-strongly-comprehensive-Pareto-optimal-Nash-stable-setminimal-collectively-comprehensive-Pareto-optimal-Nash-stable-setminimal-individually-comprehensive-Pareto-optimal-Nash-stable-setminimal-Nash-comprehensive-Pareto-optimal-Nash-stable-setminimal-individually-rational-comprehensive-Pareto-optimal-Nash-stable-setminimal-weakly-extensive-Pareto-optimal-Nash-stable-setminimal-strongly-extensive-Pareto-optimal-Nash-stable-setminimal-collectively-extensive-Pareto-optimal-Nash-stable-setminimal-individually-extensive-Pareto-optimal-Nash-stable-setminimal-Nash-extensive-Pareto-optimal-Nash-stable-setminimal-individually-rational-extensive-Pareto-optimal-Nash-stable-setminimal-weakly-retentive-Pareto-optimal-Nash-stable-setminimal-strongly-retentive-Pareto-optimal-Nash-stable-setminimal-collectively-retentive-Pareto-optimal-Nash-stable-setminimal-individually-retentive-Pareto-optimal-Nash-stable-setminimal-Nash-retentive-Pareto-optimal-Nash-stable-setminimal-individually-rational-retentive-Pareto-optimal-Nash-stable-setminimal-weakly-stable-Pareto-optimal-Nash-stable-setminimal-strongly-stable-Pareto-optimal-Nash-stable-setminimal-collectively-stable-Pareto-optimal-Nash-stable-setminimal-individually-stable-Pareto-optimal-Nash-stable-setminimal-Nash-stable-Pareto-optimal-Nash-stable-setminimal-individually-rational-stable-Pareto-optimal-Nash-stable-setminimal-weakly-Nash-stable-Pareto-optimal-Nash-stable-setminimal-strongly-Nash-stable-Pareto-optimal-Nash-stable-setminimal-collectively-Nash-stable-Pareto-optimal-Nash-stable-setminimal-individually-Nash-stable-Pareto-optimal-Nash-stable-setminimal-Nash-Nash-stable-Pareto-optimal-Nash-stable-setminimal-individually-rational-Nash-stable-Pareto-optimal-Nash-stable-setminimal-weakly-individually-rational-Pareto-optimal-Nash-stable-setminimal-strongly-individually-rational-Pareto-optimal-Nash-stable-setminimal-collectively-individually-rational-Pareto-optimal-Nash-stable-setminimal-individually-individually-rational-Pareto-optimal-Nash-stable-setminimal-Nash-individually-rational-Pareto-optimal-Nash-stable-setminimal-individually-rational-individually-rational-Pareto-optimal-Nash-stable-setminimal-weakly-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-comprehensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-comprehensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-comprehensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-comprehensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-comprehensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-comprehensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-extensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-extensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-extensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-extensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-extensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-extensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setGabrius astutoides
Gabrius astutoides is a rove beetle in the family Staphylinidae, subfamily Staphylininae. Originally described from Europe in 1946, it has established populations in eastern North America through human introduction. The species belongs to the diverse genus Gabrius, which contains numerous similar-looking species requiring careful examination for identification.
Glycaspis
Glycaspis is a genus of plant-parasitic psyllids in the family Aphalaridae. Species in this genus are strongly associated with Eucalyptus host plants. The genus includes at least two described species: Glycaspis brimblecombei (red gum lerp psyllid), a significant invasive pest of eucalyptus plantations worldwide, and Glycaspis granulata. Many Glycaspis species exhibit high host specificity, often restricted to single Eucalyptus species or closely related host groups. The genus is native to Australia, where it shows patterns of short-range endemism linked to host plant distribution.
Glyphodes pyloalis
lesser mulberry snout moth, lesser mulberry pyralid, beautiful glyphodes moth, mulberry pyralid
Glyphodes pyloalis is a small crambid moth native to Asia that has expanded its range to North America, Africa, and Central Asia. It is a specialist pest of mulberry (Morus spp.), with larvae feeding on leaves and causing significant economic damage to sericulture and mulberry cultivation. The species has shown notable range expansion in recent decades, facilitated by climate warming and wind dispersal. Population dynamics are strongly influenced by temperature extremes, with cold winters causing mass mortality of overwintering larvae.
Grapholita delineana
Eurasian hemp moth, hemp moth, hemp borer
Grapholita delineana, commonly known as the Eurasian hemp moth or hemp borer, is a tortricid moth species in the family Tortricidae. It has emerged as a significant economic pest of hemp (Cannabis sativa) in the United States following the crop's legalization for commercial production. The species is native to Eurasia but has established populations in North America where it can cause substantial damage to hemp crops, particularly those grown for cannabidiol (CBD) production.
Gynaikothrips ficorum
Cuban laurel thrips, fig leaf-rolling thrips
Gynaikothrips ficorum is a tube-tailed thrips in the family Phlaeothripidae, commonly known as the Cuban laurel thrips or fig leaf-rolling thrips. It is a gall-forming pest of Ficus species, inducing characteristic curled and folded leaf galls where it feeds, reproduces, and shelters. The species has achieved near-cosmopolitan distribution through human transport of its host plants. It was first described by Paul Marchal in 1908 as Phloeothrips ficorum.
Hemigrapsus
Hemigrapsus is a genus of varunid crabs containing fifteen recognized species. The genus is naturally distributed almost exclusively in the Pacific Ocean, with one species endemic to the Atlantic coast of South America. Two species—H. sanguineus and H. takanoi—have been introduced from East Asia to the North Atlantic region, where they have established invasive populations along European coasts and the eastern United States. One species, H. estellinensis, is considered extinct; it was endemic to a hypersaline spring in the Texas Panhandle.
Heterobostrychus
horned powder post beetles
A genus of wood-boring beetles in the family Bostrichidae, comprising six described species. Members are known for infesting timber, processed wood products, and certain crop plants. The genus has significant economic importance due to damage caused to lumber, furniture, and agricultural commodities. Several species have been introduced beyond their native ranges through international trade in wood products.
Heterobostrychus aequalis
oriental wood borer, lesser auger beetle, oriental bostrichid
Heterobostrychus aequalis is a horned powder-post beetle (family Bostrichidae) and serious pest of seasoned hardwood timber. Native to tropical and subtropical Asia, it has been introduced to multiple continents including North America, Australia, Africa, and Oceania. The species attacks starch-rich sapwood of timber and wood products, often remaining undetected until emergence holes and frass appear. In Florida, it has been intercepted at ports and is now established in the wild. In Australia, its establishment status remains ambiguous despite multiple detections, with evidence suggesting at most a tenuous local population in northern Queensland rather than widespread establishment.
Homalodisca vitripennis
Glassy-winged Sharpshooter
Homalodisca vitripennis, commonly known as the glassy-winged sharpshooter, is a large leafhopper native to the southeastern United States and northeastern Mexico. It has become a major invasive pest in California, Hawaii, and the Pacific Islands, where it vectors Xylella fastidiosa, the bacterium causing Pierce's disease of grapevine—a lethal condition for vineyards. The species exhibits complex vibrational communication during mating, with males and females engaging in duets using species-specific signals. Populations in California's Central Valley have developed significant resistance to neonicotinoid insecticides, particularly imidacloprid, creating challenges for chemical management. Classical biological control using egg parasitoids from the genus Gonatocerus (family Mymaridae) has proven effective, with parasitism rates reaching 80-100% in some areas.
Hoplocampa testudinea
apple sawfly, European apple sawfly
Hoplocampa testudinea is a sawfly species in the family Tenthredinidae that is a significant pest of apple orchards. Native to Europe, it was accidentally introduced to North America in 1939 and has since become invasive across the northeastern United States and Canada. The larvae develop inside developing apple fruitlets, causing characteristic ribbon scars or fruit drop, while adults are active during apple bloom and oviposit into the calyx of flowers.
Hyalopterus
Mealy Plum-Reed Aphids
Hyalopterus is a genus of aphids in the family Aphididae, commonly known as mealy plum-reed aphids. Species in this genus are cosmopolitan pests of stone fruit trees in the genus Prunus, causing damage through direct feeding and virus transmission. The genus has undergone recent taxonomic revision, with molecular and morphometric analyses supporting at least three distinct species: H. pruni, H. amygdali, and H. persikonus (the latter described in 2006). Species boundaries are strongly correlated with host-plant associations.
Hyblaea puera
teak defoliator
A small moth (wingspan 3–4 cm) and major defoliator of teak and mangrove ecosystems across tropical Asia. Adults have greyish-brown forewings that conceal brightly colored black and orange-yellow hindwings at rest. The species undergoes dramatic population outbreaks, with densities shifting from endemic levels to epidemic proportions seasonally. It has expanded beyond its native South and Southeast Asian range to Central America, Africa, and Australia, and has recently emerged as a serious invasive pest of mangroves in India and China.
Hyblaeoidea
Teak Moths
Hyblaeoidea is a small superfamily of moths in the order Lepidoptera, containing a single family Hyblaeidae with two genera (Hyblaea and Erythrochrus) and approximately 20 species. The superfamily has an uncertain phylogenetic position within the group Obtectomera, sometimes placed near Pyraloidea. Members are distributed in tropical and subtropical regions worldwide. The genus Hyblaea includes significant forest pests, notably Hyblaea puera, the teak defoliator.
Hypoponera punctatissima
Roger's ant
Hypoponera punctatissima, commonly known as Roger's ant, is a small ponerine ant native to Africa that has become a globally distributed tramp species. The species is strongly associated with human-modified environments, particularly heated buildings, greenhouses, and animal waste. Colonies are small, typically containing approximately 200 workers, and are often polydomous. The species is frequently encountered through alate gynes that appear above ground.
Incisitermes minor
Western Drywood Termite
Incisitermes minor, commonly known as the western drywood termite, is a drywood termite species in the family Kalotermitidae. Native to western North America, it has become established across much of the United States and has been introduced to Hawaii, China, and Japan. It is a significant structural pest, ranking as the second most destructive termite in California after the western subterranean termite (Reticulitermes hesperus), with estimated annual economic impact of approximately $250 million in California and Arizona combined. The species forms one-piece nests entirely within dry wood, does not require soil contact, and exhibits temperature-driven seasonal and daily activity patterns.
Leiobunum
harvestmen, daddy long-legs
Leiobunum is a genus of harvestmen (order Opiliones, family Sclerosomatidae) comprising over 100 described species. Members are characterized by exceptionally long legs relative to body size, with the second pair serving as sensory appendages rather than locomotory structures. The genus exhibits pronounced gregarious behavior, with many species forming dense aggregations on vertical surfaces. Leiobunum species are found across North America, Europe, and Asia, with some populations demonstrating rapid invasive spread in Europe.
Lepidosaphes ulmi
oystershell scale, apple mussel scale
Lepidosaphes ulmi, commonly known as the oystershell scale or apple mussel scale, is a widely distributed armored scale insect in the family Diaspididae. It is a significant pest of deciduous trees and woody plants, including major fruit crops such as apples, pears, plums, and peaches. The species causes damage by attaching to bark and sucking sap, potentially killing branches or entire trees under heavy infestation. It has been introduced to many regions worldwide beyond its native range and is considered invasive in numerous countries.
Leuronota fagarae
Wild Lime Psyllid
Leuronota fagarae (Wild Lime Psyllid) is a psyllid species in the family Triozidae, native to Paraguay and invasive in Florida, USA since 2001. It feeds on Zanthoxylum fagara (wild lime), a citrus relative in the Rutaceae family, causing characteristic rolled leaf edges that shelter developing nymphs. The species is taxonomically related to Diaphorina citri, the vector of Huanglongbing (citrus greening), and their ranges overlap in Florida's transition zones between wild and cultivated citrus habitats. A novel Wolbachia endosymbiont strain (wLfag-FL, supergroup B) has been characterized from this species, revealing potential nutritional provisioning roles and informing research on symbiont-based pest control strategies.
Ligia exotica
sea roach, wharf roach, wharf louse
Ligia exotica is a large intertidal isopod commonly known as sea roach or wharf roach. It inhabits rocky coasts and harbor walls in the supralittoral zone just above the high water mark. The species has a complex taxonomic history with uncertain native origins, though molecular evidence points to East and Southeast Asia as the center of diversity. It has spread globally via human maritime activity and comprises multiple cryptic genetic lineages that may represent distinct species.
Luridiblatta
Luridiblatta is a genus of cockroaches in the family Ectobiidae, established by Fernandes in 1965. The genus contains 10 recognized species, including L. trivittata which has been introduced to California and is known there as the "tiny cockroach." Most species exhibit an almost circum-Mediterranean distribution. A 2022 taxonomic revision described 7 new species and organized them into three species-groups based on morphological characteristics.
Macrancylus linearis
Macrancylus linearis is a terrestrial weevil in the family Curculionidae. The species is believed to have originated in the Pacific region and has been introduced to mainland United States. It has been documented in shoreline habitats, specifically in association with driftwood.
Macropsis fuscula
Raspberry Leafhopper
Macropsis fuscula is a leafhopper species native to Europe, first recorded in North America in 1952 on Lulu Island, British Columbia, where it established as a pest of cultivated loganberry. The species exhibits population dynamics responsive to host availability and climatic events, having crashed after a severe freeze eliminated cultivated hosts in 1955, then rebounded within three years via re-infestation from wild host plants. It is now known from multiple localities across Europe and Asia, with 71 iNaturalist observations documenting its continued presence.
Megalonotus sabulicola
Introduced Dirt-colored Seed Bug
Megalonotus sabulicola is a seed bug in the family Rhyparochromidae, native to Europe and northern Asia. It has been introduced to North America, where it is established and frequently recorded. The species is associated with sandy habitats, as indicated by its specific epithet 'sabulicola' (sand-dwelling). It is one of the more commonly observed dirt-colored seed bugs in its introduced range.
Megasternum concinnum
Megasternum concinnum is a small water scavenger beetle in the family Hydrophilidae. It has a broad native distribution across Europe, North Africa, and western Asia, and has been introduced to western North America. The species belongs to the subfamily Sphaeridiinae, whose members are generally associated with moist organic substrates rather than truly aquatic habitats.
Microplax
Microplax is a genus of true bugs in the family Oxycarenidae, established by Fieber in 1860. The genus includes species native to the Mediterranean region, with at least one species, Microplax albofasciata, having expanded its range northward to the Channel Islands and Germany, and subsequently introduced to North America. The 2012 detection in California represents the first documented occurrence in the United States.
Microtheca ochroloma
Yellow-margined Leaf Beetle, Yellowmargined Leaf Beetle
Microtheca ochroloma is a leaf beetle in the family Chrysomelidae, native to South America and adventive in North America. It is a significant agricultural pest of cruciferous crops, with documented populations in the southeastern United States. Development from egg to adult takes approximately 17–57 days depending on temperature, with optimal survival around 20–25°C. The species exhibits strong host preferences, favoring turnip and mustard while showing least preference for collard and cabbage.
Modisimus culicinus
cellar spider
Modisimus culicinus is a cellar spider (family Pholcidae) native to South America that has become established across tropical and subtropical regions worldwide through human-mediated dispersal. It is considered a pantropical, synanthropic species strongly associated with human structures. The species was previously classified under the genus Hedypsilus, which has been synonymized with Modisimus.
Molytinae
Molytinae is a large and diverse subfamily of weevils (Curculionidae) described by Carl Johan Schönherr in 1823. The subfamily contains numerous tribes and genera distributed worldwide, with particularly high diversity in tropical and subtropical regions. Many Molytinae species are associated with specific host plants, including conifers, oaks, and various flowering plants. Some species are economically important as pests of forestry and agricultural crops, while others are restricted to specialized habitats such as leaf litter, caves, or high-elevation ecosystems. The subfamily has undergone significant taxonomic revision in recent decades, with many new species described particularly from Central America, Africa, and Asia.
Myllocerus undecimpustulatus
Sri Lanka weevil, yellow-headed ravenous weevil
Myllocerus undecimpustulatus is an oriental broad-nosed weevil (Curculionidae: Entiminae) native to Sri Lanka and the Indian subcontinent. It is a polyphagous pest with a wide host range that has established invasive populations in Florida, USA, since at least 2000. The species exhibits distinctive sexual behavior including female abdominal rocking as a mate-strength test and prolonged copulatory guarding by males. It shows seasonal variation in cold tolerance and feeding activity, with winter-collected adults consuming more leaf material after cold exposure than summer-collected individuals.