Subtropical
Guides
Rhagoletotrypeta
Rhagoletotrypeta is a genus of tephritid fruit flies native to the Americas, established by Aczél in 1951. The genus currently includes nine described species distributed across the Neotropics, with records from Cuba, Argentina, and other regions. Species in this genus are fruit-infesting flies whose larvae develop inside the fruits of Celtis species (Cannabaceae). The genus is notable for its extended pupal dormancy periods, which can last approximately 12 months in some populations.
Rhyacionia subtropica
Subtropical Pine Tip Moth
Rhyacionia subtropica is a small tortricid moth with a wingspan of approximately 18 mm. It is distributed in the southeastern United States, Cuba, and Belize. The larvae are specialized feeders on pine species, attacking the terminal shoots of their hosts.
Russellaspis pustulans
oleander pit scale, fig pit scale, Akee Fringed Scale
Russellaspis pustulans, commonly known as the oleander pit scale or fig pit scale, is a parthenogenetic scale insect in the family Asterolecaniidae. It is highly polyphagous, feeding on plants across 69 families with documented preference for fig (Ficus carica) and oleander (Nerium oleander). The species is widely distributed in tropical and subtropical regions worldwide, completing 2-3 generations annually depending on environmental conditions. It is recognized as a pest of economic importance on fruit trees and ornamental plants, with feeding activity inducing characteristic pitting and galling on host tissues.
Scolioidea
Scoliid Wasp Superfamily
Scolioidea is a superfamily of Hymenoptera containing the family Scoliidae, a group of medium-sized wasps with approximately 560 species worldwide. The family is predominantly distributed in tropical and subtropical regions, with 64 species recorded in the New World. Scoliidae are ectoparasitoids of Scarabaeoidea larvae, particularly Melolonthinae (scarab beetles), which they immobilize and parasitize in terrestrial larval galleries. The superfamily has been taxonomically understudied in the Americas, with limited keys and ambiguous historical descriptions.
Strymon alea
Lacey's Scrub-Hairstreak, Alea Hairstreak
Strymon alea is a small hairstreak butterfly in the family Lycaenidae, first described by Godman and Salvin in 1887. It ranges from northwestern Costa Rica through Mexico to central and southern Texas, occupying subtropical thorn scrub habitat. The species has a wingspan of 19–29 mm. Adults feed on flower nectar, while larvae feed specifically on Bernardia myricaefolia buds and flowers.
Tanaostigmatidae
Tanaostigmatidae is a small family of chalcidoid wasps comprising approximately 90 species in 9 genera. Unlike most chalcidoids, they are almost exclusively phytophagous rather than parasitoid, inducing galls in plant stems, leaves, or seeds. The family is primarily distributed in tropical and subtropical regions. Members are typically short, squat wasps with distinctive morphological features including a protruding prepectus and a strongly arched mesonotum that renders the pronotum nearly vertical.
Tetragnatha
Stretch Spiders, Long-jawed Orb Weavers
Tetragnatha is a large genus of long-jawed orb-weaving spiders with hundreds of species distributed worldwide, though most occur in tropical and subtropical regions. These spiders are characterized by their elongated bodies, extremely long legs, and notably elongated chelicerae (jaws) with well-developed teeth. They construct horizontal orb webs, often near water, and many species can run across water surfaces. The genus exhibits remarkable variation in dispersal ability, with some species being excellent dispersers capable of colonizing islands, while others appear to have secondarily lost this capacity. Hawaiian Tetragnatha species have undergone adaptive radiation, with some lineages evolving cursorial (running) behavior and abandoning web-building entirely.
spiderorb-weaverlong-jawedaquaticstretch-spideradaptive-radiationHawaiicursorial-evolutionhorizontal-webwater-walkingcamouflagemultivoltinesocial-spiderTetragnathidaeAraneaeArachnidapredatorinsectivoreriparianwetlandmeadowtropicalsubtropicalcosmopolitandispersalisland-colonizationphylogeographychemical-species-recognitioncommunal-webbingkleptoparasitism-targetTetramorium lanuginosum
Downy Pennant Ant
Tetramorium lanuginosum is a species of ant in the family Formicidae, first described by Mayr in 1870. The species is distributed across the Caribbean region and the Galápagos Islands, with confirmed presence in Antigua and Barbuda, Barbados, Cuba, and Spain (likely referring to the Canary Islands). It belongs to the genus Tetramorium, a diverse group of myrmicine ants commonly known as pavement ants or pennant ants. The common name "Downy Pennant Ant" refers to the species' characteristic appearance.
Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic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Saddlebags, Saddlebags Gliders, Saddlebag Gliders
Tramea is a genus of dragonflies in the family Libellulidae, commonly called saddlebags or saddlebags gliders. Species are distributed across tropical and subtropical regions worldwide. They are characterized by colored bases on otherwise translucent hindwings that create the visual impression of carrying bags near the abdomen base. Members are medium to relatively large dragonflies known for sustained gliding flight and infrequent perching.
Uropygi
whip scorpions, vinegaroons, uropygids
Uropygi is an order of arachnids commonly known as whip scorpions or vinegaroons, characterized by a whip-like flagellum on the posterior end and large scorpion-like pedipalps. They lack venom glands but possess defensive glands capable of spraying acetic and caprylic acid, producing a vinegar-like odor. These nocturnal predators use only six legs for walking, with the first two pairs modified as sensory appendages. The order comprises approximately 100 species across 18 extant genera, all placed in the single family Thelyphonidae.
Xyleborus affinis
sugarcane shot-hole borer
Xyleborus affinis is a highly widespread ambrosia beetle native to the American tropics, now found in tropical and subtropical regions worldwide. It cultivates symbiotic fungi in galleries bored into decaying wood, feeding exclusively on these fungal gardens. The species exhibits facultative eusociality with delayed dispersal of adult offspring, who assist with nest maintenance and brood care. Females are the dispersing sex and are strongly attracted to ethanol and specific host plant volatiles. Despite frequent association with declining trees, it primarily colonizes wood already in early decay rather than causing primary mortality.
Xyleborus volvulus
Xyleborus volvulus is a bark beetle in the family Curculionidae. It is native to tropical and subtropical regions and has been documented in North America, the Caribbean, Middle America, and Africa. The species is an ambrosia beetle that cultivates symbiotic fungi in tunnels bored into wood. It has been observed to carry Raffaelea lauricola, the pathogen causing laurel wilt disease, which has altered its status from a wood degrader to a potential pest of avocado and other lauraceous trees.
Zizula cyna
cyna blue
Zizula cyna, commonly known as the cyna blue, is a small butterfly species in the family Lycaenidae. Adults display violet blue upperwings and pale gray underwings marked with tiny black dots. The species is active from March to November in most of its range, with adults feeding on flower nectar and larvae developing on flower buds of Acanthaceae species.