Freshwater
Guides
Corynoneura
non-biting midges
Corynoneura is a speciose genus of small, non-biting midges in the family Chironomidae. Members occur worldwide except Antarctica, inhabiting both flowing and standing freshwater across various thermal regimes. The genus has been subject to extensive taxonomic revision, with numerous species described from the Neotropical region and Oriental China in recent decades. DNA barcoding studies support the use of molecular data for species delimitation where morphology alone is insufficient.
Corynoneura lobata
Corynoneura lobata is a species of non-biting midge in the family Chironomidae, subfamily Orthocladiinae. First described by Edwards in 1924, this species belongs to a genus of small chironomids characterized by reduced wing venation and distinctive genitalic structures. The genus Corynoneura is taxonomically complex, with numerous species described from multiple biogeographic regions. C. lobata has documented distribution records in northern Europe.
Crangonyctidae
Crangonyctidae is a family of freshwater amphipod crustaceans containing both surface-dwelling and subterranean species. The family includes notable genera such as Crangonyx, Stygobromus, Bactrurus, and Synurella. Many species are obligate groundwater inhabitants adapted to cave and aquifer environments, while others occupy springs, streams, and lakes. The family has a broad geographic distribution across the Northern Hemisphere, with significant diversity in North America, Europe, and Asia.
Crangonyx
cave amphipods, spring amphipods
Crangonyx is a genus of freshwater amphipod crustaceans in the family Crangonyctidae. Species inhabit diverse aquatic environments including surface waters (marshes, swamps, lakes, rivers) and subterranean habitats (caves, springs, groundwater systems). The genus includes both native and highly invasive species, with some taxa exhibiting troglobitic adaptations such as reduced eyes and elongated appendages. Several species have been introduced outside their native ranges, notably Crangonyx pseudogracilis and C. floridanus in Europe and Asia, where they interact competitively and predatorily with native amphipods.
Crangonyx gracilis
Northern Lake Crangonyctid
Crangonyx gracilis is a small freshwater amphipod in the family Crangonyctidae, commonly known as the northern lake crangonyctid. It inhabits diverse aquatic environments across North America, including flowing water, standing water, and temporary habitats. The species was described by S.I. Smith in 1871.
Crangonyx richmondensis
Ellis Bog Crangonyctid
A small freshwater amphipod crustacean endemic to North America. The species exhibits an annual life cycle with distinct seasonal breeding patterns. Populations are restricted to specific freshwater habitats with particular substrate and vegetation characteristics. Two subspecies have been described: C. r. richmondensis and C. r. laurentianus, with the latter studied in detail in Algonquin Park, Ontario.
Crenitis
Crenitis is a genus of water scavenger beetles in the family Hydrophilidae, containing approximately 19 described species. The genus was established by Bedel in 1881. These beetles belong to the subfamily Chaetarthriinae and tribe Anacaenini. They are aquatic or semi-aquatic insects found in freshwater environments.
Cricotopus
non-biting midges
Cricotopus is a large and diverse genus of non-biting midges in the family Chironomidae, subfamily Orthocladiinae. Species occupy a wide range of freshwater and estuarine habitats globally, from pristine glacial streams to degraded urban waterways. Some species are specialized stem miners of aquatic plants, including C. lebetis which targets invasive hydrilla. The genus includes species with documented mutualistic relationships with cyanobacteria (Nostoc), as well as species used in biomonitoring and ecotoxicological research.
Cryptochironomus
non-biting midges
Cryptochironomus is a genus of non-biting midges (family Chironomidae) with over 80 described species distributed worldwide. The genus belongs to the Harnischia generic complex within the subfamily Chironominae. Larvae are aquatic and inhabit bottoms of lakes and large rivers, with some species also found in small streams. Species have been documented from North America, Europe, and Asia, including detailed studies from Lake Winnipeg, Michigan, and China.
Ctenopoda
Ctenopoda is an order of small crustaceans within the superorder Diplostraca, comprising three families: Holopediidae, Pseudopenilidae, and Sididae. Members are commonly known as water fleas and are predominantly freshwater inhabitants, though the genus Penilia is marine. The order is characterized by specialized swimming antennae and a body plan that reflects functional separation between locomotion and feeding appendages. Ctenopoda species have been documented across diverse aquatic habitats including lakes, reservoirs, wetlands, and coastal marine systems, with some species introduced to areas outside their native ranges by human activity.
Curicta
water scorpion
Curicta is a genus of aquatic insects in the family Nepidae, commonly known as water scorpions. Members of this genus are predatory true bugs found in freshwater habitats. The genus was established by Stål in 1861 and is placed in the tribe Curictini within the subfamily Nepinae. Curicta scorpio is one documented species, with records extending from Central to South America.
Cyclopoida
Cyclopoid Copepods
Cyclopoida is an order of small crustaceans within the class Copepoda, comprising approximately 30 families. Members are primarily planktonic, inhabiting both marine and freshwater environments, and are distinguished by morphological features including antennae shorter than the head and thorax combined. The order exhibits metamorphic larval development with embryos carried in paired or single sacs attached to the first abdominal somite. Molecular phylogenetic studies have reclassified the former order Poecilostomatoida as a lineage nested within Cyclopoida.
Cylloepus
riffle beetles
Cylloepus is a genus of riffle beetles in the family Elmidae, first described by Erichson in 1847. The genus contains approximately six described species distributed across the Americas. Members are aquatic beetles associated with flowing water habitats. Species have been documented from North America, the Caribbean, and South America.
Cymatia americana
water boatman
Cymatia americana is a species of water boatman in the family Corixidae, first described by Hussey in 1920. It belongs to the subfamily Cymatiinae and is one of several species in the genus Cymatia found in North America. Water boatmen in this family are aquatic true bugs that inhabit freshwater environments.
Cymatiainae
Cymatiainae is a subfamily of water boatmen within the family Corixidae. Members are aquatic insects adapted to freshwater habitats. The subfamily is distinguished by morphological features of the genitalia and wing venation. Cymatiainae represents one of several lineages within the diverse Corixidae family.
Daphnia
water fleas, water-fleas
Daphnia is a genus of small planktonic crustaceans (0.2–6.0 mm) in the order Anomopoda, commonly called water fleas due to their saltatory swimming style. The genus comprises over 200 species distributed across diverse freshwater habitats worldwide. Daphnia exhibits cyclical parthenogenesis, alternating between asexual and sexual reproduction, and serves as a keystone organism in freshwater food webs. Several species, particularly D. magna and D. pulex, are extensively used as model organisms in ecology, toxicology, and evolutionary biology research.
Daphniidae
water fleas
Daphniidae is a family of small freshwater crustaceans in the order Anomopoda, commonly known as water fleas. The family contains approximately 121 species across five genera: Ceriodaphnia, Daphnia, Megafenestra, Scapholeberis, and Simocephalus. Daphniidae species are important model organisms in ecology, toxicology, and evolutionary biology, particularly the genus Daphnia. Many species have been accidentally introduced to regions outside their native ranges through human activity.
Decapoda
decapods, ten-footed crustaceans
Decapoda is the most species-rich order of Crustacea, with over 14,500 described extant species worldwide. Members include crayfish, crabs, lobsters, prawns, and shrimp—collectively known as decapods or "ten-footed" crustaceans. The order exhibits extraordinary morphological diversity, ranging from tiny symbiotic shrimps under one centimetre to large crabs and lobsters. Decapods occupy virtually every aquatic habitat on Earth, from deep-sea trenches exceeding 5,000 metres depth to terrestrial environments, with nearly half of all species being crabs. The order includes several infraorders with distinct body plans: Brachyura (true crabs), Caridea (shrimps), Anomura (hermit crabs and allies), and others.
Derallus altus
Derallus altus is a water scavenger beetle in the family Hydrophilidae, distributed across the Americas from the United States through Central and South America to the Caribbean. As a member of the tribe Berosini, it inhabits aquatic or semi-aquatic environments where it contributes to nutrient cycling through scavenging behavior. The species was described by John Lawrence LeConte in 1855 and has been recorded in at least 12 U.S. states and multiple countries across the Neotropics.
Desmopachria
Desmopachria is a genus of predaceous diving beetles in the family Dytiscidae, tribe Hyphydrini. The genus contains over 80 described species distributed primarily in the Americas, from the United States through Central America to South America, with particular diversity in the Amazon basin and Caribbean islands. Species are associated with aquatic habitats including river floodplains and high-altitude forest streams. Multiple new species have been described in recent decades, indicating ongoing taxonomic work and likely remaining undescribed diversity.
Diacyclops
Diacyclops is a genus of cyclopoid copepods in the family Cyclopidae containing over 120 described species. Species occupy diverse aquatic habitats including freshwater lakes, wetlands, rivers, and groundwater systems. Some species are planktonic while others are benthic or stygobiontic. The genus shows broad salinity tolerance, with at least one species thriving in brackish conditions up to 5 ppt. Diacyclops species serve as prey for fish and are sensitive indicators of environmental contaminants.
Diacyclops thomasi
Diacyclops thomasi is a cyclopoid copepod species in the family Cyclopidae, first described by Forbes in 1882. The species exhibits a distinctive life cycle involving summer diapause with whole-body encystment at the copepodid IV stage. During encystment, the organism undergoes profound metabolic depression and ultrastructural reorganization of its digestive tract, including transformation of midgut epithelial cells and accumulation of lipid-rich lacunae.
Diaptomidae
Diaptomidae is a family of freshwater pelagic copepods comprising approximately 50 genera. Members are the most widespread copepods in lentic inland waters of the Palearctic region. The family contains two subfamilies: Diaptominae and Paradiaptominae, with the latter consisting almost exclusively of African taxa. Diaptomids exhibit high species richness in tropical and Mediterranean regions, with numerous endemic species.
Dibusa
microcaddisfly
Dibusa is a genus of microcaddisflies in the family Hydroptilidae, established by Ross in 1939. The genus is monotypic, containing a single described species, Dibusa angata. This species exhibits a highly specialized ecological relationship, being obligately associated with the freshwater red alga Lemanea australis throughout its larval development. The genus represents a narrow example of host-specific adaptation within the diverse microcaddisfly fauna.
Dicosmoecus
October Caddisflies
Dicosmoecus is a genus of caddisflies in the family Limnephilidae, commonly known as October Caddisflies. The genus contains approximately six described species distributed across western North America, from California and Oregon through British Columbia to the Rocky Mountains. Species such as D. gilvipes and D. atripes are among the most intensively studied North American caddisflies due to their large larval size, synchronous autumn emergence, and importance in freshwater ecosystem research and fly-fishing culture.
Dicranopselaphus
water-penny beetles
Dicranopselaphus is a genus of water-penny beetles in the family Psephenidae, comprising over 40 described species. The genus exhibits a disjunct distribution spanning Asia and North America. Members are aquatic beetles with distinctive flattened, oval larval forms adapted to clinging to submerged surfaces in flowing water.
Didactyliini
Didactyliini is a small tribe of water scavenger beetles within the family Hydrophilidae, currently containing the single genus Didactylus. Members of this tribe are aquatic beetles distinguished by reduced tarsal segmentation, with only two visible tarsomeres on the fore and mid legs. The tribe was established to accommodate this unusual morphological condition, which separates it from other Hydrophilinae. The genus occurs in the Neotropical region, where species inhabit freshwater environments.
Didymops
Didymops is a genus of dragonflies belonging to the family Macromiidae. The genus is small, containing only two described species. Members are classified within the suborder Anisoptera, representing a distinct lineage within the cruiser dragonfly family.
Dineutus
whirligig beetles
Dineutus is a genus of whirligig beetles (family Gyrinidae) characterized by relatively large body size compared to other whirligig beetles. Adults are 9–15 mm long with smooth, shallowly lined, or grooved elytra. The genus occurs across North America and other regions, with species inhabiting the surface waters of streams and rivers. Members are distinguished from the smaller genus Gyrinus by their larger size and hidden scutellum.
Dineutus angustus
whirligig beetle
Dineutus angustus is a species of whirligig beetle in the family Gyrinidae, first described by LeConte in 1878. The species is found in North America. As a member of the genus Dineutus, it is among the larger whirligig beetles, characterized by a hidden scutellum and rounded elytral apices. Like other gyrinids, it inhabits the surface of freshwater bodies where it forms aggregations called rafts.
Diplectrona modesta
Diplectrona modesta is a species of netspinning caddisfly in the family Hydropsychidae, first described by Banks in 1908. It is native to North America, with documented occurrences in the Nearctic region including Vermont. The species has been studied for its life history and production dynamics in freshwater stream habitats, particularly in Walker Branch, Tennessee.
Diplocladius
Diplocladius is a genus of non-biting midges in the bloodworm family Chironomidae. It belongs to the subfamily Orthocladiinae, a diverse group of chironomid midges commonly found in aquatic and semi-aquatic environments. The genus was established by Kieffer in 1908 and contains four recognized species. These midges are part of the benthic macroinvertebrate communities of freshwater systems.
Diplostraca
water fleas, clam shrimps
Diplostraca is a superorder of small branchiopod crustaceans encompassing over 1,000 described species, commonly known as water fleas and clam shrimps. Members range from 0.2 to 6.0 mm in length (with Leptodora reaching 18 mm), and are characterized by a folded bivalved carapace covering an unsegmented-appearing body. The group exhibits cyclical parthenogenesis, alternating between asexual and sexual reproduction to produce dormant eggs. While predominantly freshwater, eight species have colonized marine environments—the only branchiopods to do so.
Disholcaspis prehensa
clasping twig gall wasp
Disholcaspis prehensa is a gall-inducing cynipid wasp native to California that induces distinctive galls on scrub oak and leather oak. The species exhibits heterogony, alternating between sexual and asexual generations that produce two morphologically different gall types. The asexual generation forms conspicuous mushroom-shaped twig galls with clasping bases, while the sexual generation produces small, hidden bud galls.
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californica
Dixella californica is a species of meniscus midge in the family Dixidae (Order: Diptera). These small, delicate flies are associated with aquatic habitats where their larvae develop. The species was described by Johannsen in 1923 and is known from California. Dixidae larvae are recognized by their distinctive habit of hanging upside down from the water surface film, feeding on microorganisms and organic matter.
Djalmabatista
Djalmabatista is a genus of non-biting midges (Chironomidae) established by Fittkau in 1968. The genus belongs to the subfamily Tanypodinae and tribe Procladiini. Approximately 14 species have been described. These midges are aquatic in their larval stages and are part of the diverse chironomid fauna found in freshwater habitats.
Donacia caerulea
Donacia caerulea is an aquatic leaf beetle in the family Chrysomelidae, subfamily Donaciinae. It is native to North America, with confirmed records from Canadian provinces including Manitoba, New Brunswick, and Ontario. Like other members of its genus, it is associated with freshwater aquatic habitats where its larvae develop on submerged vegetation.
Donacia magnifica
magnificent aquatic leaf beetle, Magnificent pondweed beetle
Donacia magnifica is an aquatic leaf beetle in the family Chrysomelidae, first described by LeConte in 1851. It is native to North America, with records from Canada and the northern United States. The species is closely associated with pondweeds (Potamogeton spp.), on whose leaves adults feed. Like other members of the genus Donacia, it is adapted to an aquatic lifestyle.
Donacia tuberculata
Tuberculate reed beetle
Donacia tuberculata is a species of aquatic leaf beetle in the subfamily Donaciinae. It occurs in North America and is associated with aquatic and semi-aquatic habitats. Like other members of its genus, it is linked to wetland vegetation and has aquatic larvae that develop on submerged plant roots.
Donaciinae
Aquatic Leaf Beetles, Reed Beetles
Donaciinae is a subfamily of leaf beetles (Chrysomelidae) comprising approximately 100 species across six genera, including Donacia, Macroplea, and Plateumaris. Members are strongly associated with aquatic and semi-aquatic habitats, with larvae developing underwater attached to plant roots. The subfamily is characterized by distinctive morphological adaptations and an obligate symbiosis with bacteria that produce the silk-like material used to construct waterproof pupal cocoons. Donaciinae are primarily distributed in the Northern Hemisphere with extensions into Africa, Madagascar, and Australasia.
Dryopidae
long-toed water beetles
Dryopidae is a family of aquatic beetles in the superfamily Byrrhoidea, commonly known as long-toed water beetles. Adults possess dense hydrofuge hairs that enable underwater respiration and extended claws that give the family its common name. Despite being aquatic, adults cannot swim and instead cling to floating detritus. The family is distinguished from similar riffle beetles (Elmidae) by antennae that are not clubbed. Most species have terrestrial larvae, though the genus Stygoparnus has fully aquatic larvae and adults.
Dryops
long-toed water beetles
Dryops is a genus of long-toed water beetles in the family Dryopidae, with at least 20 described species and a worldwide distribution. The genus is particularly diverse in the Palaearctic region, where 34 species have been recorded. Members of this genus are aquatic beetles associated with freshwater habitats. Species such as Dryops rufipes and Dryops renateae have been documented from Turkey and Tajikistan respectively.
Dubiraphia brevipennis
Dubiraphia brevipennis is a species of riffle beetle in the family Elmidae, described by Hilsenhoff in 1973. Like other elmid beetles, it is adapted to life in flowing water environments. The species is known from North America, though specific details about its biology and ecology remain limited in the available literature.
Dubiraphia browni
Dubiraphia browni is a species of riffle beetle (family Elmidae) described by Hilsenhoff in 1973. As a member of the Elmidae, it belongs to a family of aquatic beetles adapted to living in fast-flowing freshwater habitats. The species is known from North America.
Dysmicohermes
Dysmicohermes is a genus of fishflies in the family Corydalidae, established by Munroe in 1953. The genus contains two described species: Dysmicohermes disjunctus and Dysmicohermes ingens. As members of Megaloptera, these insects represent a relatively small and ancient order of holometabolous insects. The genus is part of the diverse Corydalidae family, which includes some of the largest aquatic insects in North America.
Dytiscinae
predaceous diving beetles
Dytiscinae is a subfamily of predaceous diving beetles (family Dytiscidae) containing at least 20 genera and approximately 380 described species. Members are medium to large diving beetles found in aquatic habitats across most continents. The subfamily includes well-known genera such as Dytiscus, Cybister, Acilius, and Hydaticus. Species are predatory in both adult and larval stages, with larvae often called 'water tigers' for their voracious feeding habits.
Dytiscus fasciventris
Understriped Diving Beetle
Dytiscus fasciventris, the Understriped Diving Beetle, is a predaceous diving beetle in the family Dytiscidae. The species is found in North America, with records from Canada (Alberta, British Columbia) and the United States. Like other members of the genus Dytiscus, it is an aquatic predator inhabiting freshwater environments.
Dytiscus harrisii
Harris's diving beetle
Dytiscus harrisii, commonly known as Harris's diving beetle, is a predaceous diving beetle in the family Dytiscidae. It is one of the larger diving beetle species in North America. As a member of the genus Dytiscus, it is an aquatic predator inhabiting freshwater systems.
Dytiscus marginicollis
giant green water beetle
Dytiscus marginicollis, commonly known as the giant green water beetle, is a predaceous diving beetle in the family Dytiscidae. It is among the larger species in the genus Dytiscus and is distributed across North America. Like other members of its genus, it is an aquatic predator inhabiting freshwater systems.
Ecdyonurus
Ecdyonurus is a genus of mayflies in the family Heptageniidae, comprising approximately 66 described species distributed primarily across Europe and parts of North Africa and Asia. The genus is taxonomically well-studied, with multiple species described in recent decades from mountain regions including the Carpathians, Crimean Peninsula, and Aurès Mountains. Species within this genus are typically associated with running waters and have been subjects of ecological studies on larval growth and thermal biology.