Detritivore
Guides
Hexagenia limbata
Giant Mayfly, Golden Mayfly, Big Michigan Mayfly, Great Leadwing Drake, Fishfly
Hexagenia limbata is a large burrowing mayfly native to North America, widely distributed across lakes and slow-moving rivers. Nymphs construct U-shaped burrows in muddy substrates and serve as important prey for fish and other aquatic predators. Adults emerge in synchronized mass events known as "hatches," living only 1–3 days without feeding, solely to mate and reproduce. The species is economically significant to sport fishing and serves as a bioindicator of clean freshwater ecosystems.
Holoverticata
Woodlice and Pillbugs
Holoverticata is an infraorder of isopod crustaceans encompassing the familiar terrestrial woodlice and pillbugs. Members of this group are distinguished by their dorsoventrally flattened bodies, seven pairs of walking legs, and ability to occupy moist terrestrial habitats. The group includes species capable of conglobation (rolling into a ball) as well as those that remain flattened. This infraorder represents the most successful lineage of crustaceans to colonize land.
Hyalella
Hyalella is a genus of freshwater amphipods found in the Americas, with species distributed across North, Central, and South America. The genus contains numerous endemic species, particularly in South America, and includes the widely studied H. azteca, which serves as a standard test organism in aquatic toxicology. Members occupy benthic habitats in lakes, streams, and springs, where they function as important components of freshwater food webs.
Hyalella wellborni
Hyalella wellborni is a freshwater amphipod species in the family Hyalellidae, described in 2015 from the southeastern United States. The genus Hyalella comprises small benthic crustaceans commonly known as scuds or sideswimmers, widespread in lakes, ponds, and streams. H. wellborni represents part of a taxonomically complex group where species delineation has historically relied on morphological and molecular analyses. The species is known from a limited number of observations, reflecting both its relatively recent description and the ongoing challenges in amphipod taxonomy.
Hydatophylax
northern caddisfly
Hydatophylax is a genus of northern caddisflies (Trichoptera: Limnephilidae) comprising approximately 14 described species. Members are found in cool temperate regions of the Northern Hemisphere, including Scandinavia, Japan, and North America. The genus exhibits univoltine life cycles with larval development in freshwater streams.
Hypenula
litter moths
Hypenula is a genus of litter moths in the subfamily Herminiinae, family Erebidae. These moths are associated with forest floor habitats where their larvae feed on decaying plant matter. The genus was established by Grote in 1876 and contains multiple species distributed in North America.
Idia
litter moths, American idia moths
Idia is a genus of litter moths in the family Erebidae, subfamily Herminiinae. These moths are primarily nocturnal and are commonly attracted to light sources. The genus includes the well-known American Idia Moth (Idia americalis) and related species. Members of this genus are found across North America and are frequently documented in citizen science projects such as iNaturalist.
Idia aemula
Common Idia, Powdered Snout, Waved Tabby
A small litter moth in the family Erebidae, recognized by its gray forewings with intricate dark lines and a distinctive pale to orange-brown reniform spot. Adults are nocturnal and active from spring through fall, with multiple generations per year. The larvae feed on dead leaves, contributing to nutrient cycling in forest ecosystems. The species is widespread across eastern North America and has been reported in the Palearctic region.
Idia diminuendis
Orange-spotted Idia Moth
Idia diminuendis is a small litter moth in the family Erebidae, first described in 1918. It occurs across eastern and central North America. The species has two generations per year in most of its range and is attracted to light.
Idia forbesii
Forbes' Idia Moth
Idia forbesii is a small litter moth in the family Erebidae, first described by George Hazen French in 1894. The species is widely distributed across eastern North America, with populations exhibiting univoltine life cycles in northern regions and multivoltine cycles in southern regions. Adults are active from late spring through fall depending on latitude.
Idia julia
Julia's Idia Moth, Julia's idia
Idia julia is a small litter moth in the family Erebidae, first described by William Barnes and James Halliday McDunnough in 1918. The species is distributed across eastern North America, ranging from southern Canada to Georgia and Texas. Adults have a wingspan of approximately 17 mm. Larvae feed on detritus, particularly dead leaves.
Idia laurentii
Laurentine Idia, Appalachian Idia
Idia laurentii is a litter moth in the family Erebidae, first described by J. B. Smith in 1893. It is endemic to the Appalachian region of the eastern United States, ranging from central New York south to the mountains of North Carolina. The species has a univoltine life cycle with one generation per year. Larvae have been documented feeding on dead cherry leaves.
Idia rotundalis
Rotund Idia Moth, Chocolate Idia
Idia rotundalis is a small litter moth in the family Erebidae, subfamily Herminiinae. First described by Francis Walker in 1866, it is widespread across eastern North America. The species exhibits latitudinal variation in voltinism, with one generation annually in northern populations and two or more generations in southern populations. Larvae are detritivores that feed on dead leaves and other organic debris.
Idia scobialis
Smoky Idia Moth, smoky idia
Idia scobialis is a small litter moth in the family Erebidae, first described by Grote in 1880. It occurs across eastern North America from southern Canada to the southeastern United States. The species has a wingspan of approximately 20 mm and completes one generation per year. Larvae are detritivores, feeding on dead leaves and other organic debris.
Idoteidae
Common Valvetails
Idoteidae is a family of aquatic isopod crustaceans in the suborder Valvifera, distributed globally in marine and freshwater habitats. The family includes approximately 20 genera and numerous species, with highest diversity in temperate coastal waters. Members range from free-living forms in macroalgae and seagrass beds to commensal species associated with other marine organisms. The family has been extensively studied in Australia, New Zealand, the northeastern Pacific, and the North Atlantic.
Indiopsocus lanceolatus
Indiopsocus lanceolatus is a species of barklouse in the family Psocidae, described by Mockford and Young in 2015. The species belongs to the genus Indiopsocus, which comprises common barklice found in various habitats across North America. As a member of Psocodea, it possesses chewing mouthparts and is typically associated with dead plant material, bark, and leaf litter.
Isopoda
isopods, woodlice, pillbugs, sowbugs, sea slaters, gribbles
Isopoda is an ancient order of crustaceans encompassing over 10,000 described species across marine, freshwater, and terrestrial habitats. Members are characterized by dorsoventrally flattened bodies, seven pairs of similar walking legs (giving the group its name from Greek iso- "equal" and pod- "foot"), and two pairs of antennae. The order exhibits exceptional morphological diversity, ranging from minute interstitial forms to giant deep-sea species exceeding 30 cm in length. Isopods lack a carapace, instead possessing overlapping dorsal plates that provide flexibility and protection. Females brood eggs in a specialized marsupium formed by oostegites under the thorax.
Jaera
Jaera is a genus of small marine isopods in the family Janiridae, comprising more than 20 described species. The genus is notable for the Jaera albifrons species complex, a group of closely related, sympatric species that exhibit fine-scale habitat partitioning along intertidal shores. These isopods are euryhaline, capable of osmoregulation across wide salinity ranges from freshwater-influenced areas to fully marine conditions. The group has been extensively studied for its ecological differentiation, reproductive isolation, and as a model for understanding speciation processes in marine environments.
Julida
Snake Millipedes
Julida is an order of millipedes commonly known as snake millipedes due to their long, cylindrical body form. Members typically range from 10–120 mm in length and are characterized by having two pairs of legs per body segment, a trait distinguishing them from centipedes. The order exhibits considerable diversity with 593 species recorded from Europe alone, and includes families such as Julidae, Parajulidae, Blaniulidae, and Zosteractinidae. Many species are important decomposers in forest ecosystems.
Julus scandinavius
Julus scandinavius is a millipede species in the family Julidae, described by Latzel in 1884. It is distributed across much of western and central Europe, extending into Scandinavia. The species has been studied for its behavioral preferences regarding leaf litter composition.
Lacinipolia incurva
Lacinipolia incurva is a small owlet moth in the family Noctuidae, described by John B. Smith in 1888. The species is restricted to the southwestern United States and adjacent regions of Mexico, with records from California, Arizona, New Mexico, Texas, Utah, and Colorado. Adults have a wingspan of approximately 25 mm. The larvae are known to feed on dead leaves of Quercus hypoleucoides (silverleaf oak), indicating a detritivorous or saprophagous feeding strategy rather than typical herbivory on living plant tissue.
Lacunicambarus diogenes
devil crayfish, devil crawfish
Lacunicambarus diogenes, commonly known as the devil crayfish or devil crawfish, is a primary burrowing crayfish native to eastern North America. This species constructs and inhabits burrows in wet, muddy terrestrial habitats rather than living in permanent surface water. Its burrowing activities create refugia used by numerous other species, including documented use by eastern cicada killer wasps (Sphecius speciosus) as brooding habitat. The species ranges across the Atlantic Coastal Plain and Piedmont ecoregion from New Jersey to Georgia, with disjunct populations in Louisiana.
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harvestmen, daddy long-legs
Leiobunum is a genus of harvestmen (order Opiliones, family Sclerosomatidae) comprising over 100 described species. Members are characterized by exceptionally long legs relative to body size, with the second pair serving as sensory appendages rather than locomotory structures. The genus exhibits pronounced gregarious behavior, with many species forming dense aggregations on vertical surfaces. Leiobunum species are found across North America, Europe, and Asia, with some populations demonstrating rapid invasive spread in Europe.
Lepidostoma
Lepidostoma is a genus of caddisflies in the family Lepidostomatidae comprising over 150 described species. The genus is notable for the distinctive case-building behavior of its larvae, which construct portable cases from plant materials, primarily leaf panels in later instars. Larvae are detritivores that process allochthonous organic matter in freshwater streams. The genus has a broad geographic distribution including North America, Europe, Asia, and Africa.
Lepinotus reticulatus
reticulate-winged trogiid, reticulate-winged booklouse, granary booklouse
Lepinotus reticulatus is a species of granary booklouse in the family Trogiidae. It is one of the most widely distributed psocids, occurring across six continents in association with stored grain and dry organic materials. The species is frequently encountered in anthropogenic environments, particularly granaries, warehouses, and food storage facilities. Its common name refers to the distinctive reticulate wing venation pattern visible in winged morphs.
Lepismatidae
Typical Silverfishes
Lepismatidae is a family of primitive, wingless insects in the order Zygentoma, containing approximately 190-340 described species worldwide. The family includes the two most familiar domestic species: the silverfish (Lepisma saccharina) and the firebrat (Thermobia domestica). These ancient insects represent some of the earliest diverging lineages within Insecta, with origins dating back hundreds of millions of years. Members are characterized by elongated, flattened bodies covered in scales, three caudal filaments, and a complete absence of wings throughout their life cycle.
Leptophlebia
Early brown spinner, Sepia dun, Claret dun
Leptophlebia is a genus of mayflies in the family Leptophlebiidae, comprising approximately 11 described species distributed across the Northern Hemisphere. Nymphs are primarily detritivores that inhabit lentic waters, slow-flowing streams, and floodplain wetlands, with documented movements between river channels and temporary wetland habitats. Several species, including L. vespertina and L. cupida, have been studied as model organisms for understanding life cycle plasticity, acid tolerance, and river-floodplain connectivity in freshwater ecosystems.
Leptophlebia cupida
Early Brown Spinner, Black Quill
Leptophlebia cupida is a pronggilled mayfly species native to North America, commonly known as the early brown spinner or black quill. The species exhibits a univoltine life cycle with egg diapause during summer months. Nymphs develop through approximately 20-34 instars over 10 months, with emergence occurring from late April to mid-May. Adults are short-lived, non-feeding, and mate in swarms near streams.
Ligia oceanica
sea slater, common sea slater, sea roach
Ligia oceanica is a large intertidal isopod reaching up to 35 mm in length, making it the largest species in the suborder Oniscidea. Native to rocky Atlantic coasts of Europe, it has been introduced to eastern North America and Atlantic islands. This semelparous species inhabits the supralittoral zone, hiding in rock crevices by day and emerging nocturnally to feed. Molecular phylogenetics suggests closer affinity to marine isopod suborders Valvifera and Sphaeromatidea than to terrestrial woodlice, challenging traditional classification.
Limnephilus moestus
Limnephilus moestus is a species of northern caddisfly in the family Limnephilidae, described by Nathan Banks in 1908. Like other members of its genus, it is associated with lentic (still water) habitats. The species is recorded from North America with distribution records in the Nearctic region. As with many Limnephilus species, adults are typically active in late summer and fall.
Liogma nodicornis
Liogma nodicornis is a species of cylindrotomid crane fly native to North America. Larvae inhabit wet environments including streams, marshes, and saturated soils beneath alders, where they function as detritivores. The species exhibits an approximately two-year life cycle with adults emerging during summer months.
Lirceus
Flat Waterslaters
Lirceus is a genus of freshwater isopod crustaceans in the family Asellidae, commonly known as flat waterslaters. The genus contains at least 18 described species distributed across southern Canada and the eastern United States, extending westward to the Great Plains. Many species inhabit caves and springs, particularly in the Interior Highlands region (Ozark and Ouachita Mountains), while others occupy surface streams and leaf pack habitats. Two species are listed as endangered or vulnerable on the IUCN Red List.
Lithoseopsis hysteryx
Mystery Caddisfly
Lithoseopsis hysteryx is a species of caddisfly in the family Lepidostomatidae, described by Ross in 1956. The species is known from limited collections in western North America. Adults are small to medium-sized caddisflies with reduced wing venation characteristic of the genus. The larval stage constructs portable cases using mineral particles.
Littorophiloscia richardsonae
Western Saltmarsh Woodlouse
Littorophiloscia richardsonae is a small terrestrial isopod commonly known as the Western Saltmarsh Woodlouse. It belongs to the family Halophilosciidae and has been documented primarily in coastal saltmarsh habitats of western North America. The species was described in 1909 and represents one of the few truly halophilic (salt-tolerant) woodlice in the region.
Machilinae
Machilinae is a subfamily of bristletails within the family Machilidae, comprising one of the two major lineages of the jumping bristletail family. Members are small, wingless insects with the characteristic arched thorax and springing organ (furcula) that enables their distinctive jumping locomotion. The subfamily has been historically distinguished from the other machilid subfamily, Petrobiinae, primarily by subtle differences in abdominal appendage structure and scale patterns. Machilinae species are found across temperate and Mediterranean regions, often occupying rocky, coastal, or urban habitats.
Macrochilo litophora
Brown-lined Owlet, Angulate Fan-foot, Brown-lined Owlet Moth
Macrochilo litophora is a small litter moth in the subfamily Herminiinae, first described by Augustus Radcliffe Grote in 1873. It occurs across the eastern and central United States. The species exhibits regional variation in voltinism, with one generation annually in northern populations and two generations in parts of the Midwest. Larvae are detritivores, feeding on dead plant material.
Martyringa xeraula
Himalayan Grain Moth
Martyringa xeraula, known as the Himalayan grain moth, is a small moth in the family Lecithoceridae described by Edward Meyrick in 1910. It has an unusually broad distribution spanning Asia (India, western China, Japan) and North America (southern United States from Louisiana to South Carolina). The species is associated with stored products and detritus-feeding habits. Its transcontinental range suggests either natural dispersal or human-mediated introduction, though the mechanism remains unclear.
Megatoma ampla
carpet beetle
Megatoma ampla is a species of carpet beetle in the family Dermestidae, first described by Casey in 1900. It belongs to a genus of beetles known for infesting stored products and natural history collections. The species is documented from North America, with specific records from British Columbia, Canada. As with other dermestid beetles, it likely has a scavenging or detritivorous lifestyle, though specific ecological details for this species remain poorly documented.
Metylophorus
common barklice
Metylophorus is a genus of barklice in the family Psocidae, established by Pearman in 1932. The genus contains at least 50 described species distributed across multiple continents. As members of Psocidae, these insects are commonly found in association with tree bark and other woody substrates. The genus is taxonomically placed in the tribe Metylophorini within the subfamily Psocinae.
Milesia virginiensis
yellowjacket hover fly, Virginia flower fly, Virginia Giant Hover Fly, News Bee
A large, striking syrphid fly native to eastern North America. Adults are notable mimics of yellowjackets and hornets, complete with yellow, brown, and black coloration and a loud droning buzz. The species is active primarily in mid-summer to early fall, frequenting forest edges and meadows. Larvae develop in decaying wood. The species carries extensive American folklore, commonly known as the "News Bee" for its habit of hovering near people.
Molannidae
Hood Casemaker Caddisflies
Molannidae is a small family of caddisflies (Trichoptera) containing approximately 40 described species across three genera: Molanna, Molannodes, and Indomolannodes. The family occurs in the Holarctic and Oriental biogeographic regions. Adults are commonly known as "hood casemakers" and have a distinctive appearance in repose, resembling short branch segments. Larvae construct portable cases and inhabit lentic and slow lotic environments, primarily on sandy substrates.
Monopis monachella
White-blotched Clothes Moth
Monopis monachella is a small tineid moth with a nearly cosmopolitan distribution spanning Eurasia, Africa, Asia, the Americas, and Oceania. The species is commonly known as the White-blotched Clothes Moth and has been observed feeding on animal remains during its larval stage. Adults are active from spring through late summer in temperate regions.
Montescardia fuscofasciella
Montescardia fuscofasciella is a species of moth in the family Tineidae, described by Chambers in 1875. It belongs to a genus of small moths commonly associated with detritivorous or keratinophagous feeding habits. The species is known from limited records in the eastern United States.
Motyxia
Sierra luminous millipedes, motyxias
Motyxia is a genus of blind, cyanide-producing millipedes endemic to three mountain ranges in California. All 11 species exhibit bioluminescence, making them one of only three known bioluminescent millipede groups worldwide. Adults range 3–4 cm in length with 20 body segments and prominent lateral keels (paranota). The genus was established by Chamberlin in 1941 and belongs to the tribe Xystocheirini within the family Xystodesmidae.
Myriapoda
myriapods
Myriapoda is a subphylum of terrestrial arthropods comprising approximately 13,000–16,000 described species across four extant classes: Chilopoda (centipedes), Diplopoda (millipedes), Pauropoda, and Symphyla. All myriapods are obligate terrestrial, characterized by elongated bodies with numerous segments bearing legs. The group represents one of the earliest arthropod lineages to colonize land, with fossil evidence dating to the Late Silurian–Early Devonian boundary. Myriapods exhibit diverse ecological roles: centipedes are primarily nocturnal predators using venomous forcipules, while millipedes, pauropods, and symphylans function predominantly as detritivores in soil and leaf litter ecosystems.
Nannariini
Nannariini is a tribe of flat-backed millipedes within the family Xystodesmidae, subfamily Rhysodesminae. The tribe was established by Hoffman in 1964 and comprises several genera of small to medium-sized polydesmidan millipedes found primarily in North America. Members of this tribe are characterized by specific gonopodal modifications that distinguish them from related tribes within Rhysodesminae.
Narceus
Narceus is a genus of large cylindrical millipedes in the family Spirobolidae native to eastern North America. The genus includes some of the largest millipedes in the region, with individuals reaching up to 12 cm in length. It comprises three to four recognized species, including two Florida endemics and a widespread species complex (N. americanus/annularis) spanning eastern Canada to the southern United States. These millipedes are significant decomposers in forest ecosystems and serve as intermediate hosts for certain parasites.
Narceus americanus-annularis-complex
A species complex of large North American millipedes comprising two closely related, morphologically similar species: Narceus americanus and Narceus annularis. These are among the largest millipedes in eastern North America, reaching lengths over 100 mm. The two species are difficult to distinguish without detailed examination of gonopod morphology, leading to frequent misidentification and the recognition of this unresolved complex. They are slow-moving detritivores found in moist forest habitats.
Nearctodesmus
Nearctodesmus is a genus of small millipedes in the order Polydesmida, family Nearctodesmidae. These millipedes are characterized by their flattened bodies and reduced segmentation. The genus was established by Silvestri in 1910 and is primarily distributed in the Nearctic region. Members of this genus are part of the diverse soil fauna and contribute to decomposition processes in forest ecosystems.
Nemapogon
Fungus moths
Nemapogon is a genus of small tineid moths in the subfamily Nemapogoninae, comprising approximately 69 described species as of 2007. Species occupy woodland habitats where larvae develop within bracket fungi on dead wood. Some species are attracted to light and may occasionally be captured in pheromone traps intended for clothes moths. The genus includes species with divergent feeding habits: most are fungivores, while at least one species (N. gersimovi) has been intercepted feeding on stored seeds and grains.