Gall-wasp
Guides
Ormyrus venustus
Ormyrus venustus is a species of parasitoid wasp in the family Ormyridae, described by Hanson in 1992. Like other members of the genus Ormyrus, it is a parasitoid of gall-forming insects. The genus Ormyrus has been the subject of recent research revealing extensive cryptic species diversity, with morphologically identical populations often representing multiple genetically distinct species. O. venustus is known from scattered records across the United States including Arizona, California, Colorado, Maryland, and Massachusetts.
Paracraspis patelloides
Bowl Gall Wasp
Paracraspis patelloides, commonly known as the Bowl Gall Wasp, is a species of cynipid wasp that induces distinctive gall formation on host plants. First described by Trotter in 1910 under the genus Andricus, it was later transferred to Paracraspis. The species is part of the diverse Cynipidae family, which contains numerous gall-inducing wasps primarily associated with oaks and related plants. Observations suggest it is moderately well-documented, with over 100 records on iNaturalist.
Periclistus
Periclistus is a genus of micro-hymenopteran gall wasps (Cynipidae) comprising at least 17 described species. These wasps are obligate inquilines that occupy galls induced by other cynipid genera on plants in the rose family (Rosaceae), particularly roses (Rosa). Unlike gall inducers, Periclistus species have lost the ability to initiate galls but retain the capacity to modify existing gall tissue, inducing nutritive tissue for their own larvae. The genus exhibits a phylogenetic divide between Palaearctic and Nearctic clades, with host specificity ranging from specialists to generalists across different species.
Phanacis
Phanacis is a genus of cynipid gall wasps (Hymenoptera: Cynipidae) in the tribe Aylacini. The genus includes species that induce galls on host plants, with Phanacis taraxaci documented as forming leaf galls on common dandelion (Taraxacum officinale). These galls function as physiological sinks that actively redirect carbon and mineral nutrients from host tissues.
Phanacis hypochoeridis
Catsear Gall Wasp
Phanacis hypochoeridis is a gall wasp in the family Cynipidae. It induces galls on plants in the Asteraceae family, specifically on catsear (Hypochaeris radicata) and related species. The species is native to Europe and has been introduced to other regions including Oceania. As a gall-former, it manipulates host plant tissue to create a protective structure for its developing larvae.
Phanacis taraxaci
dandelion gall wasp
Phanacis taraxaci is a gall-forming cynipid wasp that induces galls on the leaves of common dandelion (Taraxacum officinale). The galls function as physiological sinks, actively redirecting carbon and mineral nutrients from host plant tissues. This species has been documented in Europe, northern Asia, and North America.
Philonix
oak gall wasps
Philonix is a genus of oak gall wasps in the family Cynipidae, restricted to North America (Canada, United States, and Mexico). Species in this genus induce spherical, soft, fleshy galls on white oaks (Quercus sect. Quercus), typically on the undersides of leaves. The genus was first described by Asa Fitch in 1859. Unlike many cynipid wasps, alternate sexual and asexual generations have not been documented in Philonix.
Philonix fulvicollis
Philonix fulvicollis is a gall wasp species in the family Cynipidae, first described by Fitch in 1859. It has an alternation of sexual and asexual generations, a phenomenon confirmed through molecular analysis of cytb and ITS2 sequences. The species is distributed across eastern and central North America, with records from the United States and Canada. Like other cynipid wasps, it induces gall formation on host plants, though specific host associations require further documentation.
Philonix gigas
Philonix gigas is a species of gall wasp in the family Cynipidae, first described by Weld in 1922. The genus Philonix is part of the diverse Cynipidae family, which comprises insects known for inducing plant galls. As with many Cynipidae species, specific biological details for P. gigas remain poorly documented in available literature. The species name 'gigas' suggests relatively large size compared to congeners, though this has not been quantified in accessible sources.
Phylloteras
Phylloteras is a North American genus of gall wasps in the family Cynipidae, tribe Cynipini. These insects are specialized associates of oak trees (Quercus), inducing galls on leaves. The genus contains at least 4 described species, with estimates suggesting up to 11 species may exist. As members of the oak gall wasp tribe, they exhibit the characteristic life history of alternating generations between sexual and asexual forms, often with distinct gall morphologies for each generation.
Phylloteras cupella
urn gall wasp, banded urn gall wasp
Phylloteras cupella is a cynipid wasp that induces distinctive leaf galls on multiple oak species in western North America. The wasp measures 1.3–2 mm in length. Its galls display a characteristic ombré color gradient, transitioning from lighter tones at the top to darker shades below, with new galls appearing yellow, red, or mauve, aging to purple, and ultimately turning brown. The species was originally described as Trigonaspis cupella by Weld in 1926 and later transferred to Phylloteras.
Phylloteras sigma
Phylloteras sigma is a species of gall wasp in the family Cynipidae. As a member of this family, it is an obligate gall-former on oak trees (Quercus species). The genus Phylloteras is associated with leaf galls on oaks, and species in this genus induce distinctive galls on oak foliage. Like other Cynipidae, P. sigma has a complex life cycle that typically alternates between sexual and asexual generations, though specific details for this species remain poorly documented.
Phylloteras volutellae
conical oak gall wasp
Phylloteras volutellae is a cynipid gall wasp native to North America that induces distinctive conical galls on the leaves of white oaks (Quercus section Quercus). The species is best known for the detachable, clustered galls formed on leaf undersides, each housing a single larva. Adults emerge in early January, though adult morphology remains undescribed.
Protobalandricus spectabilis
Tapered Stem Gall Wasp
Protobalandricus spectabilis is a gall wasp species in the family Cynipidae, originally described as Andricus spectabilis by Kinsey in 1922 and later transferred to the genus Protobalandricus by Nicholls, Stone & Melika in 2018. It is commonly known as the Tapered Stem Gall Wasp. The species is part of the diverse Cynipini tribe, which includes numerous gall-inducing wasps associated with oaks and related plants. As a cynipid wasp, it induces the formation of plant galls during its life cycle, though specific gall morphology and host associations for this species require further documentation.
Quadrastichus
Quadrastichus is a genus of small parasitic wasps in the family Eulophidae, subfamily Tetrastichinae. The genus includes species with diverse ecological roles: some are gall-inducing pests of trees, while others are ectoparasitoids used in biological control programs. Notable species include Quadrastichus erythrinae, an invasive gall wasp that damages Erythrina (coral trees) in tropical and subtropical regions, and Q. mendeli, an obligate ectoparasitoid of the eucalyptus pest Leptocybe invasa that has been successfully established for biological control across multiple countries.
Sphaeroteras
Sphaeroteras is a genus of cynipid wasps established by Ashmead in 1897. It is currently recognized as a synonym of Biorhiza, a genus of gall wasps that induce galls on oak trees. Records from Scandinavia suggest these wasps are associated with temperate European oak woodlands.
Sphaeroteras pulchripennis
Sphaeroteras pulchripennis is a species of gall wasp in the family Cynipidae. The species name 'pulchripennis' (Latin: 'beautiful wing') refers to its distinctive wing coloration. As with other members of Cynipidae, this species induces gall formation on host plants, though specific host associations remain incompletely documented. It is a relatively rarely observed species with limited published biological data.
Striatoandricus
Striatoandricus is a genus of Neotropical gall wasps in the family Cynipidae, established by Pujade-Villar in 2020. The genus comprises approximately six described species, with four species transferred from the genus Andricus. Despite morphological similarities to Andricus, phylogenetic analyses indicate the two genera are not closely related. The genus name refers to the striations on the metasomal segments combined with its superficial resemblance to Andricus.
Striatoandricus aciculatus
Striatoandricus aciculatus is a species of gall wasp in the family Cynipidae, recently transferred from the genus Andricus to Striatoandricus based on phylogenetic and morphological studies. Like other members of its tribe, this species induces galls on oak trees (Quercus spp.) as part of its reproductive cycle. The species was originally described in 1909 and has been documented across multiple observations, indicating established populations in regions with suitable oak host availability.
Striatoandricus furnessulus
Striatoandricus furnessulus is a species of gall wasp in the family Cynipidae. It was originally described by Weld in 1944 and later transferred to the genus Striatoandricus in 2022. The species is known from Arizona, USA. As a member of the oak gall wasps, it likely induces galls on oak species (Quercus), though specific host associations have not been documented in available sources.
Synergini
inquiline gall wasps
Synergini is a tribe of inquiline gall wasps in the family Cynipidae. Members have lost the ability to induce their own galls and instead inhabit galls created by other cynipid wasps, primarily those in tribe Cynipini. The tribe includes genera such as Synergus, Saphonecrus, Lithosaphonecrus, and Ufo. Most species are associated with oaks (Quercus) and related Fagaceae. A 2015 taxonomic revision transferred three former Synergini genera to other tribes: Ceroptres to its own tribe Ceroptresini, and Periclistus and Synophromorpha to Diastrophini.
Synergus
inquiline oak gall wasps
Synergus is a genus of inquiline gall wasps in the tribe Synergini (Hymenoptera: Cynipidae). Most species develop within galls induced by other cynipid wasps on oaks (Quercus), though some inhabit galls on chestnuts (Castanea). The genus is distributed worldwide, with notable diversity in the New World and Eastern Palaearctic. One species, S. itoensis, has independently evolved the ability to induce its own galls inside oak acorns, representing a rare transition from inquilinism to gall induction. Species are morphologically grouped by fore wing radial cell structure and mesopleural sculpture.
Tanaostigmatidae
Tanaostigmatidae is a small family of chalcidoid wasps comprising approximately 90 species in 9 genera. Unlike most chalcidoids, they are almost exclusively phytophagous rather than parasitoid, inducing galls in plant stems, leaves, or seeds. The family is primarily distributed in tropical and subtropical regions. Members are typically short, squat wasps with distinctive morphological features including a protruding prepectus and a strongly arched mesonotum that renders the pronotum nearly vertical.
Tanaostigmodes albiclavus
Tanaostigmodes albiclavus is a species of chalcid wasp in the family Tanaostigmatidae, first described by Girault in 1917. The species is known from the southwestern United States. Members of this family are gall wasps associated with plants, though specific host relationships for this species remain poorly documented.
Tetramesa
Tetramesa is a genus of minute phytophagous wasps in the family Eurytomidae comprising over 200 described species. Members are exclusively associated with grasses (Poaceae), where they typically induce stem or inflorescence galls. The genus exhibits pronounced host specificity, with most species restricted to a single grass species or closely related congeners. Several species have been deployed as biological control agents against invasive grasses, including T. romana for giant reed (Arundo donax) and candidate agents for medusahead (Taeniatherum caput-medusae) and African lovegrass (Eragrostis curvula). Adults feed on nectar. Recent phylogenomic studies have synonymized the genera Aiolomorphus and Cathilaria within Tetramesa.
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koebelei
Torymus koebelei is a species of chalcid wasp in the family Torymidae. The species was described by Huber in 1927. Torymus species are generally known as parasitoids of gall-forming insects, particularly gall wasps and gall midges. T. koebelei has been documented in the western United States, including California, where it has been reared from oak apple galls.
Torymus zabriskii
Torymus zabriskii is a species of parasitoid wasp in the family Torymidae. The species was originally described as Diomorus zabriskii by Cresson in 1878 and subsequently transferred to the genus Torymus. Torymid wasps in this genus are known to develop as parasitoids within insect-induced plant galls.
Trigonaspis
Trigonaspis is a genus of cynipid wasps in the family Cynipidae, tribe Cynipini. These insects are gall wasps that induce galls on host plants. The genus was established by Hartig in 1840 and is part of the diverse radiation of oak gall wasps and related lineages.
Trigonaspis polita
Trigonaspis polita is a species of gall wasp in the family Cynipidae, originally described as Cynips polita by Bassett in 1881. The species was later transferred to the genus Trigonaspis. Like other cynipids, it is associated with the formation of plant galls, though specific gall morphology and host associations for this species require further documentation. The genus Trigonaspis is part of the diverse Cynipidae family, which contains numerous species that induce galls on oaks and related plants.
Trigonaspis quercusforticorne
Trigonaspis quercusforticorne is a cynipid wasp that induces galls on oak trees. It is one of several oak gall wasps in the genus Trigonaspis, which are characterized by their distinctive gall formations on specific host tissues. The species has been documented across multiple European countries and is associated with Quercus species. Its biology and life history follow the typical pattern of unilocular cynipid galls with alternating sexual and asexual generations.
Trigonaspis radicola
Trigonaspis radicola is a species of gall wasp in the family Cynipidae. The species is known for inducing root galls on oak species (Quercus), distinguishing it from many cynipids that form galls on above-ground plant parts. Like other members of the tribe Cynipini, it has a complex life cycle involving alternating sexual and asexual generations, typically associated with different oak host species or plant structures. The species name 'radicola' (meaning 'root-dwelling') reflects its distinctive subterranean gall-forming habit.
Walkerella
Walkerella is a genus of non-pollinating fig wasps in the family Pteromalidae, subfamily Otitesellinae. It is the most widely distributed genus within its subfamily, with species described from the Old World. Members are associated with Ficus fig trees, specifically species in subsection Conosycea of section Urostigma. Two new species were recently described from Xishuangbanna, China.
Xanthoteras
Xanthoteras is a genus of gall-inducing wasps in the family Cynipidae, first described by William Harris Ashmead in 1897. The genus includes species that induce galls on oak trees (Quercus). The ball-tipped gall wasp, a notable member, was transferred to this genus from Trigonaspis. These wasps are part of the diverse community of cynipids that manipulate plant tissue to form protective structures for their developing larvae.
Xanthoteras eburneum
Xanthoteras eburneum is a species of gall wasp in the family Cynipidae. It is one of several North American species in the genus Xanthoteras, which induce distinctive galls on oak trees (Quercus spp.). The species is primarily known from observations of its characteristic galls rather than the adult wasps themselves.
Xanthoteras teres
Ball-tipped Gall Wasp
Xanthoteras teres is a cynipid gall wasp native to western North America. The species induces distinctive galls on oak leaves, characterized by an upright stalk topped with a fuzzy spherical structure. Formerly classified under the genus Trigonapsis, it was reclassified to Xanthoteras based on taxonomic revision. The wasp has been documented primarily in Oregon and California.
Zapatella
Zapatella is a genus of gall wasps in the family Cynipidae. The genus includes at least one well-documented species, Z. davisae, which has become a significant pest of black oak (Quercus velutina) in the northeastern United States. Gall wasps in this genus induce gall formation on oak hosts. The genus was described in 2012 by Pujade-Villar & Melika.
Zapatella davisae
black oak gall wasp
Zapatella davisae is a cynipid gall wasp (Hymenoptera: Cynipidae) described in 2016 that has become a significant pest of black oak (Quercus velutina) in the northeastern United States. Larvae develop under bark in late summer, with adults emerging in early spring. The species has undergone notable population outbreaks on Long Island, NY (first documented 1990), Cape Cod and Martha's Vineyard, MA (2008), and more recently in coastal Rhode Island and Nantucket, MA. High parasitism rates by Sycophila species can lead to rapid population collapse, as observed on Long Island in 2016.
Zapatella quercusmedullae
Oak Rough Bulletgall Wasp
Zapatella quercusmedullae is a gall wasp in the family Cynipidae that induces galls on oak trees (Quercus). The species was transferred from Andricus to Zapatella based on phylogenetic studies. It produces distinctive bullet-shaped galls on twigs and branches of host oaks. The wasp has an alternating sexual and asexual generation life cycle typical of many Cynipini.
Zapatella quercusphellos
Zapatella quercusphellos is a species of gall wasp in the family Cynipidae. It is known to induce galls on oaks (Quercus species). The species was described by Carl Robert Osten Sacken and has been documented in the northeastern United States. It belongs to a genus of cynipid wasps specialized on oak hosts.
Zopheroteras compressum
Zopheroteras compressum is a species of gall wasp in the family Cynipidae, originally described as Acraspis compressus by Gillette in 1891. Like other members of its family, it is an obligate plant parasite that induces gall formation on host plants. The species has been documented through 202 iNaturalist observations, indicating it is not uncommon but remains poorly studied in the scientific literature. As a cynipid wasp, it likely exhibits the typical life history of alternating sexual and asexual generations, though specific details for this species are not well documented.
Zopheroteras sphaerula
Zopheroteras sphaerula is a species of gall wasp in the family Cynipidae, described by Weld in 1926. The genus Zopheroteras is part of the diverse cynipid radiation associated with oak galls. Like other members of its family, this species is presumed to be a specialist on oaks (Quercus), though specific host associations for this particular species require confirmation. The species has been documented through numerous iNaturalist observations, indicating it is not rare in suitable habitats.