Endemic
Guides
Sclerobunus nondimorphicus
Sclerobunus nondimorphicus is a species of armoured harvestman (order Opiliones, suborder Laniatores) described by Briggs in 1971. It belongs to the family Paranonychidae, a group of small to medium-sized harvestmen characterized by heavy body armouring. The species is known from the Pacific Northwest region of North America.
Scleropogon huachucanus
Scleropogon huachucanus is a species of robber fly (family Asilidae) described by Hardy in 1942. The specific epithet references the Huachuca Mountains of Arizona, suggesting a southwestern U.S. distribution. Like other Asilidae, this species is presumed to be a predatory fly, though specific behavioral and ecological details remain poorly documented. The genus Scleropogon is characterized by distinctive morphological features including a bearded face and elongated body form.
Scolops californicus
California Dictyopharid Planthopper
Scolops californicus is a planthopper species in the family Dictyopharidae, endemic to California. It belongs to a genus characterized by distinctive head morphology with an extended, often bizarrely modified frons. The species is part of the diverse planthopper fauna of western North America, with records primarily from California. As a member of Hemiptera, it possesses piercing-sucking mouthparts adapted for feeding on plant vascular tissues.
Scopula aemulata
Imitator Wave, Rival Wave
Scopula aemulata is a small moth in the family Geometridae, commonly known as the imitator wave or rival wave. The species was described by George Duryea Hulst in 1896 and has a restricted distribution in the southeastern United States. It is one of approximately 237 observations recorded on iNaturalist, indicating it is encountered with moderate frequency by citizen scientists and naturalists.
Selenophorus fabricii
Selenophorus fabricii is a species of ground beetle in the family Carabidae, described in 2017. The species is known from the Caribbean region, including the Cayman Islands, Hispaniola, and the Lesser Antilles. As a member of the genus Selenophorus, it belongs to a diverse group of small to medium-sized ground beetles commonly found in tropical and subtropical regions. Very little is known about the specific biology or ecology of this recently described species.
Selonodon compositus
Selonodon compositus is a species of click beetle in the family Cebrionidae, endemic to the southern United States. The species was originally described by Fall in 1934 and has undergone taxonomic revision, with three former species (S. emarginatus, S. knausi, and S. bruesi) now synonymized under this name. As a member of the genus Selonodon, it belongs to a group of 25 recognized species within this North American genus.
Selonodon floridensis
Selonodon floridensis is a click beetle species in the family Cebrionidae, described by Galley in 1999 from Florida. It is one of 17 new species described from the southern United States in a comprehensive revision of the genus Selonodon, which recognized 25 species total. The genus belongs to the click beetle superfamily Elateroidea and exhibits a trend of phyletic size increase across its range.
Sericomyia carolinensis
Two-spotted Pond Fly
Sericomyia carolinensis is a rare hoverfly (Syrphidae) endemic to the southeastern United States. Adults are distinguished by an all-yellow face, paired narrow yellow spots on the abdomen, and a yellow pilose scutellum. The species has been observed feeding on pear blossoms. Though the larval stage remains undocumented, it likely possesses the 'rat-tailed' morphology characteristic of the genus, inhabiting nutrient-rich aquatic environments such as bog mat pools.
Serranillus
Serranillus is a genus of small ground beetles in the family Carabidae. The genus was established by Barr in 1995 and contains three described species, all endemic to the United States. These beetles belong to the subtribe Anillina, a group characterized by reduced eyes and elongate bodies adapted to subterranean or soil-dwelling habits. The genus is relatively poorly known, with limited ecological and biological data published.
Shotwellia
Shotwellia is a monotypic genus of band-winged grasshoppers in the family Acrididae. The sole described species, Shotwellia isleta, is a rare North American grasshopper known from limited locations in the Chihuahuan Desert. It has historically been difficult to classify phylogenetically due to its unique morphological characteristics. The genus is currently placed within the Chortophaga genus group based on molecular and morphological evidence.
Sigmocheir
Sigmocheir is a genus of flat-backed millipedes in the family Xystodesmidae, endemic to the foothills of the Sierra Nevada mountains in California. The genus contains three species: S. maculifer, S. calaveras, and S. furcata. All species share a distinctive color pattern of three lemon yellow spots per segment. The genus was established by R.V. Chamberlin in 1951, with a complex taxonomic history involving synonymies that were not fully resolved until 1995.
Sigmoria ainsliei
Sigmoria ainsliei is a large flat-backed millipede in the family Xystodesmidae, endemic to a narrow region of eastern Tennessee. First described in 1921, it is one of the largest xystodesmid millipedes in North America, with adults exceeding 50 mm in length. The species has a highly restricted distribution spanning approximately 50 km from the Great Smoky Mountains National Park to Knoxville.
Sigmoria nantahalae
Nantahala Cherry Millipede
Sigmoria nantahalae is a species of flat-backed millipede in the family Xystodesmidae, first described by Hoffman in 1958. It is endemic to the southern Appalachian Mountains of North America, with confirmed records from North Carolina. The species belongs to the tribe Apheloriini, a group known for producing hydrogen cyanide as a defensive secretion.
Simplicia
Simplicia is a genus of monocotyledonous plants in the grass family (Poaceae), first described by Thomas Kirk in 1896. The genus is accepted in botanical nomenclature and is classified within the order Poales. It is native to New Zealand, where it represents a distinctive element of the native grass flora.
Siro sonoma
Siro sonoma is a species of mite harvestman in the family Sironidae. It was originally described by Shear in 1980 and placed in the genus Siro. In 2022, the genus Arhesiro was erected to accommodate this species and Siro clousi, with S. sonoma becoming Arhesiro sonoma. As of 2023, taxonomic sources vary in their treatment, with some recognizing Arhesiro sonoma as the accepted name while others retain Siro sonoma. The species is known only from Sonoma County, California.
Sisyracera inabsconsalis
A small crambid moth described by Möschler in 1890. Records indicate it occurs only on Puerto Rico and Cuba. The genus Sisyracera contains relatively few described species, and this taxon appears to be one of the less frequently encountered members of the group.
Sitalcina
Sitalcina is a genus of armoured harvestmen (Opiliones) in the family Phalangodidae, established by Nathan Banks in 1911. The genus contains approximately 10 described species, all endemic to western North America, primarily distributed in California and adjacent regions. These harvestmen belong to the suborder Laniatores, a group characterized by relatively short legs and heavily sclerotized body armor. Most species have been described from cave or deep-soil habitats, reflecting a troglomorphic or endogean lifestyle.
Sminthurus mencenbergae
Sminthurus mencenbergae is a springtail species endemic to the east coast of North America, first described by Snider in 1983. It belongs to the family Sminthuridae, a group of globular springtails characterized by their rounded body shape and distinctive color patterning. The species exhibits characteristic dorsal markings that aid in identification.
Socalchemmis shantzi
Socalchemmis shantzi is a species of spider in the family Zoropsidae, commonly referred to as false wolf spiders and wandering spiders. The species was described by Platnick and Ubick in 2001. It is endemic to the United States, specifically known from California. The genus Socalchemmis was established as a new genus in the same 2001 revision, representing a previously unrecognized group of North American spiders.
Soliperla sierra
Soliperla sierra is a stonefly species in the family Peltoperlidae, first described by Stark in 1983. The species is notable for its vibrational communication system, with male call and response signals described for the first time in a 2023 study. Males produce repeated monophasic call signals consisting of one to six signals with inconsistent inter-call intervals. Females respond with sequenced monophasic answers that do not follow all male calls, and occasional three-way exchanges occur when males respond to female answers. The species is known from California.
Somatochlora calverti
Calvert's Emerald, Andre Flies
Somatochlora calverti is a rare, range-restricted dragonfly in the family Corduliidae. It is endemic to the southeastern United States, primarily known from the Florida panhandle with scattered records in Alabama, Georgia, and South Carolina. The species is a habitat specialist associated with shallow seepage streams and steephead ravines. Its nymph has never been collected in the field, and its reproductive microhabitat remains speculative based on analogy to sympatric congeners.
Somatochlora margarita
Texas emerald
Somatochlora margarita, the Texas emerald, is a dragonfly endemic to the United States with a highly restricted range in eastern Texas and western Louisiana. It inhabits river systems and is rarely observed due to its habit of perching in forest canopies. The species was described by Donnelly in 1962 and is considered one of the more poorly known North American emeralds.
Somatochlora septentrionalis
Muskeg Emerald
Somatochlora septentrionalis, known as the muskeg emerald, is a dragonfly species in the family Corduliidae endemic to Canada. Adults are medium-sized, measuring 39–48 mm in length, with metallic green, brown, and black coloration. The species inhabits open fens with pools of open water, where males patrol and females oviposit in water and floating vegetation. Adults are active from June to August. It is nearly identical to Somatochlora whitehousei, requiring examination of male cerci and female subgenital plate for definitive identification.
Sosippus placidus
Lake Placid Funnel Wolf Spider
Sosippus placidus is a web-building wolf spider in the family Lycosidae, endemic to the United States. Unlike most wolf spiders, which are nomadic hunters, this species constructs thick sheet-like webs with funnel-like retreats. It is described as a habitat specialist compared to its widespread congener Sosippus floridanus. The species is known from relatively few observations (31 records on iNaturalist as of source date), suggesting restricted distribution or specific habitat requirements.
Spanglerogyrus
Spanglerogyrus is a monotypic genus of whirligig beetles (family Gyrinidae) containing the single species S. albiventris. It represents the only living member of the subfamily Spanglerogyrinae, the earliest diverging extant lineage of gyrinid beetles. The genus was described in 1979 from specimens collected in southern Alabama, making it a rare example of a relictual aquatic beetle lineage in North America.
Speleomaster
Speleomaster is a genus of armoured harvestmen (Opiliones: Laniatores) described by Briggs in 1974. The genus contains at least two species, both endemic to subterranean lava tube environments in southern Idaho's Snake River Plain. As members of the family Cryptomastridae, these harvestmen represent a specialized lineage adapted to cave life.
Speleonychia sengeri
Speleonychia sengeri is a troglobitic harvestman (order Opiliones) described by Briggs in 1974. The genus Speleonychia is endemic to cave systems in western North America. As a member of the family Cladonychiidae, this species belongs to a group of small, eyeless, pale-colored arachnids adapted to subterranean environments. Very few observations exist—only five records on iNaturalist—reflecting its restricted habitat and cryptic lifestyle.
Speyeria callippe comstocki
Comstock's Callippe Fritillary
Speyeria callippe comstocki is a subspecies of the Callippe fritillary butterfly, endemic to the Canadian prairies. It belongs to the greater fritillary group (genus Speyeria), characterized by silvery spots on the wing undersides and obligate associations with violet host plants. The subspecies is distinguished from other S. callippe populations by geographic isolation in the northern Great Plains. Like other Speyeria species, adults are strong fliers that rely primarily on vision for habitat navigation.
Speyeria carolae
Carole's fritillary
Speyeria carolae, known as Carole's fritillary, is a brush-footed butterfly endemic to the Charleston Mountains of Clark County, Nevada. Described in 1942 by dos Passos and Grey, this species is notable for its extremely restricted range, having been recorded only from this single mountain range. Adults fly from mid-June to September in a single annual generation. The species is closely associated with Viola charlestonensis, which serves as the sole known larval host plant.
Stagmomantis californica
California mantis, California mantid
Stagmomantis californica is a native California praying mantis and one of approximately 20 native mantis species in the United States. It is one of three native mantid species in California, alongside Stagmomantis limbata and Litaneutria minor. The species is characterized by dark bands on abdominal tergites, a potentially diagnostic feature when examined with male genitalia. Basic biological information including morphology, biometry, life history, and ecology remains incomplete for this species.
Stegea mexicana
Stegea mexicana is a moth in the family Crambidae, described by Munroe in 1964. It is known from a single locality in Veracruz, Mexico. Very little is documented about this species beyond its original description and basic taxonomic placement.
Stenogomphurus consanguis
Cherokee Clubtail
Stenogomphurus consanguis, commonly known as the Cherokee clubtail, is a species of clubtail dragonfly endemic to the southeastern United States. It inhabits small rivers and streams. The species is currently listed as Least Concern by the IUCN, though it has experienced significant shifts in conservation status over recent decades, previously being assessed as endangered in 2007. Approximately 30 known populations exist, most with fewer than 50 adults.
Stenomorpha confluens
Stenomorpha confluens is a flightless darkling beetle (family Tenebrionidae, tribe Asidini) native to Mexico. The species belongs to a genus that historically included 88 species with limited modern taxonomic revision until recent work by Fran Keller, who recognized 51 valid species and established seven new subgenera. Stenomorpha confluens is part of a group of flightless species restricted to the Sierra Transvolcanica or southern Transverse range in Mexico.
Sticthippus
Sticthippus is a genus of band-winged grasshoppers in the family Acrididae, established by Scudder in 1892. The genus contains a single described species, S. californicus. It belongs to the tribe Hippiscini within the subfamily Oedipodinae. The genus is endemic to California.
Stictiella villegasi
Algodones Sand Wasp
Stictiella villegasi, commonly known as the Algodones Sand Wasp, is a species of sand wasp in the family Crabronidae (formerly Bembicidae). It is endemic to the Algodones Dunes in North America, indicating a highly restricted geographic range. The species was described by R. Bohart in 1982.
Stictocephala stimulea
Stictocephala stimulea is a treehopper species in the family Membracidae, first described by Van Duzee in 1914. Treehoppers in this genus are characterized by their enlarged, often ornate pronotum that extends over the body like a helmet or thorn. The specific epithet 'stimulea' suggests possible resemblance to or association with stinging organisms, though this refers to appearance rather than actual sting. Like other membracids, this species likely feeds on plant phloem sap using piercing-sucking mouthparts.
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braunella
Catalina Cherry Leaf Miner
Stigmella braunella is a micro-moth in the family Nepticulidae, endemic to California. First described by W.W. Jones in 1933, this species has a wingspan of 5.4–6.6 mm and completes two generations per year. The larvae are leaf miners on Prunus ilicifolia (Catalina cherry), feeding within the leaves of this host plant.
Stigmella ceanothi
Stigmella ceanothi is a pygmy moth in the family Nepticulidae, first described by Braun in 1910 under the name Nepticula ceanothi. The species is endemic to California, United States, where it develops as a leafminer on host plants in the genus Ceanothus. Like other Stigmella species, the larvae feed internally within leaves, creating characteristic mines. The moth belongs to a large genus of minute leafmining moths that are ecologically significant as specialists on particular host plant genera.
Stigmella longisacca
Stigmella longisacca is a pygmy moth in the family Nepticulidae, first described in 1982. It is endemic to California, where its larvae are leaf miners on Juglans species, particularly Juglans californica. The species has a wingspan of 3.2–4.4 mm and appears to have two to three generations per year.
Stonemyia velutina
velvety stonian horsefly, Volutine Stoneyian Tabanid Fly
Stonemyia velutina is a rare horsefly endemic to California, first described in 1892. The species was not observed between 1942 and 2023, leading to a premature declaration of extinction in 1996. Its 2023 rediscovery confirmed continued survival, though knowledge of its biology remains limited. The species belongs to the family Tabanidae, a group of biting flies whose females typically require blood meals for egg development.
Stratiomydas
Stratiomydas is a genus of mydas flies (family Mydidae) established in 1989. The genus contains four described species distributed in Mexico, with records from Colima and Nayarit states. Species in this genus were previously classified under other genera before being separated based on morphological characteristics. Like other mydid flies, members are likely large-bodied and robust, though specific ecological data remain limited.
Strigoderma knausi
A small scarab beetle endemic to the Great Plains sand dune systems of south-central Kansas, Oklahoma, and northern Texas. Males are frequently encountered by sweeping low vegetation or at light traps, where they adopt a characteristic horizontal posture with forelegs extended and antennal clubs spread to detect female sex pheromones. The species was described from specimens collected by Warren Knaus in 1923–1925 at the type locality near Medora, Kansas, and named in his honor. Females are extraordinarily rare in collections, with only a single specimen among nearly 100 examined by Bader (1992).
Strumigenys chiricahua
Strumigenys chiricahua is a species of trap-jaw ant described by Ward in 1988. The specific epithet references the Chiricahua Mountains, a mountain range in southeastern Arizona known for high endemism and biodiversity. The species belongs to a large genus of miniature predatory ants characterized by specialized mandible morphology. Like other members of Strumigenys, it likely possesses elongated mandibles capable of rapid closure to capture prey.
Stygobromus
Stygobromus is a genus of subterranean freshwater amphipods in the family Crangonyctidae, comprising 134 described species. The genus is primarily distributed in North America, with a smaller number of species in the Palearctic region including Siberia. Many species are narrow endemics restricted to specific groundwater systems, and several are listed as endangered or vulnerable by the IUCN; one species, S. lucifugus, is extinct.
Stygobromus russelli
Russell's Cave Amphipod, Russell stygobromid
Stygobromus russelli is a subterranean amphipod species in the family Crangonyctidae, endemic to Texas in the United States. As a stygobiont, it inhabits groundwater systems in cave environments. The species was originally described by Holsinger in 1967 under the basionym Stygonectes russelli. Like other members of the genus Stygobromus, it exhibits troglomorphic adaptations including reduced pigmentation and eye development associated with life in permanent darkness.
Stygoparnus
Comal Springs dryopid beetle (for sole species S. comalensis)
Stygoparnus is a monotypic genus of dryopid beetles containing the single species Stygoparnus comalensis, known as the Comal Springs dryopid beetle. The genus is endemic to central Texas spring systems fed by the Edwards Aquifer. Its sole member is a federally endangered, subterranean-obligate aquatic beetle with vestigial eyes and reduced pigmentation. The genus represents the only known aquatic lineage within the family Dryopidae.
Styloniscidae
Styloniscidae is a family of terrestrial isopods (woodlice) within the suborder Oniscidea. The family comprises at least twelve genera distributed across tropical and subtropical regions, with exceptional diversity documented in Brazilian karst systems. Multiple genera contain troglobitic (exclusively cave-dwelling) species, including Xangoniscus, Cylindroniscus, Pectenoniscus, and Chaimowiczia. These cave-dwelling representatives exhibit classic troglomorphic traits: anophthalmy (eye loss), depigmentation, and elongated appendages.
Stylurus townesi
Townes' Clubtail, Townes's Clubtail
Stylurus townesi is a species of clubtail dragonfly endemic to the United States. It is a member of the family Gomphidae, a group characterized by their distinctive clubbed abdomens. The species is associated with riverine habitats, where it completes its life cycle. Like other members of the genus Stylurus, it is presumed to have aquatic nymphal stages and aerial adult stages typical of dragonflies.
Sumitrosis triplehorni
Sumitrosis triplehorni is a small leaf beetle in the family Chrysomelidae. It is known only from southern Florida and the Florida Keys, making it a geographically restricted species. Adults are among the smallest members of the genus, measuring under 3.5 mm in length.
Sunius
Sunius is a genus of rove beetles (Staphylinidae: Paederinae) in the Palaearctic region, currently comprising approximately 138 described species and two subspecies. The genus exhibits notable diversity in Turkey, where 25 species have been documented. Multiple species are micropterous (wingless) and appear to be locally endemic, particularly in Anatolia. Taxonomic revisions continue to expand the known fauna, with recent descriptions from Turkey, Iran, and Iraq.