Harpalinae
Guides
Amara aurata
Amara aurata is a species of ground beetle in the family Carabidae, first described by Dejean in 1828. It belongs to the large genus Amara, which contains numerous species distributed across North America and other regions. As a member of Carabidae, it is part of a diverse family of predatory beetles commonly known as ground beetles.
Athrostictus
Athrostictus is a genus of ground beetles in the family Carabidae, established by Henry Walter Bates in 1878. The genus comprises 18 described species. As members of the tribe Harpalini within the subfamily Harpalinae, these beetles are part of a diverse group of predominantly ground-dwelling predatory beetles.
Badister parviceps
Badister parviceps is a small ground beetle in the family Carabidae, described by Ball in 1959. The species is found in North America, with records from Canada and the United States. As a member of the genus Badister, it belongs to a group of predatory beetles commonly known as bombardier beetles or hunting beetles, though specific ecological details for this species remain poorly documented.
Bradycellus conformis
Conforming Harp Ground Beetle
Bradycellus conformis is a species of ground beetle in the family Carabidae, commonly known as the Conforming Harp Ground Beetle. It occurs in North America, with records from both Canada and the United States. As a member of the genus Bradycellus, it belongs to a group of small to medium-sized carabid beetles often associated with open, disturbed, or sandy habitats. The species was described by Fall in 1905.
Bradycellus congener
Congeneric Harp Ground Beetle
Bradycellus congener is a ground beetle species in the family Carabidae, first described by John Lawrence LeConte in 1847. It belongs to a genus of small to medium-sized carabid beetles commonly referred to as harp ground beetles. The species is distributed across North America, with records from both Canada and the United States. Like other members of Bradycellus, it is likely associated with open, dry habitats, though specific ecological details remain poorly documented.
Corticotomus depressus
Corticotomus depressus is a species of ground beetle in the family Carabidae. The genus Corticotomus is a small group within the tribe Harpalini, characterized by species adapted to particular microhabitats. Very little specific information is available about this particular species in the accessible literature. The species epithet "depressus" suggests a flattened body form, which is common among ground beetles that inhabit tight spaces under bark or in leaf litter.
Cratacanthus dubius
Dubious Harp Ground Beetle
Cratacanthus dubius is a species of ground beetle in the family Carabidae, and the sole member of the genus Cratacanthus. It is a moderately-sized beetle with a distinctive harp-like pronotal shape that gives the genus its name. The species has a broad distribution across North America, with records from both Canada and the United States. Its common name reflects both its taxonomic placement in the harp ground beetle lineage and the uncertainty that surrounded its early classification.
Cymindis americana
Cymindis americana is a species of ground beetle in the family Carabidae, subfamily Harpalinae. It was described by Pierre François Marie Auguste Dejean in 1826. The genus Cymindis comprises numerous species of small to medium-sized predatory beetles distributed primarily in the Holarctic region. As a member of the Harpalinae, this species likely exhibits the cursorial (running) locomotion and predatory habits characteristic of this diverse subfamily.
Cymindis arizonensis
Cymindis arizonensis is a species of ground beetle in the subfamily Harpalinae, described by Schaeffer in 1910. The genus Cymindis comprises numerous species of ground beetles distributed across the Northern Hemisphere. Like other members of Carabidae, this species is likely a nocturnal predator. Records indicate presence in the United States and North America, though detailed ecological information remains limited.
Cymindis borealis
Cymindis borealis is a species of ground beetle in the subfamily Harpalinae, described by John Lawrence LeConte in 1863. The genus Cymindis comprises small to medium-sized carabid beetles, many of which are nocturnal predators. Species in this genus are typically associated with open, dry habitats and are often found under stones or debris.
Cymindis californica
Cymindis californica is a species of ground beetle in the subfamily Harpalinae, described by George Henry Horn in 1895. It belongs to the genus Cymindis, a group of small to medium-sized carabid beetles commonly known as ant-loving beetles or simply cymindines. The species epithet indicates a California distribution.
Cymindis laticollis
A ground beetle species in the subfamily Harpalinae, described by Thomas Say in 1830. Records indicate presence in Canada and the United States. As a member of genus Cymindis, it likely shares the nocturnal predatory habits characteristic of this genus, though specific behavioral observations for this species remain limited.
Cymindis seriata
Cymindis seriata is a ground beetle species in the subfamily Harpalinae, described by Hatch in 1953. It belongs to the genus Cymindis, a diverse group of predatory beetles commonly known as hairy ground beetles. The species is known from the United States, though detailed biological information remains limited.
Cymindis vaporariorum
Cymindis vaporariorum is a species of ground beetle in the subfamily Harpalinae, first described by Carl Linnaeus in 1758. It belongs to the genus Cymindis, a group of predatory beetles within the family Carabidae. Adult activity peaks in June and July, though the species likely remains present throughout the summer months. Its specific epithet 'vaporariorum' suggests a historical association with greenhouses or steamy environments, though the precise origin of the name is not documented in available sources.
Diplocheila oregona
Diplocheila oregona is a species of ground beetle in the family Carabidae. It is found in western North America, ranging from Nevada and Utah northward to British Columbia and Manitoba. As a member of the genus Diplocheila, it belongs to a group of ground beetles adapted to moist riparian and shoreline habitats.
Discoderus congruens
Discoderus congruens is a species of ground beetle in the family Carabidae, described by Casey in 1914. It belongs to the genus Discoderus, a group of small to medium-sized carabids within the tribe Harpalini. The species is known from limited records in the United States, with few documented observations.
Discoderus pinguis
Discoderus pinguis is a ground beetle species in the family Carabidae, described by Thomas L. Casey in 1884. It belongs to the genus Discoderus, which is part of the tribe Harpalini within the subfamily Harpalinae. The species is known from the United States, with records indicating presence in North America. Very little specific information about its biology, ecology, or behavior has been documented in available sources.
Harpalini
ground beetles
Harpalini is a large tribe of ground beetles (family Carabidae, subfamily Harpalalinae) containing over 1,900 described species across approximately 103 genera. Members are predominantly nocturnal ground beetles distributed across all major biogeographic regions including the Palaearctic, Oriental, Afrotropical, Neotropical, Nearctic, and Australian regions. The tribe includes economically significant genera such as Harpalus, a diverse group with numerous Palaearctic species, and Trichotichnus, which comprises over 260 species worldwide. Taxonomic research on Harpalini focuses heavily on species descriptions, distribution records, and generic revisions, with recent work describing new species from China, Brazil, India, and Australia.
Harpalitae
Harpalitae was historically recognized as a supertribe of ground beetles within the subfamily Harpalinae of family Carabidae. Following taxonomic reorganization of Carabidae, the supertribe designation fell out of use. The four tribes formerly grouped under Harpalitae are now classified directly under Harpalinae. The term persists primarily in older literature.
Harpalus amputatus
Harpalus amputatus is a ground beetle species in the family Carabidae, described by Thomas Say in 1830. It belongs to the subfamily Harpalinae and is one of numerous species in the large genus Harpalus. The species exhibits a transcontinental distribution spanning northern Asia and North America, with recognized subspecies in Russia, China, Mongolia, and North America. Like other Harpalus species, it is a ground-dwelling beetle typically found in open, dry habitats where it hunts small invertebrates and scavenges opportunistically.
Harpalus atrichatus
Harpalus atrichatus is a species of ground beetle in the subfamily Harpalinae, described by Hatch in 1949. As a member of the large genus Harpalus, it belongs to the family Carabidae, one of the most diverse beetle families. The species epithet 'atrichatus' suggests a lack of hair or setae, though this has not been explicitly documented in the available sources. Very little specific information about its biology, ecology, or distribution has been published.
Harpalus cohni
Harpalus cohni is a species of ground beetle in the subfamily Harpalalinae, described by George E. Ball in 1972. It belongs to the large genus Harpalus, one of the most diverse genera within the Carabidae family. The species is known from the United States, with distribution records indicating presence in North America. Like other members of Harpalus, it is likely a nocturnal or crepuscular predator, though specific ecological details remain poorly documented due to limited study.
Harpalus compar
Harpalus compar is a species of ground beetle in the subfamily Harpalalinae, described by John Lawrence LeConte in 1848. It belongs to the large genus Harpalus, which contains over 400 described species and represents one of the most diverse genera within the family Carabidae. The species is known from North America, with records from both the United States and Canada.
Harpalus cordatus
Harpalus cordatus is a species of ground beetle in the family Carabidae. It is found in the United States, with records primarily from North America. The species has been historically placed in the genus Ophonus as O. (Metophonus) cordatus based on Duftschmid's 1812 description, but modern taxonomy recognizes it under Harpalus. Information on its biology and ecology is sparse, reflecting limited study of this particular species.
Harpalus cordifer
Harpalus cordifer is a species of ground beetle in the family Carabidae, subfamily Harpalalinae, described by Notman in 1919. It belongs to the large genus Harpalus, one of the most diverse genera of ground beetles in North America. The species is documented from Alaska and northern North America, with records extending into Canada and the contiguous United States. Like other members of the genus, it is a nocturnal ground beetle likely associated with open, disturbed habitats where it preys on small invertebrates and may consume seeds.
Harpalus ellipsis
Harpalus ellipsis is a species of ground beetle in the family Carabidae, subfamily Harpalinae. First described by John Lawrence LeConte in 1848, this beetle belongs to one of the largest genera of ground beetles in North America. The genus Harpalus contains numerous species that are primarily granivorous as adults, feeding on weed seeds, while larvae are predatory on soil-dwelling insects. Members of this genus are commonly found in agricultural and disturbed habitats across temperate regions.
Harpalus erythropus
Harpalus erythropus is a species of ground beetle in the subfamily Harpalinae, first described by Pierre François Marie Auguste Dejean in 1829. It belongs to the large and diverse genus Harpalus, which contains numerous species of ground beetles distributed across North America and other regions. Like other members of its genus, this species is likely adapted to ground-dwelling habits in various terrestrial habitats. The specific epithet "erythropus" refers to reddish legs, a characteristic feature of this species.
Harpalus faunus
Harpalus faunus is a species of ground beetle in the subfamily Harpalinae, first described by Thomas Say in 1823. It belongs to the large and diverse genus Harpalus, which contains numerous species across North America and other regions. Like other members of its genus, it is a ground-dwelling predator that inhabits open terrestrial environments. The species has been recorded from Canada and the United States, with observations documented in the northern parts of its range.
Harpalus fraternus
Harpalus fraternus is a species of ground beetle in the subfamily Harpalinae. It was described by John Lawrence LeConte in 1852. The species is native to western Canada and the United States. Like other members of the genus Harpalus, it is a ground-dwelling predator.
Harpalus fulvilabris
Harpalus fulvilabris is a species of ground beetle in the subfamily Harpalinae, described by Carl Gustaf Mannerheim in 1853. It belongs to the genus Harpalus, one of the most diverse genera of Carabidae with over 400 described species. The species is classified within the newly described subgenus Meroharpalus, established with H. fulvilabris as its type species. This subgenus is part of a broader systematic reorganization of the genus Harpalus based on adult morphological characters.
Harpalus fuscipalpis
Harpalus fuscipalpis is a species of ground beetle in the subfamily Harpalinae, described by Sturm in 1818. It belongs to the genus Harpalus, one of the largest genera of ground beetles. The species has a Holarctic distribution, with records from Alaska, Armenia, Austria, and Azerbaijan. As a member of the Carabidae family, it is likely a predatory beetle, though specific ecological details remain poorly documented.
Harpalus gravis
Harpalus gravis is a species of ground beetle in the subfamily Harpalinae, described by John Lawrence LeConte in 1858. It belongs to the large and diverse genus Harpalus, which contains numerous ground beetle species found across North America. Like other members of this genus, H. gravis is likely a nocturnal or crepuscular predator that forages on the ground surface. The specific epithet 'gravis' (Latin for 'heavy' or 'serious') may allude to some aspect of its physical characteristics or behavior.
Harpalus herbivagus
Harpalus herbivagus is a ground beetle species in the family Carabidae, subfamily Harpalinae. First described by Thomas Say in 1823, this species is native to North America with documented occurrence in Canada and the United States. Like other members of the genus Harpalus, it is a ground-dwelling beetle typically found in terrestrial habitats. The species name 'herbivagus' suggests herbivorous or plant-associated feeding habits, though specific ecological details remain poorly documented.
Harpalus indianus
Harpalus indianus is a ground beetle species in the subfamily Harpalalinae, described by Csiki in 1932. It belongs to the diverse genus Harpalus, which contains numerous species of ground beetles distributed across North America. The species has been recorded in Canada and the United States.
Harpalus innocuus
Harpalus innocuus is a species of ground beetle in the family Carabidae, subfamily Harpalinae. It was described by John Lawrence LeConte in 1863. The species belongs to the large genus Harpalus, which contains numerous ground beetle species distributed across North America and other regions. Like other members of its genus, it is a ground-dwelling predatory beetle.
Harpalus katiae
Harpalus katiae is a species of ground beetle in the subfamily Harpalinae, described by F. Battoni in 1985. It belongs to the large genus Harpalus, which contains numerous ground beetle species distributed across multiple continents. As with most members of this genus, it is likely a nocturnal predator inhabiting open, dry habitats, though specific details about its biology remain poorly documented.
Harpalus lewisii
Lewis' harpaline beetle
Harpalus lewisii is a ground beetle species in the subfamily Harpalinae, described by John Lawrence LeConte in 1865. It is a medium-sized carabid beetle measuring 11–15 mm in length. The species occurs in northeastern North America, with documented records from both Canada and the United States. As a member of the large genus Harpalus, it shares the general ground beetle morphology but is distinguished by specific characteristics that allow identification to species level.
Harpalus megacephalus
Harpalus megacephalus is a ground beetle species in the family Carabidae, subfamily Harpalalinae. It was described by John Lawrence LeConte in 1848. The species epithet "megacephalus" (Greek for "large-headed") likely refers to a relatively enlarged head capsule compared to congeners. As a member of the large genus Harpalus, it belongs to a group of primarily nocturnal, ground-dwelling beetles commonly found in open habitats across North America.
Harpalus obnixus
Harpalus obnixus is a species of ground beetle in the subfamily Harpalalinae, described by Thomas Lincoln Casey in 1924. As a member of the genus Harpalus, it belongs to one of the largest genera within the Carabidae family. The species is documented from North America, with records from both Canada and the United States. Like other ground beetles in this genus, it is presumed to be a ground-dwelling predator, though specific ecological details remain poorly documented in the available literature.
Harpalus opacipennis
Harpalus opacipennis is a species of ground beetle in the subfamily Harpalinae, described by Haldeman in 1843. It belongs to the large and diverse genus Harpalus, which contains numerous species of medium-sized ground beetles distributed across the Northern Hemisphere. The species name 'opacipennis' refers to the dull or opaque appearance of the elytra (wing covers). Like other members of Harpalus, this species is likely a generalist predator and scavenger, though specific ecological details remain poorly documented.
Harpalus providens
Harpalus providens is a species of ground beetle in the subfamily Harpalinae, described by Thomas L. Casey in 1914. As a member of the large genus Harpalus, it belongs to one of the most diverse groups of carabid beetles in North America. The species is known from limited occurrence records in Canada and the United States, with observations documented through citizen science platforms. Like other Harpalus species, it likely occupies open, dry habitats where it functions as an omnivorous ground-dwelling predator and seed-feeder.
Harpalus puncticeps
Harpalus puncticeps is a species of ground beetle in the family Carabidae, subfamily Harpalinae. The species was described by Thomas Lincoln Casey Jr. in 1914. Taxonomic records indicate some nomenclatural complexity, with the species also referenced under Stephens (1828) as Glanodes puncticeps in some databases. It is currently classified within the genus Harpalus, though some sources list it under the subgenus Ophonus (Metophonus).
Harpalus retractus
Harpalus retractus is a ground beetle species in the family Carabidae, subfamily Harpalinae, described by John Lawrence LeConte in 1863. It is recorded from Mexico and the United States. Like other members of the genus Harpalus, it is a nocturnal ground beetle. The species is part of a large genus containing over 400 species worldwide, with many species being granivorous as adults.
Harpalus rubripes
Red-legged Harpalus
Harpalus rubripes is a medium-sized ground beetle in the subfamily Harpalinae, characterized by its distinctive reddish legs that give the species its name. Native to the Palearctic region, it has established populations across much of Europe, extending eastward through Siberia and Central Asia to Anatolia. The species was introduced to North America, with records dating from 1987 onward. Like other members of the genus Harpalus, it is primarily nocturnal and ground-dwelling.
Harpalus somnulentus
Harpalus somnulentus is a ground beetle species in the family Carabidae, subfamily Harpalinae. It was first described by Pierre François Marie Auguste Dejean in 1829. The species belongs to the large genus Harpalus, which contains numerous ground beetle species distributed across North America and other regions. As a member of the Carabidae family, it is a predatory beetle likely active in terrestrial habitats.
Harpalus texanus
Harpalus texanus is a species of ground beetle in the subfamily Harpalinae, described by Casey in 1914. As a member of the large genus Harpalus, it belongs to the family Carabidae, one of the most diverse beetle families. The species is known from the United States, with records indicating presence in North America. Like other Harpalus species, it is likely a nocturnal ground-dwelling beetle, though specific ecological details remain poorly documented.
Harpalus ventralis
Harpalus ventralis is a ground beetle species in the family Carabidae, subfamily Harpalinae. It was described by John Lawrence LeConte in 1848. The species is native to North America, with records from Canada and the United States. Like other members of the genus Harpalus, it is a nocturnal, ground-dwelling beetle.
Hartonymus alternatus
Hartonymus alternatus is a ground beetle species in the family Carabidae, first described by LeConte in 1863. It is native to North America and has been recorded from the United States. As a member of the Harpalinae subfamily, it belongs to a diverse group of ground beetles. Very little specific biological information is available for this species.
Nomius
Nomius is a genus of ground beetles (Carabidae) in the tribe Psydrini, subfamily Psydrinae. The genus contains at least one well-documented species, Nomius pygmaeus, which exhibits a remarkable disjunct distribution across North America, Europe, and Asia. Members are characterized by their isolated phylogenetic position within the Harpalinae subfamily.
Notiobia mexicana
Notiobia mexicana is a species of ground beetle in the family Carabidae. It occurs across a broad geographic range from the southwestern United States through Mexico and Central America to Panama. As a member of the tribe Notiobiini, it belongs to a group of carabid beetles often associated with riparian and wetland habitats.
Oodini
Oodini is a tribe of ground beetles (Carabidae: Harpalinae) containing over 400 species in 43 genera, with highest diversity in tropical Africa. Members are generally associated with standing water habitats. The tribe is distinguished by two diagnostic morphological characters: a laterally coadunate metepisternum and an elytral ridge formed by fusion of elytral intervals 7 and 8.
Oxycrepis intercepta
Oxycrepis intercepta is a species of ground beetle in the family Carabidae. It occurs in North America, with records from Mexico and the United States. As a member of the tribe Harpalini, it belongs to a diverse group of predominantly ground-dwelling beetles. Published information on its biology and ecology remains limited.
Pelmatellus
Pelmatellus is a genus of ground beetles in the family Carabidae, established by Henry Walter Bates in 1882. The genus contains at least 20 described species and is the type genus of the tribe Pelmatellini within the subfamily Harpalinae. As with other ground beetles, species in this genus are likely predatory, though specific ecological details remain poorly documented.
Philodes rectangulus
Rectangular Harp Ground Beetle
Philodes rectangulus is a species of ground beetle in the family Carabidae, commonly known as the Rectangular Harp Ground Beetle. It is native to North America, with records from both Canada and the United States. The species was originally described as Acupalpus rectangulus by Chaudoir in 1868 and later transferred to the genus Philodes. It belongs to the subfamily Harpalinae, a diverse group of ground beetles often associated with open habitats.
Piosoma setosum
Piosoma setosum is a species of ground beetle in the family Carabidae, notable as the sole member of its monotypic genus. First described by LeConte in 1847, this beetle occurs in North America. The genus name derives from Greek, reflecting its distinctive setose (bristly) characteristics. Despite its taxonomic uniqueness, detailed biological information remains limited.
Pterostichini
Woodland Ground Beetles
Pterostichini is a tribe of ground beetles in the subfamily Harpalinae, commonly known as woodland ground beetles. The tribe contains approximately 180 genera and over 3,800 described species, though estimates vary. Members are primarily ground-dwelling beetles with diverse ecological roles as predators. Research on certain genera, particularly Pterostichus, has revealed complex life history patterns including iteroparity, flexible breeding periods, and responses to environmental conditions such as drought.
Pterostichus
Blackclock Ground Beetles
Pterostichus is a large genus of ground beetles (Carabidae) in the subfamily Harpalinae, comprising over 1,200 species with a Holarctic distribution. These beetles are primarily predatory and are commonly found in ground-level habitats, often beneath rocks and debris. The genus exhibits considerable ecological diversity, with species showing varied substrate preferences ranging from sandy soils to subterranean environments.
Selenophorus breviusculus
Selenophorus breviusculus is a ground beetle in the family Carabidae, described by George Henry Horn in 1880. It belongs to the genus Selenophorus, a group of nocturnal, soil-dwelling beetles commonly known as false bombardier beetles. The species is recorded from the United States.
Selenophorus chaparralus
Selenophorus chaparralus is a ground beetle species described by Purrington in 2000. It belongs to the genus Selenophorus, a group of nocturnal ground beetles within the family Carabidae. The species epithet 'chaparralus' suggests an association with chaparral vegetation, a type of Mediterranean scrubland found in parts of western North America. As a recently described species with limited observation records, much of its biology remains undocumented.
Selenophorus fabricii
Selenophorus fabricii is a species of ground beetle in the family Carabidae, described in 2017. The species is known from the Caribbean region, including the Cayman Islands, Hispaniola, and the Lesser Antilles. As a member of the genus Selenophorus, it belongs to a diverse group of small to medium-sized ground beetles commonly found in tropical and subtropical regions. Very little is known about the specific biology or ecology of this recently described species.
Selenophorus hylacis
Selenophorus hylacis is a species of ground beetle in the family Carabidae. It was described by Thomas Say in 1823. The species occurs in North America, with records from both the United States and Canada. Like other members of its genus, it is a nocturnal ground-dwelling beetle.
Selenophorus mexicanus
Selenophorus mexicanus is a ground beetle species in the family Carabidae, described by Putzeys in 1878. It belongs to the genus Selenophorus, a group of nocturnal carabid beetles commonly known as false bombardier beetles or night-stalking tiger beetles. The species is native to Central America and southern Mexico.
Stenocrepis tibialis
Stenocrepis tibialis is a ground beetle species in the family Carabidae, described by Chevrolat in 1834. It has a broad Neotropical distribution extending into the southern United States. The species occurs in diverse habitats ranging from Caribbean islands to South American mainland regions.
Stenolophina
seedcorn beetles (for genus Stenolophus)
Stenolophina is a subtribe of ground beetles in the family Carabidae, comprising approximately 9 genera and at least 50 described species. The subtribe includes the genus Stenolophus, commonly known as seedcorn beetles. Members are small to medium-sized carabids found in diverse terrestrial habitats.
Stenolophus binotatus
Stenolophus binotatus is a ground beetle species in the family Carabidae. It occurs in North America, with records from the United States. The species has been documented in blacklighting surveys in Kansas during early spring. It belongs to the genus Stenolophus, commonly known as seedcorn beetles.
Synuchus impunctatus
Synuchus impunctatus is a ground beetle species in the family Carabidae, described by Thomas Say in 1823. It belongs to the subfamily Platyninae (sometimes referenced as Harpalinae in older literature) and is part of the diverse genus Synuchus, which contains numerous similar-looking species. The species is documented from Canada and the United States.
Trichotichnus fulgens
Trichotichnus fulgens is a ground beetle species in the family Carabidae, originally described by Csiki in 1932. The species is currently considered a synonym of Trichotichnus nitidulus according to GBIF taxonomy. It is recorded from North America, specifically Canada and the United States. As a member of the tribe Harpalini, it belongs to a diverse group of ground beetles commonly found in terrestrial habitats. The genus Trichotichnus contains multiple species distributed across the Holarctic region.