Lotic
Guides
Heptagenioidea
Heptagenioidea is a superfamily of mayflies (Ephemeroptera) distributed across most global regions except the Arctic and Antarctic. The superfamily likely originated in the Jurassic period, coinciding with a shift from still-water to flowing-water breeding habitats. This ecological transition drove rapid radiation and morphological adaptations, including modifications to mouthparts and forelegs for passive feeding in current. Heptageniidae represents the most widely distributed family within this group.
Hesperoperla hoguei
Banded Stone
Hesperoperla hoguei is a species of stonefly in the family Perlidae, commonly known as the banded stone. It is a member of the subfamily Acroneuriinae, a group of predatory stoneflies often associated with flowing freshwater habitats. The species was described by Baumann and Stark in 1980 and is known from western North America, particularly California. As with other perlid stoneflies, it likely inhabits cool, well-oxygenated streams.
Hydropsyche betteni
Common Netspinner
Hydropsyche betteni is a species of netspinning caddisfly in the family Hydropsychidae, described by Ross in 1938. It is one of the most widespread and commonly encountered members of its genus in North America. Like other hydropsychids, the larvae construct fixed capture nets to filter food from flowing water. The species is well-documented in freshwater biomonitoring studies due to its abundance and sensitivity to water quality.
Hydropsyche incommoda
Hydropsyche incommoda is a species of net-spinning caddisfly in the family Hydropsychidae. Described by Hermann Hagen in 1861, this species is part of a large genus of aquatic insects whose larvae construct fixed capture nets in flowing water. Adults are nocturnal and attracted to light. The species is documented from North America.
Hydropsyche occidentalis
Hydropsyche occidentalis is a filter-feeding caddisfly species in the family Hydropsychidae, found in lotic (flowing water) streams of North America. It constructs fixed capture nets to filter organic particles from the water column, playing a significant role in stream ecosystem function. The species has been studied for its bioenergetics and contribution to controlling organic transport in streams, where it can comprise a substantial portion of invertebrate biomass in suitable habitats.
Hydropsyche scalaris
Hydropsyche scalaris is a species of net-spinning caddisfly in the family Hydropsychidae, first described by Hagen in 1861. It belongs to one of the largest genera of caddisflies, with larvae that construct fixed retreats and capture nets in flowing water to filter food particles. The species is documented from North America within the Nearctic region.
Hydropsyche slossonae
Hydropsyche slossonae is a caddisfly species in the family Hydropsychidae, known from freshwater lotic habitats in eastern and midwestern North America. The species is univoltine, with adults emerging synchronously from late May through early July. Larvae construct silk capture nets to filter suspended organic matter from flowing water and overwinter as fifth-instar larvae.
Hydropsychinae
net-spinning caddisflies
Hydropsychinae is a subfamily of caddisflies (Trichoptera) within the family Hydropsychidae, commonly known as net-spinning caddisflies. Larvae construct fixed capture nets in flowing water to filter food particles. The subfamily is taxonomically diverse, with generic revisions documented in multiple regions including New Zealand and the Americas.
Isogenoides
springflies
Isogenoides is a genus of springflies (family Perlodidae) comprising eight described species distributed across North America. These stoneflies inhabit cool, lotic freshwater systems and exhibit species-specific vibrational communication through drumming. The genus demonstrates considerable life history diversity, including extended diapause periods and semivoltine development in some populations.
Isonychia campestris
Isonychia campestris is a filter-feeding mayfly species described by McDunnough in 1931. The immature nymphs inhabit lotic aquatic systems where they employ specialized morphological structures to capture suspended particulate matter from the water column. This species has been studied specifically for its filter feeding mechanisms, which involve interception and diffusive deposition of particles smaller than the pore spaces between filtering structures. The species occurs in the Nearctic region of North America.
Isonychia rufa
Isonychia rufa is a brushlegged mayfly species in the family Isonychiidae, first described by McDunnough in 1931. It belongs to the bicolor group within its genus. The species is found in larger rivers and streams of the Mississippi drainage system in central North America.
Isoperla sobria
Colorado Stripetail
Isoperla sobria is a species of stonefly in the family Perlodidae, commonly known as the Colorado Stripetail. It is a member of the genus Isoperla, which comprises numerous small to medium-sized predatory stoneflies found in running waters of the Northern Hemisphere. The species has been documented across western North America from Alaska to California. Like other perlodids, its larvae develop in freshwater streams and are indicators of good water quality.
Isoperlinae
Stripetail Stoneflies, Springflies
Isoperlinae is a subfamily of stoneflies in the family Perlodidae, distributed across the Palaearctic and Nearctic regions. Members are commonly known as stripetails or springflies. The subfamily includes approximately 12 genera, with Isoperla being the most speciose and well-known. These stoneflies are aquatic insects whose nymphs inhabit running waters.
Lacunicambarus diogenes
devil crayfish, devil crawfish
Lacunicambarus diogenes, commonly known as the devil crayfish or devil crawfish, is a primary burrowing crayfish native to eastern North America. This species constructs and inhabits burrows in wet, muddy terrestrial habitats rather than living in permanent surface water. Its burrowing activities create refugia used by numerous other species, including documented use by eastern cicada killer wasps (Sphecius speciosus) as brooding habitat. The species ranges across the Atlantic Coastal Plain and Piedmont ecoregion from New Jersey to Georgia, with disjunct populations in Louisiana.
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cupida
Early Brown Spinner, Black Quill
Leptophlebia cupida is a pronggilled mayfly species native to North America, commonly known as the early brown spinner or black quill. The species exhibits a univoltine life cycle with egg diapause during summer months. Nymphs develop through approximately 20-34 instars over 10 months, with emergence occurring from late April to mid-May. Adults are short-lived, non-feeding, and mate in swarms near streams.
Leuctrinae
Leuctrinae is a subfamily of stoneflies (order Plecoptera) within the family Leuctridae. Members of this group are small to medium-sized stoneflies commonly known as 'rolled-winged stoneflies' due to their distinctive wing posture at rest. The subfamily was established by Klapálek in 1905 and represents one of the major lineages within the Nemouroidea superfamily. Leuctrinae species are primarily associated with cool, clean running waters and are widely distributed across the Holarctic region.
Liodessus
Liodessus is a genus of small diving beetles in the family Dytiscidae, tribe Bidessini. Species occur across the Americas from North America through the Andes to southern South America, with notable diversity in high-altitude wetlands and páramo ecosystems. Many species inhabit shallow, exposed pools and temporary water bodies, showing phenotypic plasticity in body form correlated with habitat permanence. The genus contains numerous species, with several new species described from the high Andes of Peru and Colombia in recent years. Taxonomic resolution relies heavily on male genital morphology.
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net tube caddisfly
Lype is a genus of net tube caddisflies in the family Psychomyiidae, containing more than 20 described species. Adults are small to medium-sized caddisflies with reduced mouthparts. Larvae construct fixed, silken retreat nets on submerged surfaces in flowing water. The genus has been studied for female identification and reproductive biology in Denmark.
Macronematinae
net-spinning caddisflies
Macronematinae is a subfamily of net-spinning caddisflies (Trichoptera: Hydropsychidae) established by Ulmer in 1905. Larvae construct silk webs to capture prey drifting in flowing water. The subfamily includes genera such as Macronema, Macrostemum, Leptonema, Leptopsyche, and the Neotropical endemic Centromacronema. Members occupy diverse freshwater habitats across multiple continents.
Macronychus
riffle beetles
Macronychus is a genus of riffle beetles in the family Elmidae, comprising approximately 11 described species. Members of this genus inhabit flowing water environments, particularly riffle zones of streams. The genus was established by Müller in 1806 and includes species distributed across North America, Europe, and Asia.
Macronychus glabratus
riffle beetle
Macronychus glabratus is a species of riffle beetle in the family Elmidae, described by Say in 1825. It inhabits lotic freshwater environments, specifically stream riffles in North America. Research has documented its life cycle, growth, survival, and production in streams of northwest Arkansas and southeast Texas.
Macrostemum
Glossy Wing Sedges
Macrostemum is a cosmopolitan genus of net-spinning caddisflies in the family Hydropsychidae, comprising at least 90 described species. Adults are recognized by distinctive dark and light contrasting wing patterns. Larvae construct silken retreats with capture nets to filter fine particulate organic matter from flowing water. The genus is particularly diverse in the Neotropical region, with species documented across Africa, Asia, Madagascar, and the Americas.
Megarcys subtruncata
Truncate Springfly
Megarcys subtruncata, commonly known as the truncate springfly, is a stonefly species in the family Perlodidae. It was described by Hanson in 1942. The species occurs in western North America, with documented records from British Columbia, California, Idaho, Montana, and Nevada. It belongs to a genus of predatory stoneflies typically associated with cold, well-oxygenated streams.
Metrobates artus
Metrobates artus is a water strider species in the family Gerridae, first described by Anderson in 1932. It belongs to the subfamily Trepobatinae, a group of gerrids adapted to life on flowing water surfaces. The genus Metrobates is primarily Nearctic in distribution. This species has been recorded from both Middle America and North America.
Metrobates hesperius
Metrobates hesperius is a water strider (family Gerridae) distributed across eastern North America and the Caribbean. It belongs to the subfamily Trepobatinae, a group of semiaquatic bugs that inhabit flowing water habitats. The species was described by Uhler in 1871 and includes three recognized subspecies with distinct geographic distributions.
Nebrioporus macronychus
Nebrioporus macronychus is a predaceous diving beetle in the family Dytiscidae. The species was described by Shirt & Angus in 1992. It belongs to the genus Nebrioporus, a group of small to medium-sized diving beetles primarily associated with running waters. The specific epithet 'macronychus' refers to enlarged claws, a distinctive morphological feature of this species.
Neophylax aniqua
Neophylax aniqua is a species of caddisfly (order Trichoptera) first described by Ross in 1947. The species occurs in North America, with documented populations in southern Ontario and Vermont. Larvae inhabit cool headwater sections of streams and construct protective cases using silk and gathered materials. Adults emerge in September and October following a summer prepupal diapause.
Neureclipsis
tube maker caddisflies
Neureclipsis is a genus of net-spinning caddisflies in the family Polycentropodidae, containing at least eight described species distributed across the Northern Hemisphere and Australia. Larvae construct elaborate silken catchnets to capture drifting invertebrate prey in lotic freshwater habitats. The genus includes the well-studied species N. bimaculata, whose life history and feeding ecology have been investigated in North American and European populations.
Neurocordulia alabamensis
Alabama Shadowdragon
Neurocordulia alabamensis, the Alabama Shadowdragon, is a dragonfly species in the family Corduliidae described in 1955. It belongs to a genus of shadowdragons known for crepuscular flight activity. The species is endemic to the southeastern United States, with documented populations in Alabama and adjacent regions. Like other Neurocordulia species, it is associated with riverine habitats and exhibits distinctive flight behavior during low-light conditions.
Octogomphus specularis
Grappletail, Club-tailed dragonfly
Octogomphus specularis, commonly known as the grappletail or club-tailed dragonfly, is the sole species in its genus. It is a North American dragonfly distinguished by a club-like widening at the end of its abdomen. Larvae are specialized to lake outlet streams, where warmer thermal conditions support development at the northern range limit. The species has a three-year life cycle and reaches its northernmost distribution in British Columbia, Canada.
Oplonaeschna
Riffle Darners
Oplonaeschna is a genus of dragonflies in the family Aeshnidae, commonly known as riffle darners. The genus comprises approximately five described species, including both extant and fossil taxa. The type species, Oplonaeschna armata, has been the subject of limited ecological study. The genus was established by Selys in 1883.
Oreodytes snoqualmie
Oreodytes snoqualmie is a species of predaceous diving beetle in the family Dytiscidae. It was described by Hatch in 1933 and was originally placed in the genus Hydroporus. The species is known from western North America, with records from Alberta and British Columbia in Canada. As a member of the genus Oreodytes, it belongs to a group of small diving beetles typically associated with lotic (flowing water) habitats.
Ostrocerca prolongata
Bent Forestfly
Ostrocerca prolongata, commonly known as the bent forestfly, is a species of spring stonefly in the family Nemouridae. It was described by Claassen in 1923. The species is found in eastern North America, with records from the northeastern United States and adjacent Canada. Like other members of Nemouridae, it is associated with forested stream habitats.
Oulimnius
Oulimnius is a genus of riffle beetles in the family Elmidae, containing approximately 15 described species. These beetles are aquatic and inhabit running water environments. The genus has been the subject of molecular phylogenetic studies, particularly for the O. tuberculatus species group, which revealed cryptic diversity and supported the re-validation of O. perezi as a distinct species.
Parachaetocladius
non-biting midges
Parachaetocladius is a genus of non-biting midges in the subfamily Orthocladiinae of the family Chironomidae, established by Wolfgang Wülker in 1959 and revised in 2020. The genus contains approximately 10 described species distributed across the Holarctic region, including both Palaearctic and Nearctic representatives. Species are typically associated with lotic freshwater habitats, particularly sandy substrates in springs and groundwater-fed streams. The genus is distinguished from related orthocladiine genera primarily through adult male genitalic characters, particularly features of the hypopygium.
Parapsyche apicalis
Parapsyche apicalis is a species of net-spinning caddisfly in the family Hydropsychidae, originally described as Arctopsyche apicalis by Banks in 1908. The species belongs to the subfamily Arctopsychinae and is found in North America. As a member of Hydropsychidae, larvae construct fixed retreats and capture nets to filter food particles from flowing water.
Petrophila confusalis
confusing petrophila moth
Petrophila confusalis is a semi-aquatic moth in the family Crambidae, commonly known as the confusing petrophila moth. Adults are small, with forewing lengths of 5–11 mm, and are active from April to September. The species is notable for its specialized aquatic larval stage, during which caterpillars feed on algae and diatoms in fast-moving streams. The moth serves as a host for the parasitic aquatic wasp Tanychela pilosa and has been observed in large grazing zones that contribute to epilithic spatial patchiness. In 2021, it was listed on the Nevada Division of Natural Heritage watch list due to population decline.
Podmosta
spring stoneflies
Podmosta is a genus of spring stoneflies in the family Nemouridae, comprising approximately six described species distributed across northern North America and the Russian Far East. The genus is notable for its adaptation to cold climates, including documented embryonic diapause in at least one species. Adults are active in spring, with nymphal development occurring primarily during winter months.
Polycentropus
tube maker caddisflies
Polycentropus is a large genus of caddisflies (Trichoptera) in the family Polycentropodidae, containing more than 190 described species. The genus is commonly referred to as 'tube maker caddisflies' due to the silken retreat structures constructed by larvae. Species occur across multiple continents including Europe, Asia, and South America. Larvae are predatory and inhabit running waters, where they build silken retreats with capture nets to trap prey.
Potamanthidae
Hackle-gilled Burrower Mayflies
Potamanthidae is a family of burrowing mayflies comprising approximately 23 species across three to four genera (Anthopotamus, Potamanthus, Rhoenanthus, and Stygifloris). Larvae are fossorial, inhabiting interstitial spaces in gravel and pebble substrates of streams and rivers, and possess distinctive mandibular tusks used for excavation and defense. Adults are aerial and short-lived. The family has a disjunct distribution spanning North America and East Asia.
Prosimulium mixtum
Mixed-up Black Fly
Prosimulium mixtum is a species of black fly in the family Simuliidae, commonly known as the mixed-up black fly. It is native to North America and has been studied in the context of spring snowmelt environments and physiological responses to environmental stressors. The species is part of the Prosimulium fuscum/mixtum complex, which has historically caused taxonomic confusion. Adults are small blood-feeding flies, with females requiring blood meals for egg development.
Psephenus herricki
Herrick's Water Penny Beetle
Psephenus herricki is a water penny beetle found in eastern North America. Larvae are highly flattened and inhabit stream riffles, where they occupy interstitial crevices and undersides of stones. The species exhibits specific behavioral and morphological adaptations to flowing water environments, including thigmokinesis that inhibits movement when wedged in substrate. Adults are associated with wave-splashed rocks that serve as mating sites and oviposition substrates.
Pseudiron centralis
White Sand-river Mayfly
Pseudiron centralis is a predaceous mayfly species endemic to sandy freshwater river habitats in North America. It exhibits a univoltine summer life cycle with an extended egg stage lasting 9–10 months. Larvae are active epibenthic predators specialized for feeding on psammophilous chironomid larvae, with distinct microhabitat shifts during development: early instars occupy marginal sands, while later instars inhabit shifting sand dunes in main channel areas. The species shows low population density but also low mortality, with random dispersion patterns in mature larvae.
Pteronarcys
Giant Stoneflies, Salmonflies
Pteronarcys is a genus of giant stoneflies in the family Pteronarcyidae, commonly known as salmonflies. The genus comprises approximately 8 described species distributed across North America and Far Eastern Russia. These are among the largest stoneflies, with nymphs reaching substantial sizes in lotic freshwater habitats. Life cycles are notably long, ranging from 1 to 5 years depending on species and thermal conditions, with multiple larval diapause stages and temperature-dependent egg development documented in several species.
Rhagovelia becki
Rhagovelia becki is a species of small water strider in the family Veliidae, described by Drake and Harris in 1936. Like other members of the genus Rhagovelia, this species is adapted to life on the water surface, utilizing the middle and hind legs for propulsion. The species has been recorded from Middle America and North America, though detailed ecological studies remain limited. It belongs to a diverse genus containing over 100 described species, many of which occupy specialized microhabitats along streams and rivers.
Rhagovelia choreutes
Rhagovelia choreutes is a species of small water strider in the family Veliidae. It belongs to the infraorder Gerromorpha, a group of semiaquatic true bugs adapted to life on the water surface. The species was described by Hussey in 1925. It occurs in running water habitats in parts of North and Central America.
Rhagovelia obesa
riffle bug
Rhagovelia obesa is a small aquatic true bug in the family Veliidae, commonly known as a riffle bug or smaller water strider. It inhabits flowing water environments where it uses a unique swimming plume to move underwater. The species has an unusual life history among aquatic Hemiptera, with four nymphal instars rather than five and eggs as the overwintering stage. It has been successfully reared in laboratory conditions from first instar to adult.
Rheopelopia
Rheopelopia is a genus of non-biting midges in the family Chironomidae, subfamily Tanypodinae. Members are aquatic in their larval stage and belong to the tribe Pentaneurini. The genus was established by Fittkau in 1962. Larvae are generally found in running water habitats.
Rhyacophila acutiloba
Rhyacophila acutiloba is a species of free-living caddisfly in the family Rhyacophilidae, first described by Morse & Ross in 1971. As a member of the genus Rhyacophila, it belongs to a group commonly known as "caseless caddis" or "free-living caddisflies" whose larvae do not construct portable cases. The species is known from North America, with records from Vermont and other parts of the Nearctic region. Like other rhyacophilids, it is associated with lotic (flowing water) habitats.
Sericostomatoidea
Sericostomatoidea is a superfamily of caddisflies (Trichoptera) containing approximately 13 families. The superfamily includes diverse case-making caddisflies, notably the Helicopsychidae (snail-case caddisflies) and the Australasian endemic family Conoesucidae. Molecular and morphological studies have confirmed the monophyly of at least some constituent families, though relationships among families remain under investigation.