Cynipidae
Guides
Disholandricus lasius
Disholandricus lasius is a cynipid gall wasp species that induces galls on oak hosts. Formerly classified in the genus Andricus, this species was reassigned to Disholandricus based on phylogenetic and morphological studies. Like other members of the Cynipidae family, the wasp manipulates host plant tissue to form a protective structure that houses and nourishes its developing larvae. The species exhibits the complex life cycle typical of oak gall wasps, involving both sexual and asexual generations that alternate between different plant structures or host species.
Disholcaspis
Disholcaspis is a genus of gall wasps in the family Cynipidae containing more than 40 described species. Species in this genus induce characteristic galls on oak trees (Quercus spp.), with gall morphology varying by species and generation. Some species produce galls that secrete honeydew, establishing facultative mutualisms with ants that defend the galls from parasitoids. The genus exhibits heterogony, with alternating sexual and parthenogenetic asexual generations that typically produce different gall forms.
Disholcaspis bassetti
Disholcaspis bassetti is a gall wasp species in the family Cynipidae, a group known for inducing distinctive plant growths. Like other members of the genus Disholcaspis, this species induces galls on oak species (Quercus). The genus is characterized by alternation of generations between sexual and asexual (agamic) forms, with the latter producing the more conspicuous galls. Specific details about D. bassetti's host associations and gall morphology are not well-documented in available sources.
Disholcaspis canescens
Round Honeydew Gall Wasp
Disholcaspis canescens is a species of gall wasp in the family Cynipidae, known for inducing distinctive galls on oak trees. Like other members of the genus Disholcaspis, this species produces round, detachable galls on oak twigs. The genus is notable for having eliminated males from the life cycle in all known species, with females reproducing parthenogenetically and producing only female offspring. The specific epithet 'canescens' refers to a grayish or hoary appearance, though the precise application to this species' galls or wasps is not detailed in available sources.
Disholcaspis cinerosa
Mealy oak gall wasp
Disholcaspis cinerosa, commonly known as the mealy oak gall wasp, is a species of gall wasp in the family Cynipidae. It induces distinctive galls on oak trees (Quercus spp.), specifically known as "mealy oak galls." The species has been documented to possess exceptionally large venom glands relative to its body size—when unfolded, the venom gland is approximately 10 times longer than the abdomen—making it a notable subject in studies of gall induction mechanisms. Like other cynipid wasps, it exhibits an alternation of generations life cycle with both sexual and asexual (parthenogenetic) generations.
Disholcaspis eldoradensis
Honeydew Gall Wasp
Disholcaspis eldoradensis is a gall wasp in the family Cynipidae that induces distinctive galls on valley oak (Quercus lobata). The species is notable for producing sucrose-secreting galls that attract ant mutualists, particularly invasive Argentine ants, which defend the galls against parasitoids. This mutualism creates a complex trophic community involving multiple ant species, parasitoids, and other associated insects. The species exhibits variable ecological outcomes depending on local conditions, with ant defense benefits varying across years and gall characteristics influencing parasitism rates.
Disholcaspis globosa
Globular root gall
A cynipid gall wasp that induces distinctive globular galls on oak roots. Like other members of the genus Disholcaspis, it exhibits parthenogenetic reproduction in at least one generation, with females capable of producing offspring without mating. The species is one of approximately twelve described in the genus, all of which appear to have eliminated males from their life cycles. Root galls differ from the more commonly observed twig and leaf galls of related species, occurring below ground where they are rarely encountered unless excavated.
Disholcaspis lacuna
Disholcaspis lacuna is a species of gall wasp in the family Cynipidae. Like other members of its genus, it induces the formation of characteristic galls on oak trees (Quercus spp.). The species exhibits the typical cynipid life cycle involving alternation of generations, with separate sexual and parthenogenetic (agamic) generations that produce distinct gall forms. Its biology and gall morphology are less documented than those of congeners such as D. quercusmamma and D. quercusglobulus.
Disholcaspis pedunculoides
Disholcaspis pedunculoides is a species of gall wasp in the family Cynipidae, described by Weld in 1926. Like other members of the genus Disholcaspis, it induces gall formation on oak trees (Quercus spp.), though specific host associations and gall morphology for this species remain poorly documented in published literature. The species follows the typical cynipid life cycle with alternating sexual and asexual generations.
Disholcaspis perniciosa
Disholcaspis perniciosa is a gall wasp species in the family Cynipidae, first described by Bassett in 1890. As a member of the genus Disholcaspis, it is part of a diverse group of oak-associated gall wasps found in North America. The genus is known for producing distinctive bullet-shaped or spherical galls on oak trees. Like other cynipids, this species likely exhibits an alternation of generations life cycle with both sexual and agamic (asexual) generations producing different gall types on their host plants.
Disholcaspis prehensa
clasping twig gall wasp
Disholcaspis prehensa is a gall-inducing cynipid wasp native to California that induces distinctive galls on scrub oak and leather oak. The species exhibits heterogony, alternating between sexual and asexual generations that produce two morphologically different gall types. The asexual generation forms conspicuous mushroom-shaped twig galls with clasping bases, while the sexual generation produces small, hidden bud galls.
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quercusomnivora
Disholcaspis quercusomnivora is a species of gall wasp in the family Cynipidae, described by Ashmead in 1885. Like other members of the genus Disholcaspis, this species induces distinctive galls on oak trees (Quercus spp.), though specific gall morphology and host associations for this particular species are not well documented in available sources. The genus is known for producing hard, bullet-shaped or spherical galls on oak twigs and branches. As with many cynipid wasps, D. quercusomnivora likely exhibits an alternation of generations life cycle with both sexual and asexual phases.
Disholcaspis rubens
Disholcaspis rubens is a species of gall wasp in the family Cynipidae, first described by Gillette in 1893. Like other members of its genus, it induces distinctive galls on oak host plants. Species in Disholcaspis are known for producing twig galls rather than leaf galls, and many exhibit parthenogenetic reproduction with all-female generations. The genus is characterized by enlarged venom glands, which are implicated in the chemical induction of gall formation.
Disholcaspis simulata
Dried Peach Gall Wasp
Disholcaspis simulata is a species of gall wasp in the family Cynipidae, first described by Alfred C. Kinsey in 1922. Like other members of its genus, this species induces the formation of characteristic galls on oak trees (Quercus spp.). The species exhibits the complex alternation of generations typical of many cynipid wasps, with both sexual and agamic (parthenogenetic) generations. The specific epithet 'simulata' and common name 'Dried Peach Gall Wasp' suggest the gall's resemblance to a dried peach in appearance.
Disholcaspis spissa
Disholcaspis spissa is a gall-inducing wasp in the family Cynipidae, first described by Weld in 1957. Like other members of the genus Disholcaspis, it induces galls on oak trees (Quercus spp.), though specific host associations for this species are not well documented in the provided sources. The species is part of a diverse radiation of cynipid wasps that manipulate plant growth to create protective structures for their developing larvae. Research on related Disholcaspis species indicates these wasps possess enlarged venom glands, implicating venom compounds in the induction of gall formation.
Druon
Druon is a genus of cynipid gall wasps re-established in 2021, containing 15 Nearctic species that induce galls on oak trees (Quercus). Most species are known only from asexual generations, though two species (D. ignotum and D. quercuslanigerum) have documented alternating sexual and asexual generations. The genus was previously synonymized under Andricus but restored based on morphological and molecular evidence. Species are characterized by woolly galls and specific metasomal sculpturing patterns.
Druon fullawayi
yellow wig gall wasp
Druon fullawayi, commonly called the yellow wig gall wasp, is a species of cynipid gall wasp native to western North America. The species was originally described by William Beutenmüller in 1913 under the genus Andricus and later transferred to Druon. It induces distinctive woolly, yellowish galls on oak trees, particularly valley oak (Quercus lobata). The species has been documented in California and Zacatecas, Mexico.
Druon ignotum
Druon ignotum is a gall wasp that induces galls on white oaks in central and eastern North America. The species exhibits a complex life cycle with alternating sexual and asexual generations, each producing morphologically distinct galls. First described as Cynips ignota in 1881, it has undergone several taxonomic revisions before its current placement in Druon.
Druon quercuslanigerum
Druon quercuslanigerum is a cynipid gall wasp that alternates between sexual and asexual generations on live oaks. The sexual generation forms small galls on catkins in spring, while the asexual generation produces distinctive woolly galls on leaf undersides. It is one of only two species in the genus Druon with documented alternating generations.
Dryocosmus albidus
Dryocosmus albidus is a species of gall wasp in the family Cynipidae, described by Weld in 1944. Members of this genus are known to induce galls on oak trees (Quercus spp.). The species name 'albidus' (Latin for 'whitish') likely refers to some aspect of the adult's coloration or the gall structure. As with other cynipids, this species has a complex life cycle typically involving both sexual and asexual generations, with associated morphological and ecological differences between generations.
Dryocosmus archboldi
Dryocosmus archboldi is a species of gall wasp in the family Cynipidae, described in 2021 by Melika and Abrahamson. It belongs to a genus whose members are associated with oak trees (Quercus species) and induce the formation of galls. The species name honors Archbold Biological Station in Florida, where the type specimen was collected. As a recently described species, detailed biological information remains limited.
Dryocosmus asymmetricus
Dryocosmus asymmetricus is a species of gall wasp in the family Cynipidae, first described by Kinsey in 1922. The genus Dryocosmus comprises gall-forming wasps associated with oak trees (Quercus species). Members of this genus induce galls on various parts of host oaks, though specific details about D. asymmetricus remain limited in published literature.
Dryocosmus cinereae
Dryocosmus cinereae is a species of gall wasp in the family Cynipidae, first described by Ashmead in 1887. The genus Dryocosmus comprises gall-forming wasps associated with oak trees (Quercus species). As a member of Cynipidae, this species likely induces gall formation on host plant tissue, though specific gall morphology and host associations for D. cinereae require verification from primary sources. The species has been documented in citizen science observations, suggesting established populations in regions where it occurs.
Dryocosmus favus
Dryocosmus favus is a species of gall wasp in the family Cynipidae, first described by Beutenmueller in 1911. Members of the genus Dryocosmus are known to induce galls on oak trees (Quercus spp.). The species has been documented through a limited number of observations.
Dryocosmus quercuslaurifoliae
Dryocosmus quercuslaurifoliae is a species of gall wasp in the family Cynipidae. Species in this genus are associated with oak trees (Quercus), inducing the formation of galls on various plant tissues. The specific epithet "quercuslaurifoliae" indicates an association with Quercus laurifolia, the laurel oak.
Dryocosmus quercuspalustris
Succulent Oak Gall Wasp
Dryocosmus quercuspalustris, commonly known as the Succulent Oak Gall Wasp, is a species of gall wasp in the family Cynipidae. The species was described by Osten Sacken in 1861. It induces galls on oak trees, with the specific epithet referencing Quercus palustris (pin oak) as a host. The species has been recorded from eastern North America including Canada and the northeastern United States.
Feron bakkeri
Pinched Leaf Gall Wasp
Feron bakkeri is a gall wasp in the family Cynipidae that induces distinctive pink leaf galls on oaks. The species was formerly classified under the genus Andricus and has been reclassified to Feron based on phylogenetic studies. It is considered relatively uncommon compared to other oak gall wasps in its range.
Feron crystallinum
Crystalline Gall Wasp
Feron crystallinum is a cynipid gall wasp that induces distinct galls on oak species in California. The species exhibits complex alternation of generations, producing different gall morphologies and wasp forms across seasonal cycles. The parthenogenic generation forms large, pink, hairy galls on leaf undersides, while the bisexual generation produces smaller, less hairy galls. Formerly classified as Andricus crystallinus, this species has been reclassified to the genus Feron based on phylogenetic relationships.
Feron izabellae
Pink Bow-Tie Gall Wasp
Feron izabellae is an oak gall wasp described in 2023 from Northern California. The species induces distinctive pink spangle galls on white oak leaves, with galls folding at maturity to resemble bow-ties. Only the asexual generation is known. The species was named in memory of the senior author's mother and granddaughter.
Heteroecus dasydactyli
Heteroecus dasydactyli is a species of gall wasp in the family Cynipidae. The species was originally described by Ashmead in 1896 under the name Andricus dasydactyli before being transferred to the genus Heteroecus. Like other members of Cynipidae, this species induces gall formation on host plants, though specific host associations require verification. The species has been documented in iNaturalist observations, indicating ongoing contemporary presence.
Heteroecus lyoni
Lyon's Gall Wasp
Heteroecus lyoni is a gall wasp in the family Cynipidae. Species in this genus are known to induce galls on oaks (Quercus), though specific details for H. lyoni are limited in available literature. The species was named in honor of someone with the surname Lyon, a common practice in taxonomic nomenclature. Like other cynipids, it likely exhibits a complex life cycle with alternating generations between sexual and parthenogenetic forms, though this has not been explicitly documented for this species.
Heteroecus pacificus
beaked spindle gall wasp
Heteroecus pacificus, commonly known as the beaked spindle gall wasp, is a species of cynipid gall wasp native to western North America. It induces distinctive galls on canyon live oak (Quercus chrysolepis) and huckleberry oak (Quercus vaccinifolia). The species exhibits a complex life cycle with two annual generations: one sexual generation producing males and females, and one parthenogenetic generation consisting solely of females.
Holocynips
Holocynips is a genus of oak gall wasps in the family Cynipidae, established by Kieffer in 1910. The genus occurs in the Nearctic region and contains species that induce galls on oak trees (Quercus). A 2023 description of H. illinoiensis prompted preliminary discussion on the coherence of the genus, suggesting taxonomic boundaries may require revision.
Kokkocynips difficilis
Kokkocynips difficilis is a species of gall wasp in the family Cynipidae, originally described by Ashmead in 1887 as Andricus difficilis. It belongs to the tribe Cynipini, which comprises gall-inducing wasps associated with oaks (Quercus spp.). The genus Kokkocynips was established relatively recently (2021) following taxonomic revision of the oak gall wasps. Like other members of this genus, K. difficilis likely induces galls on oak species, though specific host associations and gall morphology remain poorly documented in the available literature.
Kokkocynips rileyi
Kokkocynips rileyi is a species of gall wasp in the family Cynipidae. The species was originally described as Andricus rileyi by Ashmead in 1896 and has undergone taxonomic revisions, with recent treatments placing it in the genus Kokkocynips. Like other cynipid wasps, this species induces gall formation on host plants, though specific gall morphology and host associations require further documentation. The species name honors an individual, following the Latin convention for male eponyms ending in -i.
Liposthenes glechomae
Glechoma gall wasp
Liposthenes glechomae is a gall wasp in the family Cynipidae that induces galls on its host plant Glechoma hederacea (ground ivy). The species is native to Europe and Northern Asia but has been introduced to North America, where both the wasp and its host plant are considered invasive. Research on this species has contributed to understanding gall induction mechanisms in cynipid wasps.
Loxaulus
Loxaulus is a genus of gall wasps in the family Cynipidae, tribe Cynipini. Members are small, specialized insects that induce the formation of plant galls on host plants. The genus was established by Mayr in 1881. As with other cynipine gall wasps, species likely exhibit complex life cycles involving alternating sexual and asexual generations on different host plants, though detailed studies remain limited.
Melikaiella flora
Live Oak Petiole Gall Wasp
Melikaiella flora is a cynipid gall wasp that induces distinctive leaf galls on live oak trees (Quercus agrifolia, Q. wislizeni, and Q. chrysolepis) in California. The species exhibits a complex life cycle involving two generations, with second-generation females ovipositing into acorns rather than leaf tissue. The galls typically engulf the entire petiole and may extend into the leaf blade, representing one of the more dramatic gall morphologies among North American Cynipidae. Formerly classified as Callirhytis milleri, the species was reclassified to reflect improved understanding of cynipid phylogeny.
Neuroterus
Neuroterus is a genus of gall wasps in the family Cynipidae that induce galls on oak species. The genus was first described by Theodor Hartig in 1840. Some species, notably N. saltatorius, produce distinctive 'jumping galls' that detach from the host plant and move along the ground due to larval activity within. Like other oak gall wasps, Neuroterus species exhibit complex life cycles with alternating sexual and asexual (agamic) generations, with each generation typically inducing galls on different parts of the host plant.
Neuroterus alexandrae
Neuroterus alexandrae is a species of gall wasp in the family Cynipidae, described by Nicholls and Melika in 2021. As a member of the genus Neuroterus, it induces distinctive galls on oak leaves. The genus Neuroterus is known for producing some of the most recognizable oak galls, including spangle galls and jumping galls.
Neuroterus aliceae
Neuroterus aliceae is a species of gall wasp in the family Cynipidae, described in 2021. Like other members of the genus Neuroterus, this species induces gall formation on oak leaves. The genus Neuroterus is known for creating distinctive leaf galls, including the jumping oak galls where larval movement can cause detached galls to move. This species is part of the diverse radiation of oak-associated cynipid wasps, with over 800 species of oak gall wasps documented in North America.
Neuroterus fragilis
succulent gall wasp
Neuroterus fragilis, commonly known as the succulent gall wasp, is a species of oak gall wasp in the family Cynipidae. It induces distinctive galls on California oak species, with only the spring bisexual generation currently documented. The species is notable for its relatively large galls compared to its tiny adult body size.
Neuroterus minutus
Neuroterus minutus is a species of gall wasp in the family Cynipidae. Members of this genus are known to induce galls on oak trees (Quercus species). The species has been documented through citizen science observations, with records available on platforms such as iNaturalist. As with many cynipid wasps, detailed biological studies of this particular species appear limited in the available literature.
Neuroterus niger
jumping gall wasp, black jumping gall wasp
Neuroterus niger is a species of gall wasp in the family Cynipidae that induces distinctive galls on oak leaves. This species exhibits an alternating sexual and asexual life cycle, with different generations producing different gall types. The asexual generation creates small, spherical galls that detach from leaves and fall to the ground, where the larva's movements cause the gall to jump—giving rise to the common name "jumping gall." The sexual generation produces inconspicuous leaf galls. The species has been documented across much of eastern and central North America.
Neuroterus quercusmajalis
Neuroterus quercusmajalis is a species of gall wasp in the family Cynipidae that induces galls on oak leaves. As a member of the genus Neuroterus, it is part of a diverse group of cynipid wasps specialized on oaks. The species was described by Beutenmüller in 1909. Like other Neuroterus species, it exhibits complex life cycles involving both sexual and asexual generations that produce different gall types on oak tissues.
Neuroterus quercusminutissimus
Neuroterus quercusminutissimus is a species of cynipid gall wasp in the genus Neuroterus. The species was described by Ashmead in 1885 and is associated with oak trees (Quercus species), as indicated by its specific epithet. Like other Neuroterus species, it induces the formation of galls on oak tissue. The genus Neuroterus contains approximately 50-60 species, many of which are known for their complex life cycles involving alternating generations that produce different gall types on oak leaves or other plant parts.
Neuroterus saltarius
Jumping Gall Wasp
Neuroterus saltarius is a species of gall wasp in the family Cynipidae, commonly known as the jumping gall wasp. It induces distinctive leaf galls on oak trees that detach from the plant and fall to the ground. The developing larva inside the fallen gall can cause the gall to jump when it moves, a unique behavioral trait that gives the species its common name. This species is part of the diverse Neuroterus genus, which contains many oak-associated gall wasps with complex life cycles.
Neuroterus saltatorius
Jumping Gall Wasp
Neuroterus saltatorius, commonly known as the jumping gall wasp, is a North American oak gall wasp in the family Cynipidae. The species is notable for producing small, round galls that detach from leaves and exhibit a distinctive jumping behavior caused by larval movements. It has a complex life cycle with alternating bisexual and parthenogenetic generations, each producing different gall types on oak hosts.
Paracraspis patelloides
Bowl Gall Wasp
Paracraspis patelloides, commonly known as the Bowl Gall Wasp, is a species of cynipid wasp that induces distinctive gall formation on host plants. First described by Trotter in 1910 under the genus Andricus, it was later transferred to Paracraspis. The species is part of the diverse Cynipidae family, which contains numerous gall-inducing wasps primarily associated with oaks and related plants. Observations suggest it is moderately well-documented, with over 100 records on iNaturalist.
Phanacis
Phanacis is a genus of cynipid gall wasps (Hymenoptera: Cynipidae) in the tribe Aylacini. The genus includes species that induce galls on host plants, with Phanacis taraxaci documented as forming leaf galls on common dandelion (Taraxacum officinale). These galls function as physiological sinks that actively redirect carbon and mineral nutrients from host tissues.