Europe
Guides
Astacidae
Astacid Crayfishes, Freshwater Crayfish
Astacidae is a family of freshwater crayfish comprising four extant genera: Astacus, Austropotamobius, Pontastacus, and Pacifastacus. Members are native to Europe, western Asia, and western North America. The family is distinguished by a gonopore positioned on the coxa of the third pair of walking legs in males (fifth pair in females), a trait shared with all Astacoidea. Several species, particularly Pacifastacus leniusculus (signal crayfish), have become invasive outside their native ranges, carrying crayfish plague (Aphanomyces astaci) that threatens native European populations.
Atheta longicornis
Atheta longicornis is a rove beetle species in the family Staphylinidae, first described by Gravenhorst in 1802. The species has a broad native distribution across Europe, North Africa, and parts of Asia, and has been introduced to North America (Canada and the United States), Australia, and other regions. As a member of the hyperdiverse genus Atheta, it inhabits various terrestrial habitats where it contributes to decomposition processes. The species is characterized by its relatively long antennae, which inspired its specific epithet.
Athetini
Athetini is a large tribe of rove beetles (Staphylinidae: Aleocharinae) comprising at least 50 genera and 430 described species. The tribe is taxonomically complex, with ongoing research into its limits and internal relationships; it was not recovered as monophyletic in molecular phylogenetic studies. Multiple independent origins of coastal habitat specialization have been documented within the tribe.
Atomaria lederi
silken fungus beetle
Atomaria lederi is a species of silken fungus beetle in the family Cryptophagidae, described by Johnson in 1970. The species has a broad geographic distribution spanning three continents: Europe and Northern Asia (excluding China), North America, and Southern Asia. As a member of the Cryptophagidae, it belongs to a family commonly known as silken fungus beetles due to their association with fungal growth and the silky secretions produced by some species.
Atomaria lewisi
silken fungus beetle
Atomaria lewisi is a species of silken fungus beetle in the family Cryptophagidae. The species was described by Reitter in 1877 and is native to Europe. It belongs to a family commonly associated with fungal habitats.
Atomaria mesomela
silken fungus beetle
Atomaria mesomela is a species of silken fungus beetle in the family Cryptophagidae, native to Europe. It belongs to a genus of small beetles commonly found in association with fungal growth. The species was described by Herbst in 1792. Like other members of Cryptophagidae, it is likely associated with decaying organic matter and fungal substrates, though specific ecological details for this species remain poorly documented.
Atomaria subangulata
Atomaria subangulata is a species of silken fungus beetle in the family Cryptophagidae, described by J. Sahlberg in 1926. The genus Atomaria comprises small beetles commonly associated with fungal habitats. This species is documented from northern Europe, with confirmed records from Denmark, Norway, and Sweden. Like other members of Cryptophagidae, it is presumed to inhabit decaying organic matter and fungal environments, though specific ecological details for this species remain limited.
Atomaria testacea
silken fungus beetle
Atomaria testacea is a species of silken fungus beetle in the family Cryptophagidae. It has a broad geographic distribution spanning Africa, Europe and Northern Asia (excluding China), and North America. The species was described by Stephens in 1830. As a member of the Cryptophagidae, it is associated with fungal habitats, though specific ecological details for this species remain limited in the available literature.
Atrichopogon
biting midges, grass punkies
Atrichopogon is a genus of small biting midges in the family Ceratopogonidae, subfamily Forcipomyiinae. Species occur across multiple continents with documented records from the Neotropical region, Patagonia, Europe, and Asia. Some species are ectoparasites of larger insects, while others develop on aquatic vegetation or in terrestrial habitats such as rotting wood. Immature stages show habitat-specific distribution patterns: larvae of A. wirthi occur on water lettuce leaves, and larvae of European Meloehelea subgenus species inhabit rotting pine wood.
Atrichopogon minutus
Atrichopogon minutus is a species of biting midge in the family Ceratopogonidae, first described by Johann Wilhelm Meigen in 1830. Originally placed in the genus Ceratopogon, it was later transferred to Atrichopogon. The species belongs to the subfamily Forcipomyiinae, a group whose members are often associated with various ecological roles including pollination and parasitism. Records indicate presence in European and Atlantic island localities including the Azores.
Aulogymnus
Aulogymnus is a genus of chalcidoid wasps in the family Eulophidae, first described by Förster in 1851. Members of this genus are small parasitoid wasps, part of a diverse family that primarily parasitizes other insects. The genus has been recorded from Europe and Asia. Specific biological details for the genus as a whole remain poorly documented in accessible literature.
parasitoidEulophidaeChalcidoideaHymenopteraPalearcticwaspsinsectsarthropodsentomologytaxonomyFörster-1851DenmarkSpainTibetChinaEuropeAsiaminute-waspschalcid-waspsEulophinaeTerebrantesApocritaHexapodaAnimaliaArthropodaInsectaAulogymnussmall-waspstiny-waspsparasitic-waspsbiological-controlinsect-parasitoidssystematicsmorphologyidentificationkeysNearcticOrientalXizangcitizen-scienceiNaturalistobservationsrecordsdistributionFörster1851genusacceptedvalidsynonymychalcidoidchalcidchalcidseulophideulophidseulophid-waspseulophinesparasitoid-waspsparasitic-Hymenopterabiological-control-agentsinsect-diversitybiodiversityfaunaentomologicalhymenopteranarthropodhexapodhexapodspterygotepterygotesendopterygoteendopterygotesholometabolousholometabolatiny-insectsminute-insectssmall-insectsmicrohymenopteramicro-waspsmicro-parasitoidsmicro-chalcidsmicro-eulophidswing-venationantennaethoraxdiagnostic-characterstaxonomic-keysidentification-keysgeneric-keysNearctic-faunaEuropean-faunaAsian-faunaTibetan-faunaSpanish-faunaDanish-faunapoorly-knowndata-deficientunderstudiedcryptic-diversityhost-unknownbiology-unknownlife-history-unknownecology-unknowndistribution-recordsoccurrence-recordsspecimen-recordsmuseum-recordsdatabase-recordsGBIFCatalogue-of-LifeNCBIWikipediaUniversal-Chalcidoidea-DatabaseKey-to-Nearctic-eulophid-generaliteraturesourcesreferencescitationsbibliographyoriginal-descriptiontype-speciestype-localitynomenclaturesystematic-entomologyhymenopterologychalcidologyparasitologybiological-control-researchintegrated-pest-managementIPMagricultural-entomologyforest-entomologymedical-entomologyveterinary-entomologyurban-entomologyconservation-entomologyinsect-ecologycommunity-ecologypopulation-ecologybehavioral-ecologyevolutionary-ecologyphylogeneticsphylogenymolecular-systematicsDNA-barcodingtaxonomy-and-phylogenyclassificationbiodiversity-informaticsbiogeographyhistorical-biogeographyphylogeographydispersalvicariancespeciationdiversificationevolutionadaptationnatural-selectionsexual-selectionlife-history-evolutionhost-parasitoid-interactionscoevolutiontritrophic-interactionsfood-websecosystem-servicesnatural-enemiesbiocontrolaugmentative-biological-controlclassical-biological-controlconservation-biological-controlinvasive-species-managementpest-managementsustainable-agricultureorganic-farmingagroecologyecosystem-healthenvironmental-monitoringbioindicatorsindicator-speciesclimate-changeglobal-changehabitat-lossfragmentationconservation-statusIUCNred-listnot-evaluatedresearch-needsknowledge-gapsfuture-researchprioritiesspecimen-collectionvoucheringmuseum-collectionsnatural-history-collectionsdigitizationdata-sharingopen-scienceFAIR-principlescitizen-science-contributionscommunity-sciencepublic-engagementscience-communicationeducationoutreachnatural-historyinsect-watchingwasp-watchingnature-observationbiodiversity-appreciationAxinotarsus pulicarius
soft-winged flower beetle
A small soft-winged flower beetle (Melyridae) measuring 2.75–3.75 mm, distinguished by a reddish-yellow pronotum with a broad black median band contrasting with dark grey to nearly black head and elytra. Native to Europe with broad distribution excluding northern latitudes and southern Mediterranean regions; introduced to North America where established in the Pacific Northwest and Great Lakes region. Known for elaborate courtship involving excitatory gland displays and mutual frontal tapping between sexes.
Bactra lancealana
Black-blotched Bactra Moth
Bactra lancealana is a small tortricid moth with a wingspan of 11–20 mm, distributed across Europe. Adults are active from May to October. Larvae feed primarily on rushes in the genera Juncus and Scirpus. The species is morphologically similar to Bactra furfurana, requiring careful examination for identification.
Ballistura tuberculata
Ballistura tuberculata is a species of springtail in the family Isotomidae, described by J. Stach in 1947. It belongs to the order Entomobryomorpha, one of the major lineages of Collembola. The specific epithet 'tuberculata' suggests the presence of tubercle-like structures, though detailed morphological descriptions remain limited in available literature.
Barynotus obscurus
Ground Weevil
Barynotus obscurus is a species of weevil in the family Curculionidae, native to Europe. The species has been introduced to North America, with established populations in British Columbia, New Brunswick, and other regions. It is commonly known as the Ground Weevil.
Bathyplectes infernalis
Bathyplectes infernalis is a parasitoid wasp in the family Ichneumonidae. It is a biological control agent of the alfalfa weevil (Hypera postica), a major agricultural pest in western North America. The species has been introduced and released in the United States as part of classical biological control programs. It is one of several Bathyplectes species used to manage weevil populations in alfalfa fields.
Bellardia vulgaris
Pale-veined Emerald-bottle
Bellardia vulgaris is a species of blow fly in the family Calliphoridae, commonly known as the Pale-veined Emerald-bottle. It is a metallic green fly distributed across parts of Europe including Belgium, Denmark, Norway, and Sweden. As a member of the Calliphoridae, it is likely associated with decomposing organic matter, though specific ecological details remain poorly documented in the provided sources.
Bembidion bruxellense
Bembidion bruxellense is a species of ground beetle in the family Carabidae, first described by Wesmael in 1835. It is native to Europe and has been recorded from North America (Canada and USA), suggesting either a native Holarctic distribution or possible introduction. The species belongs to the large genus Bembidion, which contains over 1,200 described species of small to medium-sized predatory beetles.
Bembidion lampros
A small ground beetle native to Europe, now established in North America. Adults overwinter and reproduce in early spring in arable fields, where they function as generalist predators of cereal aphids. The species exhibits wing dimorphism with three phenotypes, though macropterous individuals rarely possess functional flight muscles. Low lifetime fecundity (approximately 10 eggs per female) and high juvenile mortality (66%) make population dynamics sensitive to conditions during larval development.
Bembidion tetracolum
Seine Riverbank Ground Beetle
Bembidion tetracolum is a species of ground beetle in the family Carabidae, first described by Thomas Say in 1823. It is native to Europe and has been introduced to other regions including Australia. The species is commonly known as the Seine Riverbank Ground Beetle, reflecting its association with riparian habitats. It belongs to the large genus Bembidion, which contains numerous small, often metallic ground beetles.
Berginus
Berginus is a genus of hairy fungus beetles in the family Mycetophagidae, established by Erichson in 1846. The genus contains at least three described species distributed across North America and Europe. Members of this genus are associated with fungal habitats. The genus is classified in the subfamily Bergininae, which bears its name.
Beris fuscipes
Short-horned Black Legionnaire
Beris fuscipes, commonly known as the short-horned black legionnaire, is a European soldier fly in the family Stratiomyidae. The species is distinguished by its bright shining green thorax and scutellum, black abdomen, and extensively dark dull orange or dusky legs. It inhabits damp woodlands, marshes, and fens, with adults active from May through September.
Bibloplectus
ant-loving beetles
Bibloplectus is a genus of minute rove beetles (Staphylinidae: Pselaphinae) commonly known as ant-loving beetles. The genus contains at least 10 described species distributed in North America and Europe. Seven new species were recently described from the eastern United States based on specimens from museum collections. Members of this genus are typically associated with ants and are characterized by their small size and reduced morphology.
Billaea
Billaea is a genus of tachinid flies comprising approximately 80 described species. Most larvae are parasitoids of beetle larvae, particularly cerambycids and lucanids, with some species attacking lepidopteran hosts in the family Pyralidae. Several species have been investigated as biological control agents for agricultural and forestry pests, including palm weevils and sugarcane borers.
Tachinidaeparasitoidbiological-controlCerambycidaeDryophthoridaeCrambidaeviviparousplanidiaforest-pestagricultural-pestoil-palmsugarcanepalm-weevilwood-borersoutheast-AsiaSouth-AmericaEuropeBulgariaBrazilTurkeyMexicoPeruintegrated-pest-managementnatural-enemyhost-searchingkairomonessuperparasitismmass-rearingGalleria-mellonellalaboratory-hostfrassgallerytunnelpupal-parasitoidlarval-parasitoidectoparasitoidendoparasitoidColeopteraLepidopteraDipteraBrachyceraDexiinaeDexiiniRobineau-Desvoidy-183080-speciesglobal-distributionforestryagricultureornamental-palmsdate-palmcoconutRhynchophorusDiatraeaPrionusXylotrechusclimate-matchingestablishment-potentialMediterraneanMiddle-EastCaliforniainvasive-speciesred-palm-weevilAmerican-palm-weevilred-ring-diseasevector-controlnatural-mortalitypest-managementsustainable-agriculturehost-specificitytrophic-associationnew-host-recordparasitism-ratefield-parasitismlaboratory-rearingadult-longevityfecunditysex-ratiopupal-viabilitypupal-durationplanidia-lengthfood-resourcespollenhoneysucroseflowering-plantsspontaneous-floracompanion-plantshabitat-managementconservation-biological-controlaugmentative-biological-controlclassical-biological-controlquarantinehost-rangecircumstantial-evidencetaxonomic-confusionB.-rhynchophoraeB.-menezesiB.-adelphaB.-claripalpisB.-rutilansB.-cerambycivoraB.-steiniB.-fortisB.-irrorataB.-pectinataB.-setosaB.-trianguliferaB.-lataB.-intermediaB.-biserialisB.-giganteaB.-rubensB.-maritimaB.-marmorataB.-aurifronsB.-verticalisB.-flavaB.-chinensisB.-brevicaudaB.-kurahashiiB.-micronychiaB.-papeiB.-setigeraB.-yintiaolingB.-rufescensB.-triquetrusB.-rubidaB.-rufiventrisB.-brasiliensisB.-friburgensisB.-giacomeliB.-kosteraeB.-plaumanniB.-shannoniB.-africanaB.-argentaureaB.-atkinsoniB.-capensisB.-communisB.-decisaB.-edwardsiB.-erectaB.-ficorumB.-grandisB.-impigraB.-interruptaB.-kolomyetziB.-luteolaB.-malayanaB.-minorB.-monohammiB.-montanaB.-morosaB.-nipigonensisB.-orbitalisB.-ovataB.-quadrinotaB.-rhingiaeformisB.-robustaB.-satisfactaB.-sibleyiB.-sjostedtiB.-solivagaB.-trivittataB.-trochanterataB.-vanemdeniB.-velutinaB.-versicolorB.-villeneuveiB.-vitripennisB.-ziminiB.-lateralisB.-lativentrisBlaniulus
Blaniulus is a genus of millipedes in the family Blaniulidae, order Julida. The genus was established by Gervais in 1836 and contains multiple species. Members are characterized by their elongated, cylindrical bodies with numerous segments. The genus has been documented across parts of Europe including Germany, Italy, the Czech Republic, and Croatia.
Blaniulus guttulatus
Spotted Snake Millipede
Blaniulus guttulatus, the spotted snake millipede, is a small, eyeless millipede native to Central and Western Europe that has been widely introduced to North America, Australia, and remote Atlantic islands. It is characterized by its slender, pale body with distinctive red ozadenes (defensive glands) on each segment. The species thrives in anthropogenic habitats and can become an agricultural pest, particularly during drought conditions.
Blastodacna atra
Apple Pith Moth
Blastodacna atra, commonly known as the apple pith moth, is a small moth in the family Elachistidae. It is native to most of Europe and has been introduced to North America, where it has been recorded in Massachusetts and Ontario. The species is notable for its larval habit of mining inside the young shoots of apple trees (Malus species), which can cause localized damage to orchards. Adults are active from May through September in western Europe.
Boloria freija freija
Freija Fritillary
The Freija Fritillary (Boloria freija freija) is a subspecies of butterfly in the family Nymphalidae. It is found in both Europe and North America, with distribution records from Belarus and broader European regions as well as North American localities. As a member of the genus Boloria, it shares characteristics with other fritillary butterflies, including complex life cycles with specific host plant relationships. The subspecies is relatively poorly documented compared to congeners such as Boloria selene.
Bootanomyia
Bootanomyia is a genus of parasitoid wasps in the family Megastigmidae. Members are small chalcidoid wasps that develop as parasitoids within oak galls induced by cynipid gall wasps. Two European species morphologically keyed to B. dorsalis have been introduced to North America: one clade found in New York, and another in the Pacific Northwest. These introductions represent the first documented cases of Bootanomyia establishment in North America.
Borboropsis
Borboropsis is a genus of small flies in the family Heleomyzidae, established by Czerny in 1902. The genus contains at least four described species distributed across Europe and East Asia. Species in this genus are part of a family commonly associated with decaying organic matter and shaded, moist habitats.
Brachycentrus
Grannom Caddisflies, Humpless Casemaker Caddisflies
Brachycentrus is a genus of caddisflies in the family Brachycentridae, commonly known as grannoms or humpless casemaker caddisflies. The genus contains at least 30 described species distributed across North America and Eurasia. Larvae are aquatic filter-feeders that construct portable cases from silk and environmental materials such as sand, small stones, or plant fragments. The genus is ecologically significant as an important prey item for trout and other freshwater fish.
Brachypterus urticae
Brachypterus urticae is a species of short-winged flower beetle in the family Kateretidae. It is native to Europe and has been recorded in North America as present. The species belongs to a group of beetles commonly associated with flowers and decaying plant material.
Brachysomus echinatus
Brachysomus echinatus is a European weevil species in the family Curculionidae, subfamily Entiminae. First described by Bonsdorff in 1785, it was established as the senior synonym of B. hirsutus in a 2022 taxonomic revision. The species has been documented across northern and eastern Europe including Russia (Adygey, Altay, Arkhangel'sk, Bashkortostan, Buryat) and is one of 56 species included in a comprehensive morphological review of the genus.
Brassicogethes aeneus
common pollen beetle, rape pollen beetle, rape blossom beetle
Brassicogethes aeneus is a small pollen beetle in the family Nitidulidae and a major pest of oilseed rape (canola) and other Brassica crops across Europe, Northern Asia, and North America. Adults are 2–3 mm long, metallic black with greenish reflections, while larvae reach 3 mm and are white with brown sclerotized plates. The species causes significant economic damage through adult feeding and oviposition in flower buds, which can lead to bud abortion and yield losses up to 80% in spring oilseed rape. It was previously classified under the genus Meligethes. Populations show low genetic structure across Europe, indicating substantial gene flow, and are subject to biological control by hymenopteran parasitoids.
Broscus cephalotes
coastal ground beetle
Broscus cephalotes is a nocturnal, flightless ground beetle in the family Carabidae. It is native to coastal regions of temperate Europe and western Siberia, and was introduced to eastern Canada around 1975, subsequently spreading into the United States. The species is recognized as beneficial in agricultural contexts due to its predatory habits.
Bruchus rufimanus
Broadbean Weevil, Broad Bean Beetle, Broad Bean Seed Beetle
Bruchus rufimanus is a univoltine leaf beetle and major agricultural pest of faba beans (Vicia faba). Adults feed on pollen and nectar, particularly from Fabaceae, while larvae develop inside seeds where they are protected from control measures. The species exhibits precise phenological synchrony with its host plant, with adult colonization timed to crop flowering. Sexual maturation is triggered by photoperiod and pollen consumption, with females becoming reproductive after ingesting faba bean pollen. Overwintering occurs as dormant adults in protected sites or as larvae/pupae within stored seeds.
Brumus quadripustulatus
Cream-spot ladybird, Cream-spot lady beetle
Brumus quadripustulatus is a small ladybird beetle native to Europe and northern Asia, now established in North America. It is recognized by its distinctive cream-colored spots on a dark, nearly black background. The species inhabits various wooded and shrubby environments and has been observed preying on aphids and scale insects.
Caenocara
puffball beetles
Caenocara is a genus of small beetles in the family Ptinidae (subfamily Dorcatominae), commonly known as puffball beetles. The genus was established by C.G. Thomson in 1859. Members are associated with fungi, particularly puffball mushrooms, which serve as their primary habitat and food source. The genus has been documented across northern Europe including Denmark, Norway, and Sweden.
Calaphis flava
Yellow Dark-veined Birch Aphid
Calaphis flava is a species of aphid in the family Aphididae, first described by Mordvilko in 1928. It is commonly known as the Yellow Dark-veined Birch Aphid. The species has been recorded in parts of Europe including Belgium, Denmark, Norway, and the Azores (São Miguel). As a member of the genus Calaphis, it is associated with birch (Betula) host plants, though specific host records for this species are limited in the available sources.
Callima
Callima is a genus of concealer moths in the family Oecophoridae, established by Clemens in 1860. The genus includes species such as Callima argenticinctella, commonly known as the orange-headed epicallima. These are small moths that are frequently encountered at blacklight setups during nocturnal surveys. The genus has been documented from multiple continents including North America and Europe.
Callipallene brevirostris
Long-necked Sea Spider
Callipallene brevirostris is a sea spider species in the family Callipallenidae, commonly known as the Long-necked Sea Spider. It was originally described as Pallene brevirostris by Johnston in 1837 and later transferred to the genus Callipallene. The species is known from European waters and has two recognized subspecies: C. b. brevirostris and C. b. producta.
Callipterinella
Callipterinella is a genus of aphids in the family Aphididae, established by van der Goot in 1913. The genus contains three recognized species: C. calliptera, C. minutissima, and C. tuberculata. Species are distributed across Europe and North America. The genus belongs to the tribe Calaphidini within the subfamily Calaphidinae.
Callipterinella calliptera
Black-banded Birch Aphid
Callipterinella calliptera is an aphid species commonly known as the Black-banded Birch Aphid. It belongs to the family Aphididae and is associated with birch trees (Betula species). The species has been documented in parts of northwestern Europe including Belgium, Denmark, and Norway. It is a relatively infrequently recorded species with limited published ecological data.
Callopistromyia
Peacock flies
Callopistromyia is a genus of small picture-winged flies (family Ulidiidae) known for the distinctive courtship displays of males, which hold their patterned wings perpendicular to the body while sidling along substrates. The genus contains two species: C. annulipes (the peacock fly) and C. strigula. Adults are typically 3.5–5.5 mm in length. C. annulipes has expanded from its native North American range to become an established alien species in Europe since 2007.
Calosoma sycophanta
forest caterpillar hunter, Agreeable Caterpillar Hunter
Calosoma sycophanta is a large, metallic ground beetle renowned for its specialized predation on caterpillars. Adults reach 21–35 mm in length and display striking iridescent green elytra with a bluish scutellum, though coloration shifts with light angle to show blue, bronze, copper, gold, or black tones. Both adults and larvae are arboreal hunters, ascending trees to prey on caterpillars including Lymantria dispar (gypsy moth), Thaumetopoea processionea (oak processionary), and Thaumetopoea pityocampa. The species is diurnal and visually orients toward dark objects and tree bases in forests. Native to Europe, it has been introduced to North America for biological control and is now established in the northeastern United States.
Camptomyia
Camptomyia is a genus of gall midges (Cecidomyiidae) established by Kieffer in 1894. At least one species, C. pseudotsugae, has been described from Douglas-fir cones in western North America. The genus belongs to the tribe Asynaptini within the subfamily Porricondylinae.
Campylomma
mullein bugs
Campylomma is a genus of plant bugs (family Miridae, tribe Nasocorini) comprising at least 12 recognized species. The genus is best known from detailed studies of C. verbasci, the mullein bug, which exhibits a predator-phytophage feeding strategy and has been investigated for pheromone-based mating disruption in orchard systems. Japanese species have been taxonomically revised, with emphasis on genitalic characters for identification. The genus occurs across temperate regions of the Northern Hemisphere including North America, Europe, and East Asia.
Capitophorus elaeagni
Thistle Aphid, Artichoke Aphid, Oleaster-Thistle Aphid
Capitophorus elaeagni is a species of aphid in the family Aphididae, commonly known as the thistle aphid, artichoke aphid, or oleaster-thistle aphid. It is found in Europe, with distribution records also from the Azores (São Miguel), Brazil (Paraná, São Paulo), Czech Republic, Denmark, and Norway. The specific epithet 'elaeagni' suggests an association with plants in the genus Elaeagnus, though this relationship requires confirmation.
Caplothorax
sap beetles
Caplothorax is a genus of sap beetles in the family Nitidulidae. The genus comprises 11 described species distributed across North America and Europe. It was established by Kirejtshuk in 1997. GBIF lists Caplothorax as a synonym of Carpophilus, though iNaturalist maintains it as a valid genus with over 500 observations.
Capperia
Capperia is a genus of plume moths in the family Pterophoridae, established by Tutt in 1905. Species are small with wingspans ranging from approximately 12.5 to 20.5 mm. The genus is distributed across Europe, extending into parts of Asia including Kazakhstan and Turkey. Members exhibit bivoltine flight periods with adults active in late spring and mid-summer.