Multivoltine

Guides

Recurvaria
Recurvaria is a genus of small moths in the family Gelechiidae, established by Haworth in 1828. The genus contains approximately 40 described species distributed across the Holarctic region. Several species are economically significant as pests of coniferous trees and pistachio, with larvae that mine needles or feed internally within fruits. The genus exhibits diverse life history strategies, including univoltine and bivoltine cycles, with some species requiring two years to complete development.
Gelechiidae Litini needle-miner pistachio-pest forest-pest Holarctic internal-feeder diapause multivoltine semivoltine
Renia salusalis
Dotted Renia Moth, dotted renia
Renia salusalis, commonly known as the Dotted Renia Moth, is a litter moth in the family Erebidae. It was first described by Francis Walker in 1859. The species occurs across the eastern and central United States, where its larvae feed on detritus. Adults are active from late spring through early autumn, with generation timing varying by latitude.
litter-moth detritivore multivoltine eastern-North-America forest-floor
Silba
lance flies
Silba is a genus of lance flies in the family Lonchaeidae, containing more than 120 described species. The genus has a cosmopolitan distribution, occurring on every continent except South America and Antarctica. Species within Silba exhibit diverse larval development strategies, with some species associated with figs (including the economically significant pest Silba adipata), while others develop in flower buds, fungal fruiting bodies, or other substrates. The genus is taxonomically active, with numerous new species described in recent decades, particularly from the Afrotropical and Asian regions.
Lonchaeidae lance-flies fig-pests invasive-species cosmopolitan multivoltine
Slossonella
Slossonella is a genus of slug caterpillar moths (family Limacodidae) described by Harrison Gray Dyar in 1904. The genus contains species formerly placed in Hypargyria, including Slossonella slossonella (formerly Hypargyria slossonella), a snout moth known from Florida and Mexico. Species in this genus have multiple generations annually and are associated with specific host plants.
Limacodidae slug-caterpillar-moth North-America Florida Mexico multivoltine Hippocratea-volubilis
Sparganothoides lentiginosana
Lentiginos Moth
Sparganothoides lentiginosana, the lentiginos moth, is a small tortricid moth and the sole member of the Lentiginosana Group within its genus. It occurs across the southeastern and south-central United States south to Mexico. Adults are active nearly year-round with multiple generations per year. Sexual dimorphism in forewing length is documented.
Tortricidae Sparganothini United-States Mexico multivoltine year-round-activity
Stenelmis crenata
riffle beetle
Stenelmis crenata is a species of riffle beetle in the family Elmidae, occurring in North America. It inhabits fast-flowing stream environments, particularly riffle habitats in tallgrass prairie streams. The species exhibits a multivoltine life cycle with overlapping cohorts and year-round larval presence. Adults are active from late spring through autumn.
riffle-beetle aquatic-insect benthic-macroinvertebrate prairie-stream multivoltine Elmidae North-America fast-flowing-water submerged-substrates
Stictocephala stimulea
Stictocephala stimulea is a treehopper species in the family Membracidae, first described by Van Duzee in 1914. Treehoppers in this genus are characterized by their enlarged, often ornate pronotum that extends over the body like a helmet or thorn. The specific epithet 'stimulea' suggests possible resemblance to or association with stinging organisms, though this refers to appearance rather than actual sting. Like other membracids, this species likely feeds on plant phloem sap using piercing-sucking mouthparts.
Membracidae treehopper Auchenorrhyncha Cicadomorpha Stictocephala Ceresini Smiliinae Hemiptera sap-feeder pronotum helmet thorn-mimicry plant-feeding-insect phloem-feeder cryptic-coloration camouflage Van-Duzee-1914 Nearctic North-America Florida North-Carolina South-Dakota understudied-species data-deficient taxonomy entomology insect arthropod true-bug auchenorrhynchan cicadomorph leafhopper-relative plant-hopper ornate-pronotum pronotal-extension dorsal-helmet insect-morphology insect-systematics biodiversity natural-history conservation insect-ecology insect-behavior insect-development insect-distribution biogeography species-description type-specimen original-description historical-taxonomy museum-specimen collection-record occurrence-data GBIF iNaturalist citizen-science observation-record range-map distribution-data geographic-range endemic native-species regional-fauna local-biodiversity habitat-specificity host-plant feeding-guild phytophagous herbivore plant-insect-interaction coevolution mimicry defense crypsis evolutionary-adaptation morphological-adaptation exoskeleton-modification structural-coloration insect-identification field-guide diagnostic-character taxonomic-key species-complex cryptic-species morphological-variation geographic-variation phenotypic-plasticity life-history reproduction oviposition nymph instar metamorphosis hemimetabolous incomplete-metamorphosis population-dynamics community-ecology food-web trophic-level primary-consumer ecosystem-function nutrient-cycling energy-flow biological-control pest-status agricultural-relevance forestry urban-ecology invasive-potential climate-change habitat-loss conservation-status data-gap research-need taxonomic-revision phylogenetics molecular-systematics DNA-barcoding integrative-taxonomy natural-history-collection specimen-digitization open-data FAIR-data biodiversity-informatics biological-database taxonomic-authority nomenclature zoological-nomenclature International-Code-of-Zoological-Nomenclature type-locality type-specimen-deposition author-citation original-combination current-combination synonymy homonymy nomenclatural-act taxonomic-history classification-system phylogenetic-classification evolutionary-relationships monophyly clade lineage stem-group crown-group divergence-time molecular-clock fossil-record paleontology extant extinct living-fossil evolutionary-history biogeographic-history dispersal vicariance range-expansion range-contraction refugium glacial-relict postglacial-colonization anthropogenic-impact human-disturbance land-use-change urbanization agricultural-intensification pesticide-use habitat-fragmentation connectivity corridor protected-area conservation-planning systematic-conservation-planning priority-area hotspot endemic-area center-of-origin center-of-diversity evolutionary-significant-unit management-unit population metapopulation source-sink-dynamics demography vital-rates survival fecundity generation-time population-growth-rate carrying-capacity density-dependence regulation limiting-factor environmental-stochasticity demographic-stochasticity genetic-stochasticity inbreeding-depression outbreeding-depression genetic-diversity heterozygosity allelic-richness effective-population-size genetic-drift gene-flow migration isolation-by-distance isolation-by-resistance landscape-genetics population-genomics phylogeography historical-demography coalescent-theory approximate-Bayesian-computation species-distribution-model ecological-niche-model habitat-suitability bioclimatic-envelope climate-envelope niche-conservatism niche-evolution realized-niche fundamental-niche biotic-interaction competition predation parasitism mutualism commensalism amensalism facilitation apparent-competition indirect-interaction trophic-cascade top-down-control bottom-up-control mesopredator-release keystone-species ecosystem-engineer foundation-species dominant-species characteristic-species indicator-species umbrella-species flagship-species surrogate-species focal-species target-species pest-species invasive-species naturalized-species alien-species non-native-species introduced-species cryptogenic-species range-shift poleward-shift elevational-shift phenological-shift temporal-mismatch trophic-mismatch phenology seasonality voltinism univoltine bivoltine multivoltine partial-generation prolonged-diapause aestivation hibernation overwintering overwintering-strategy cold-hardiness freeze-tolerance freeze-avoidance supercooling cryoprotectant thermal-biology thermoregulation behavioral-thermoregulation microclimate-selection habitat-selection oviposition-site-selection host-selection mate-choice sexual-selection sperm-competition cryptic-female-choice reproductive-isolation prezygotic-isolation postzygotic-isolation speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation reinforcement hybridization introgression hybrid-zone cline ecotone contact-zone secondary-contact exogenous-selection endogenous-selection ecological-speciation mutation-order-speciation sexual-conflict sexual-dimorphism sexual-size-dimorphism sexual-dichromatism polymorphism color-polymorphism morph phase gregarious-phase solitary-phase density-dependent-polyphenism reaction-norm genotype-environment-interaction plasticity canalization genetic-assimilation acclimation acclimatization hardening stress-response heat-shock-protein oxidative-stress hypoxia desiccation starvation immune-response innate-immunity humoral-immunity cellular-immunity encapsulation nodulation phagocytosis melanization antimicrobial-peptide phenoloxidase prophenoloxidase lysozyme defensin cecropin diptericin attacin immune-gene immune-pathway Toll-pathway Imd-pathway JAK-STAT-pathway NF-κB relish dorsal immune-memory specific-immunity transgenerational-immune-priming gut-microbiome symbiosis obligate-symbiont facultative-symbiont primary-symbiont secondary-symbiont vertical-transmission horizontal-transmission mixed-mode-transmission nutritional-symbiosis defensive-symbiosis digestive-symbiosis detoxification metabolism biosynthesis vitamin-synthesis amino-acid-synthesis steroid-synthesis cuticle-synthesis chitin-synthesis molting ecdysis ecdysteroid juvenile-hormone insulin-signaling TOR-signaling nutrient-sensing energy-storage lipid-metabolism carbohydrate-metabolism protein-metabolism flight-metabolism respiration tracheal-system spiracle gas-exchange water-balance osmoregulation excretion Malpighian-tubule hindgut rectum cryptonephridial-system ion-transport water-reabsorption nitrogenous-waste uric-acid urea ammonia detoxification-enzyme cytochrome-P450 glutathione-S-transferase carboxylesterase mixed-function-oxidase resistance insecticide-resistance acaricide-resistance herbicide-resistance antibiotic-resistance resistance-management resistance-evolution resistance-mechanism target-site-resistance metabolic-resistance behavioral-resistance cuticular-resistance resistance-allele resistance-gene fitness-cost fitness-benefit pleiotropy trade-off life-history-trade-off allocation resource-allocation energy-allocation time-allocation foraging optimal-foraging central-place-foraging patch-use marginal-value-theorem predator-avoidance antipredator-defense masquerade Batesian-mimicry Müllerian-mimicry automimicry intraspecific-mimicry intersexual-mimicry aggressive-mimicry social-parasitism brood-parasitism kleptoparasitism defense-behavior thanatosis death-feigning startle-display deimatic-display eyespot deflection-mark autotomy reflex-bleeding regurgitation defensive-secretion venom toxin allomone kairomone synomone semiochemical pheromone sex-pheromone aggregation-pheromone alarm-pheromone trail-pheromone territorial-pheromone recognition-pheromone signature-mixture cuticular-hydrocarbon volatile-organic-compound olfaction gustation mechanoreception thermoreception hygroreception photoreception vision compound-eye ommatidium rhabdom visual-pigment opsin color-vision polarization-vision motion-detection optomotor-response phototaxis photoperiodism circadian-rhythm biological-clock entrainment zeitgeber circannual-rhythm photoperiod day-length critical-day-length critical-photoperiod diapause-induction diapause-termination diapause-maintenance diapause-development endocrine-control neuroendocrine corpus-allatum corpus-cardiacum prothoracic-gland ecdysiotropin ecdysiostatin allatostatin allatotropin neuropeptide hormone hormone-receptor signal-transduction gene-expression transcription-factor regulatory-gene structural-gene gene-network gene-regulatory-network developmental-gene Hox-gene homeobox segment-polarity-gene gap-gene pair-rule-gene maternal-effect-gene zygotic-gene embryonic-development embryogenesis cleavage blastoderm gastrulation segmentation tagmosis appendage-development imaginal-disc holometabolous complete-metamorphosis larva pupa adult apolysis teneral sclerotization tanning cuticle exoskeleton epicuticle procuticle exocuticle endocuticle resilin sclerotin chitin protein lipid wax cement-layer wax-layer surface-lipid waterproofing anti-wetting anti-fouling self-cleaning structural-color pigmentary-color ommin pteridine carotenoid melanin papiliochrome flavonoid anthocyanin betalain iridescence photonic-crystal diffraction-grating thin-film-interference multilayer-reflector scattering Tyndall-scattering Rayleigh-scattering Mie-scattering non-iridescent-structural-color biochrome biofluorescence bioluminescence luciferin luciferase photoprotein bioluminescent-system counter-illumination burglar-alarm lure communication mate-recognition predator-deterrence aposematism warning-coloration honest-signal dishonest-signal signal-evolution signal-design receiver-psychology sensory-bias sensory-exploitation sensory-trap amplifier attenuator signal-efficacy signal-honesty handicap-principle index-signal conventional-signal assessment-signal territorial-signal dominance-signal submissive-signal aggressive-signal courtship-signal copulatory-signal postcopulatory-signal antagonistic-coevolution chase-away-selection sensory-drive reproductive-character-displacement genetic-exchange admixture hybrid-fitness hybrid-breakdown hybrid-vigor heterosis genetic-load mutational-meltdown founder-effect bottleneck population-expansion invasion-genetics propagule-pressure introduction-effort establishment spread lag-phase exponential-growth logistic-growth Allee-effect demographic-Allee-effect component-Allee-effect strong-Allee-effect weak-Allee-effect critical-population-size minimum-viable-population extinction-risk population-viability-analysis stochastic-model deterministic-model individual-based-model agent-based-model spatially-explicit-model metapopulation-model source-sink-model integrated-population-model state-space-model hierarchical-model Bayesian-model machine-learning artificial-intelligence computer-vision image-recognition automated-identification deep-learning neural-network convolutional-neural-network transfer-learning data-augmentation training-set validation-set test-set accuracy precision recall F1-score confusion-matrix receiver-operating-characteristic area-under-curve model-selection model-validation cross-validation bootstrapping permutation-test null-model randomization-test Monte-Carlo-simulation Markov-chain-Monte-Carlo Gibbs-sampling Metropolis-Hastings-algorithm Bayesian-inference prior-distribution posterior-distribution likelihood-function evidence model-comparison Bayes-factor information-criterion Akaike-information-criterion corrected-AIC Bayesian-information-criterion deviance-information-criterion Watanabe-Akaike-information-criterion cross-validation-information-criterion stacking model-averaging ensemble-model prediction forecasting scenario-analysis sensitivity-analysis uncertainty-analysis propagation-of-uncertainty error-budget confidence-interval credible-interval prediction-interval tolerance-interval statistical-power sample-size effect-size significance-test hypothesis-testing null-hypothesis alternative-hypothesis type-I-error type-II-error false-positive false-negative multiple-comparison Bonferroni-correction false-discovery-rate q-value p-value significance-level alpha-level beta-level statistical-test parametric-test nonparametric-test robust-test exact-test asymptotic-test bootstrap-test jackknife resampling subsampling leave-one-out k-fold stratified-sampling balanced-sampling imbalanced-data class-imbalance oversampling undersampling synthetic-minority-oversampling SMOTE ADASYN cost-sensitive-learning ensemble-method bagging boosting random-forest gradient-boosting XGBoost LightGBM CatBoost support-vector-machine kernel-method radial-basis-function polynomial-kernel linear-kernel sigmoid-kernel decision-tree classification-tree regression-tree pruning cost-complexity-pruning minimum-description-length rule-based-classifier association-rule frequent-pattern apriori-algorithm FP-growth sequential-pattern temporal-pattern spatial-pattern spatiotemporal-pattern time-series-analysis trend-analysis seasonal-decomposition autoregressive-model moving-average-model ARIMA SARIMA exponential-smoothing state-space-model Kalman-filter particle-filter hidden-Markov-model dynamic-Bayesian-network recurrent-neural-network long-short-term-memory gated-recurrent-unit transformer attention-mechanism self-attention multi-head-attention encoder decoder encoder-decoder-architecture sequence-to-sequence-model generative-model generative-adversarial-network variational-autoencoder normalizing-flow diffusion-model energy-based-model Boltzmann-machine restricted-Boltzmann-machine deep-belief-network autoencoder denoising-autoencoder sparse-autoencoder contractive-autoencoder variational-inference expectation-maximization maximum-likelihood maximum-a-posteriori maximum-entropy Occam's-razor parsimony model-complexity overfitting underfitting bias-variance-tradeoff regularization L1-regularization L2-regularization elastic-net dropout early-stopping batch-normalization layer-normalization group-normalization instance-normalization weight-decay mixup cutout cutmix random-erasing autoaugment fast-autoaugment population-based-augmentation neural-architecture-search automated-machine-learning hyperparameter-optimization grid-search random-search Bayesian-optimization sequential-model-based-optimization multi-fidelity-optimization knowledge-distillation teacher-student-model model-compression quantization knowledge-transfer domain-adaptation few-shot-learning zero-shot-learning one-shot-learning meta-learning learning-to-learn metric-learning siamese-network triplet-loss contrastive-learning self-supervised-learning unsupervised-learning semi-supervised-learning weakly-supervised-learning active-learning online-learning incremental-learning continual-learning lifelong-learning curriculum-learning multi-task-learning multi-modal-learning cross-modal-learning representation-learning feature-learning embedding-learning manifold-learning dimensionality-reduction principal-component-analysis t-SNE UMAP word-embedding sentence-embedding document-embedding knowledge-graph-embedding graph-neural-network graph-convolutional-network graph-attention-network message-passing-neural-network relational-graph-convolutional-network inductive-learning transductive-learning homophily heterophily graph-isomorphism Weisfeiler-Lehman-test subgraph-isomorphism graph-matching network-alignment network-embedding node-embedding edge-embedding graph-classification node-classification link-prediction graph-generation graph-synthesis molecular-generation de-novo-design inverse-design retrosynthesis reaction-prediction property-prediction quantitative-structure-activity-relationship QSAR quantitative-structure-property-relationship QSPR molecular-descriptor fingerprint Morgan-fingerprint extended-connectivity-fingerprint circular-fingerprint atom-pair-fingerprint topological-torsion-fingerprint pharmacophore shape-similarity electrostatic-similarity docking scoring-function binding-affinity-prediction protein-ligand-interaction protein-protein-interaction protein-structure-prediction AlphaFold protein-folding contact-prediction distance-prediction angle-prediction torsion-prediction secondary-structure-prediction tertiary-structure-prediction quaternary-structure-prediction protein-design protein-engineering directed-evolution rational-design computational-design de-novo-protein-design enzyme-design catalytic-antibody ribozyme aptamer DNAzyme synthetic-biology genetic-circuit metabolic-engineering metabolic-flux-analysis flux-balance-analysis genome-scale-metabolic-model dynamic-metabolic-model kinetic-modeling stoichiometric-modeling constraint-based-modeling optimization linear-programming mixed-integer-linear-programming quadratic-programming nonlinear-programming dynamic-programming stochastic-programming robust-optimization multi-objective-optimization Pareto-optimality evolutionary-algorithm genetic-algorithm genetic-programming differential-evolution particle-swarm-optimization ant-colony-optimization simulated-annealing tabu-search variable-neighborhood-search iterated-local-search greedy-randomized-adaptive-search scatter-search path-relinking memetic-algorithm hyper-heuristic metaheuristic local-search neighborhood-search hill-climbing random-walk randomized-search Monte-Carlo-tree-search upper-confidence-bound Thompson-sampling multi-armed-bandit contextual-bandit reinforcement-learning policy-gradient actor-critic Q-learning deep-Q-network policy-iteration value-iteration temporal-difference-learning SARSA expected-SARSA double-Q-learning dueling-Q-network prioritized-experience-replay distributed-reinforcement-learning impala A3C asynchronous-advantage-actor-critic PPO proximal-policy-optimization TRPO trust-region-policy-optimization DDPG deep-deterministic-policy-gradient TD3 twin-delayed-deep-deterministic-policy-gradient SAC soft-actor-critic model-based-reinforcement-learning model-predictive-control guided-policy-search imitation-learning inverse-reinforcement-learning apprenticeship-learning behavior-cloning dataset-aggregation DAgger generative-adversarial-imitation-learning third-person-imitation-learning one-shot-imitation-learning meta-reinforcement-learning hierarchical-reinforcement-learning option skill subgoal intrinsic-motivation curiosity-driven-exploration empowerment information-gain prediction-error novelty-seeking diversity-seeking multi-agent-reinforcement-learning mean-field fictitious-play correlated-equilibrium Nash-equilibrium game-theory mechanism-design auction-theory incentive-design social-choice-theory voting-theory fair-division matching-theory market-design algorithmic-game-theory computational-social-choice computational-complexity NP-completeness NP-hardness approximation-algorithm approximation-ratio inapproximability fixed-parameter-tractability parameterized-complexity kernelization treewidth graph-minor forbidden-minor-characterization well-quasi-ordering Robertson-Seymour-theorem graph-structure-theorem bidimensionality-theory meta-theorem Courcelle's-theorem automata-theory finite-automaton pushdown-automaton Turing-machine computability decidability halting-problem Rice's-theorem Gödel's-incompleteness-theorems lambda-calculus combinatory-logic type-theory proof-theory model-theory set-theory forcing large-cardinal axiom-of-choice continuum-hypothesis independence-result consistency-strength reverse-mathematics constructive-mathematics intuitionistic-logic modal-logic temporal-logic epistemic-logic deontic-logic description-logic ontology knowledge-representation semantic-web linked-data RDF SPARQL OWL knowledge-graph entity-linking relation-extraction event-extraction coreference-resolution named-entity-recognition part-of-speech-tagging dependency-parsing constituency-parsing semantic-parsing natural-language-inference textual-entailment natural-language-generation machine-translation summarization question-answering dialogue-system conversational-AI chatbot virtual-assistant speech-recognition speech-synthesis speaker-recognition emotion-recognition sentiment-analysis opinion-mining aspect-based-sentiment-analysis target-dependent-sentiment-analysis stance-detection rumor-detection fake-news-detection fact-checking claim-verification evidence-retrieval argument-mining persuasion-detection deception-detection personality-prediction mental-health-detection suicide-prevention crisis-intervention social-media-mining network-analysis community-detection influence-maximization information-diffusion cascade-modeling epidemic-modeling SEIR-model SIR-model agent-based-epidemic-model compartmental-model network-epidemic-model metapopulation-epidemic-model spatial-epidemic-model stochastic-epidemic-model branching-process renewal-equation generation-interval serial-interval incubation-period latent-period infectious-period basic-reproduction-number effective-reproduction-number herd-immunity-threshold vaccination-coverage vaccine-efficacy vaccine-effectiveness breakthrough-infection waning-immunity immune-escape variant-of-concern variant-of-interest variant-under-monitoring phylogenetic-analysis molecular-clock-analysis phylodynamics viral-evolution antigenic-drift antigenic-shift zoonotic-spillover reservoir-host intermediate-host amplifying-host dead-end-host vector-borne-disease mosquito-borne-disease tick-borne-disease flea-borne-disease lice-borne-disease midge-borne-disease sandfly-borne-disease tsetse-borne-disease blackfly-borne-disease triatomine-borne-disease kissing-bug Chagas-disease trypanosomiasis leishmaniasis malaria dengue yellow-fever Zika-virus chikungunya West-Nile-virus Japanese-encephalitis tick-borne-encephalitis Lyme-disease Rocky-Mountain-spotted-fever ehlichiosis anaplasmosis babesiosis tularemia plague murine-typhus epidemic-typhus endemic-typhus scrub-typhus Q-fever trench-fever bartonellosis cat-scratch-disease rat-bite-fever leptospirosis brucellosis anthrax melioidosis glanders tularaemia francisellosis coxiellosis rickettsiosis orientia-infection ehrlichiosis neoehrlichiosis neorickettsiosis anaplasmataceae rickettsiaceae coxiellaceae bartonellaceae brucellaceae francisellaceae pasteurellaceae legionellaceae chlamydiaceae mycoplasmataceae ureaplasmataceae spiroplasmataceae achaenoplasmataceae entomoplasmataceae haloplasmataceae tenericutes mollicutes bacteroidetes fusobacteria spirochaetes synergistetes deferribacteres thermotogae aquificae caldiserica caldithrix deferrisoma desulfurobacterium thermovibrio thermocrinis hydrogenobacter sulfurihydrogenibium persephonella hydrogenobaculum hydrothermus thermothrix sulfuricurvum sulfurimonas sulfurovum thiovulum thiomicrospira pseudomonas vibrio aeromonas photobacterium aliivibrio gracilibacillus halobacillus halomonas chromohalobacter cobetia salinicola kushneria halotalea halovibrio salinivibrio thioalkalivibrio thiohalophilus ectothiorhodospira halorhodospira thiorhodospira halochromatium thioflavicoccus marichromatium thiocapsa lamprocystis thiodictyon thiopedia amoebobacter thiocystis chromatium allo-chromatium thermochromatium rhabdochromatium thiobaca thiohalocapsa halocapsa thiolamprovum thioploca thiomargarita beggiatoa methylococcus methylomonas methylomicrobium methylosinus methylocystis methylocella methylocapsa methylacidiphilum verrucomicrobia planctomycetes chlamydiae lentisphaerae kiritimatiellaeota calditrichaeota fervidibacteria
Stigmella rhoifoliella
Stigmella rhoifoliella is a minute leaf-mining moth in the family Nepticulidae, native to the eastern United States. The species is notable for its extremely small size and specialized larval feeding behavior on poison ivy (Toxicodendron radicans). It produces distinctive serpentine leaf mines on its host plant. The moth has a multivoltine life cycle with two to three generations annually, with adult flight periods concentrated in June and August.
leaf-miner microlepidoptera specialist-herbivore multivoltine Nearctic
Strephonota tephraeus
Pearly-gray Hairstreak
Strephonota tephraeus is a small lycaenid butterfly with a wingspan of 22–29 mm, commonly known as the Pearly-gray Hairstreak. The species ranges from Mexico through Central America to the Amazon basin and northern South America, with rare vagrant records in southern Texas. It inhabits lowland tropical forests, particularly riparian corridors, and exhibits multivoltine breeding in Mexico with adults active from May through December.
Lycaenidae hairstreak Neotropical riparian multivoltine vagrant
Syssphinx bicolor
honey locust moth
Syssphinx bicolor, commonly known as the honey locust moth, is a North American saturniid moth with variable wing coloration ranging from grayish to yellow, orange, or dark reddish brown. The species exhibits three annual broods with adults of different color morphs appearing sequentially—grayish first, then yellow to orange-brown, and finally darker with more spotting. Larvae feed on honey locust and Kentucky coffee tree, with distinctive color changes between gregarious young stages and solitary older instars.
Saturniidae Ceratocampinae North-America woodland legume-feeder multivoltine color-polymorphism underground-pupation
Tetanocera ferruginea
common buff snailkiller
Tetanocera ferruginea is a medium to large sciomyzid fly with light brown coloration and no obvious wing markings. It is an obligate malacophage, with larvae preying on non-operculate snails in aquatic and semi-aquatic habitats. The species has a Holarctic distribution and is multivoltine, overwintering in the puparial stage. Despite forming large populations, individuals exhibit very limited movement within habitats.
Diptera Sciomyzidae malacophagy aquatic snail-predator multivoltine Holarctic
Tetragnatha
Stretch Spiders, Long-jawed Orb Weavers
Tetragnatha is a large genus of long-jawed orb-weaving spiders with hundreds of species distributed worldwide, though most occur in tropical and subtropical regions. These spiders are characterized by their elongated bodies, extremely long legs, and notably elongated chelicerae (jaws) with well-developed teeth. They construct horizontal orb webs, often near water, and many species can run across water surfaces. The genus exhibits remarkable variation in dispersal ability, with some species being excellent dispersers capable of colonizing islands, while others appear to have secondarily lost this capacity. Hawaiian Tetragnatha species have undergone adaptive radiation, with some lineages evolving cursorial (running) behavior and abandoning web-building entirely.
spider orb-weaver long-jawed aquatic stretch-spider adaptive-radiation Hawaii cursorial-evolution horizontal-web water-walking camouflage multivoltine social-spider Tetragnathidae Araneae Arachnida predator insectivore riparian wetland meadow tropical subtropical cosmopolitan dispersal island-colonization phylogeography chemical-species-recognition communal-webbing kleptoparasitism-target
Tetragnathidae
Long-jawed Orb Weavers, Long-jawed Orbweavers
Long-jawed orb weavers are a family of spiders characterized by elongated bodies, long legs, and often extraordinarily long chelicerae (jaws). Most species construct orb webs in a horizontal or near-horizontal plane, distinguishing them from the vertical webs typical of Araneidae. The family includes diverse ecological specialists: meadow-dwelling species that camouflage as grass stems, riparian species that build webs over water, and cave-dwelling species adapted to dimly lit environments. Some species exhibit social behavior, forming communal webs spanning extensive areas.
spiders orb-weavers riparian cave-dwelling social-spiders camouflage multivoltine horizontal-webs
Tricorythodes
little stout crawler mayflies
Tricorythodes is a genus of small, robust mayflies in the family Leptohyphidae, commonly known as little stout crawler mayflies. The genus comprises approximately 16 described species distributed across the Americas. Nymphs are primarily found in stream habitats where they function as collector-gatherers, processing fine particulate organic matter. Species within this genus exhibit multivoltine life histories with development rates strongly influenced by temperature.
Ephemeroptera Leptohyphidae mayflies aquatic-insects stream-ecology collector-gatherers multivoltine Neotropical Nearctic
Trigonotylus
rice leaf bug
Trigonotylus is a genus of plant bugs in the family Miridae, tribe Stenodemini, containing approximately 18 described species. The genus has a cosmopolitan distribution with species in North America, Europe, Asia, and Hawaii. Several species are associated with grasses (Poaceae), with T. caelestialium being a significant agricultural pest of rice in Japan. The genus exhibits variation in host plant use, oviposition behavior, and seasonal life history strategies.
grass-feeder agricultural-pest multivoltine egg-diapause olfactory-orientation Japan rice-pest Poaceae Stenodemini bet-hedging
Trigonotylus caelestialium
Rice Leaf Bug
Trigonotylus caelestialium is a mirid plant bug known as the rice leaf bug, native to Europe, Northern Asia (excluding China), and North America. It is a multivoltine pest of rice in Japan, where it causes pecky-rice damage by feeding on developing grains. The species has a broad host range within Poaceae, utilizing various grasses as alternative hosts before invading rice paddies during the flowering stage.
agricultural-pest rice-pest multivoltine diapause olfactory-orientation Poaceae Japan endophyte-resistance bet-hedging-strategy
Uresiphita reversalis
Genista Broom Moth, Sophora Worm
Uresiphita reversalis is a multivoltine crambid moth native to Mexico and the southwestern United States that has expanded its range north and east across North America. The caterpillars feed diurnally in groups on leguminous host plants, particularly members of the tribe Genisteae, and sequester quinolizidine alkaloids for chemical defense. The species has gained notoriety as both a pest of ornamental plants and a potential biocontrol agent for invasive broom species. Adults are small moths with distinctive white bodies and bright yellow or orange hindwings.
multivoltine aposematic alkaloid-sequestration diurnal-larvae group-feeding legume-feeder biocontrol-potential range-expansion pest-of-ornamentals chemical-defense quinolizidine-alkaloids carrier-mediated-uptake
Urophora affinis
Knapweed Gall Fly
Urophora affinis is a tephritid fruit fly native to Eurasia that was introduced to North America as a biological control agent for spotted knapweed (Centaurea maculosa) and diffuse knapweed (C. diffusa). The species forms galls in knapweed seed heads, where larvae develop and reduce seed production by up to 95%. First released in Montana in 1973, it has established populations but remains concentrated near release sites with limited dispersal ability. The fly is multivoltine, with adults emerging in June and larvae overwintering in seed heads.
biocontrol invasive-species-management gall-former knapweed Tephritidae multivoltine interspecific-competition
Virbia opella
tawny holomelina, Tawny Virbia Moth
Virbia opella is a small tiger moth in the family Erebidae, historically known as the tawny holomelina. It was described by Grote in 1863 and later transferred from Holomelina to Virbia following phylogenetic revision. The species occurs in eastern and central North America, where it inhabits oak-dominated forests and produces multiple generations annually.
moth tiger-moth oak-forest multivoltine North-America
Wallengrenia otho
southern broken dash, broken dash skipper
Wallengrenia otho is a small skipper butterfly in the family Hesperiidae, distributed across the Americas from the southeastern United States through Central America to Argentina. The species exhibits multivoltine flight patterns in temperate regions and year-round activity in tropical areas. Larvae develop on grasses in the genera Paspalum and Stenotaphrum, while adults feed on nectar from wetland and riparian flowers.
skipper grass-skipper multivoltine nectar-feeder grassland wetland