Biological-control
Guides
Anacharitinae
Anacharitinae is a subfamily of small parasitoid wasps within Figitidae. Members are known to attack aphid predators, specifically larvae of Syrphidae (hoverflies) and Chrysopidae (lacewings). This subfamily comprises approximately 100 species across four genera: Anacharis, Aegilips, Callaspidia, and Xyalaspis. The group has a worldwide distribution with greatest diversity in temperate regions.
Anacrabro ocellatus
square-headed wasp
Anacrabro ocellatus is a small (6-7 mm) fossorial wasp in the family Crabronidae. Unlike most related genera in tribe Crabronini that hunt flies, this species specializes in capturing plant bugs (family Miridae) to provision its subterranean nest cells. It is widespread east of the Rocky Mountains in North America, with three recognized subspecies. The species is notable for its potential role in biological control of agricultural pests.
Anagrapha
Anagrapha is a genus of moths in the family Noctuidae, subfamily Plusiinae, established by McDunnough in 1944. The genus contains at least one confirmed species, Anagrapha falcifera (celery looper), which serves as the original host for AfMNPV (Anagrapha falcifera multiple nucleopolyhedrovirus), a broad host-spectrum baculovirus with significant importance in biological control research. The virus isolated from this species has been shown to infect 43 species across 13 lepidopteran families and is genetically identical to the Rachiplusia ou virus isolate.
Anagrapha falcifera
Celery Looper, Celery Looper Moth
Anagrapha falcifera, the celery looper, is a noctuid moth native to North America. The species was first described by William Kirby in 1837. Adults have a wingspan of 35–40 mm. Larvae are generalist feeders on low-growing plants including agricultural crops. The species gained scientific significance in 1985 when specimens collected in Missouri were found infected with a naturally occurring baculovirus, which became the first patented baculovirus and contributed to the development of biological insect control products.
Anagrus
fairyflies
Anagrus is a genus of fairyflies (Mymaridae) comprising over 90 described species, many of which are important biological control agents of agricultural pests. Adults are minute parasitoids, typically 1-2 mm in length, with coloration ranging from brown and orange to black and pale. The genus is divided into three subgenera—Anagrella, Anagrus, and Paranagrus—for taxonomic purposes. Species in this genus are specialized egg parasitoids, primarily attacking Hemiptera (leafhoppers and planthoppers), with some species also recorded from Odonata eggs. Their role in regulating populations of economically significant pests like the brown planthopper and glassy-winged sharpshooter has made them valuable in integrated pest management programs worldwide.
Anagyrus
Anagyrus is a large genus of parasitic wasps in the family Encyrtidae, containing at least 247 described species distributed worldwide. These wasps are specialized parasitoids primarily targeting mealybugs (Hemiptera: Pseudococcidae), with several species commercially important for biological control of agricultural pests. The genus includes notable species such as A. pseudococci, A. lopezi, and A. vladimiri, which have been extensively studied and deployed for managing mealybug infestations in crops including cassava, citrus, and grapes. A subgenus, Nesoanagyrus, was established by Beardsley in 1969.
Anagyrus aper
Anagyrus aper is a species of parasitoid wasp in the family Encyrtidae, described by Noyes & Menezes in 2000. It belongs to the genus Anagyrus, a group of wasps known primarily as parasitoids of mealybugs (Hemiptera: Pseudococcidae). The species has been recorded from New Hampshire, Ontario, and Texas, suggesting a Nearctic distribution. Like other members of its genus, it likely functions as a biological control agent of pest mealybugs, though specific host associations and biological details remain poorly documented in the available literature.
Anagyrus californicus
Anagyrus californicus is a small parasitoid wasp in the family Encyrtidae, originally described by Compere in 1947 as Apoanagyrus californicus. It is a specialized parasitoid of mealybugs (Hemiptera: Pseudococcidae), with documented records from California, Arizona, Florida, and Hawaii. The species has been utilized in biological control programs targeting mealybug pests in agricultural systems.
Anagyrus paralia
Anagyrus paralia is a species of parasitoid wasp in the family Encyrtidae, described by Noyes and Menezes in 2000. It belongs to the genus Anagyrus, which contains numerous species used in biological control of mealybugs and other scale insects. The specific epithet "paralia" suggests a coastal or shore-associated habitat, though detailed ecological data remain limited. Like other Anagyrus species, it is presumed to be a primary parasitoid of mealybugs (Pseudococcidae), but host records specific to this species have not been documented in the available literature.
Anagyrus pulchricornis
Anagyrus pulchricornis is a species of parasitoid wasp in the family Encyrtidae, first described by Howard in 1894. It belongs to a genus widely used in biological control programs targeting mealybugs (Pseudococcidae). The species is known from the Caribbean region, with records from Jamaica, the Grenadines, and Saint Vincent and the Grenadines. Like other members of Anagyrus, it likely parasitizes mealybug hosts, though specific host associations for this species remain poorly documented in the available literature.
Anagyrus shahidi
Anagyrus shahidi is a species of parasitoid wasp in the family Encyrtidae, first described by Hayat in 1979. It belongs to a genus widely used in biological control programs targeting mealybugs (Hemiptera: Pseudococcidae). The species has been recorded from India (Andhra Pradesh, Karnataka, Rajasthan), Florida (USA), and Sulawesi Tengah (Indonesia). Like other Anagyrus species, it likely functions as a primary parasitoid of mealybugs, though specific host associations for A. shahidi remain incompletely documented.
Anagyrus trinidadensis
Anagyrus trinidadensis is a small parasitoid wasp in the family Encyrtidae, originally described from Trinidad. The species belongs to a genus of significant economic importance for biological control of mealybugs and other scale insects. Like congeners, it likely functions as a primary parasitoid of Pseudococcidae, though specific host records for this species remain limited in the available literature.
Anaphes
fairyflies
Anaphes is a genus of fairyflies (family Mymaridae), among the smallest known insects, first described by Alexander Henry Haliday in 1833. The genus has a cosmopolitan distribution and is divided into two subgenera: Anaphes (Anaphes) with approximately 37 recognized species and Anaphes (Patasson) with approximately 56 species in the Palearctic region alone. Species are egg parasitoids of various insects, with documented hosts including weevils (Curculionidae), leafhoppers (Cicadellidae), and true bugs (Miridae). Several species have been studied for biological control of agricultural pests, particularly the carrot weevil (Listronotus oregonensis) and lygus bugs (Lygus spp.).
Anasa
squash bugs
Anasa is a genus of squash bugs in the family Coreidae, containing more than 70 described species distributed throughout North, Central, and South America. The genus includes major agricultural pests, particularly Anasa tristis, which causes significant damage to cucurbit crops and vectors cucurbit yellow vine disease. Members of this genus are specialized herbivores of plants in the family Cucurbitaceae.
Anastatus
eupelmid egg parasitoids
Anastatus is a large genus of chalcidoid wasps in the family Eupelmidae, comprising approximately 160 species globally with about 18 species recorded from North America north of Mexico. All species are egg parasitoids, attacking the eggs of various insects including true bugs, moths, and other arthropods. The genus exhibits extreme sexual dimorphism: females are brachypterous or apterous with powerful jumping abilities, while males are fully winged and capable of flight. Several species have been introduced as biological control agents, including A. disparis for spongy moth management and A. orientalis for spotted lanternfly control.
Anastatus semiflavidus
Anastatus semiflavidus is a minute egg parasitoid wasp in the family Eupelmidae. It is a specialist parasitoid of the range caterpillar (Hemileuca oliviae), a pest of rangeland grasses in western North America. The species exhibits arrhenotokous parthenogenesis, with females developing from fertilized eggs and males from unfertilized eggs. Laboratory studies demonstrate strong temperature-dependence in its development and reproduction.
Anastoechus barbatus
Anastoechus barbatus is a species of bee fly in the family Bombyliidae, distributed across much of North America. The species occurs from the Yukon east to Ontario in Canada, and across most of the United States from Massachusetts west to California and Texas, though it is absent from the southeastern United States. Its larvae are predators of grasshopper eggs, representing an important biological control mechanism.
Anastrepha
fruit flies, tephritid fruit flies
Anastrepha is a highly diverse genus of tephritid fruit flies native to the American tropics and subtropics, comprising over 300 described species. The genus includes nine major agricultural pest species that inflict substantial damage on commercial fruit crops including citrus, mango, guava, and papaya. Species exhibit variation in oviposition behavior, with females depositing eggs in developing fruit, mature fruit, or seeds depending on the species. The genus has been reorganized taxonomically to include species formerly placed in Toxotrypana, and contains notable cryptic species complexes such as the Anastrepha fraterculus group.
Anatis
Giant Lady Beetles
Anatis is a genus of ladybird beetles (Coccinellidae) commonly known as "giant lady beetles." The genus contains approximately ten extant species distributed primarily in North America, with at least two fossil species described. Members are relatively large compared to most lady beetles and are predatory as adults and larvae. The genus includes notable species such as Anatis mali and Anatis labiculata, which have been studied for their biological control potential.
Anatis mali
Eye-spotted Lady Beetle
Anatis mali, commonly known as the eye-spotted lady beetle, is a North American lady beetle species in the family Coccinellidae. Adults measure 7.3–10 mm in length and display variable coloration from yellow to brownish red with black markings. The species has been documented as a specialized predator of aphids in balsam fir plantations, where it plays a significant role in biological control. Research indicates that larvae require approximately 296 aphids to complete development and pupate.
Anaxipha
Brown Sword-tail Crickets
Anaxipha is a genus of small swordtail crickets distributed across tropical and subtropical regions worldwide, including the Americas, Africa, Asia, Australia, and western Pacific islands. Members are known as "brown sword-tail crickets" and are characterized by their acoustic communication systems, which in several studied species include both calling songs and previously undocumented courtship songs with substrate-borne vibrational signals (drumming). Some species occupy agricultural habitats such as rice fields, where they function as generalist predators.
Ancistrocerus
mason wasps, potter wasps
Ancistrocerus is a genus of solitary mason wasps in the subfamily Eumeninae. Females construct nests using mud, either in pre-existing cavities or as free-form mud cells attached to hard surfaces. They provision nests with paralyzed caterpillars as food for their larvae. Males are distinguished by hooked antennae tips. The genus has a nearly worldwide distribution, with notable species in North America, Europe, and Asia.
Ancylis
Ancylis is a genus of tortricid moths in the subfamily Olethreutinae, tribe Enarmoniini. Species are commonly known as leafrollers or leaf folders, with larvae that construct silk shelters by folding or rolling host plant leaves. The genus has a Holarctic distribution with some species also recorded from the Oriental region and South America. Several species are economically significant agricultural pests, including the strawberry leafroller (A. comptana) and the jujube leafroller (A. sativa). Taxonomic revision using DNA barcoding has clarified species boundaries, revealing that presumed Holarctic distributions were often due to misidentification.
Anisodactylus carbonarius
Anisodactylus carbonarius is a ground beetle in the family Carabidae, native to North America. It is a granivorous species that feeds on weed seeds, contributing to natural weed suppression in agricultural and garden settings. The species has been documented across Canada and the United States. Like other members of its genus, it plays a role in ecosystem services through seed predation.
Anisolabididae
Anisolabididae is a family of earwigs (order Dermaptera, suborder Neodermaptera) comprising 38 genera across 13 subfamilies. Members include both cosmopolitan and geographically restricted species, with some showing specialized adaptations for subcortical or coastal habitats. Several genera, particularly Euborellia, are recognized for their predatory behavior and potential as biological control agents of agricultural pests. The family exhibits diverse morphologies ranging from typical earwig body plans to dorsoventrally flattened forms adapted for life under bark.
Anisopteromalus
Genus of small parasitoid wasps in the family Pteromalidae, comprising species that attack larvae of stored-product beetles. Species within this genus are ectoparasitoids, with females laying eggs on host larvae concealed inside grains or other substrates. The genus includes at least two morphologically similar sibling species, A. calandrae and A. quinarius, that require molecular or cytogenetic methods for reliable identification. Members of this genus have been investigated as biological control agents against pests of stored grain.
Anomaloninae
Anomaloninae is a cosmopolitan subfamily of parasitoid wasps in the family Ichneumonidae. Members are slender, ranging from small to large, and typically black or brownish with yellow markings on the face and legs. The subfamily is divided into two tribes: Anomalonini, which parasitize Coleoptera larvae (Tenebrionidae and Elateridae) and some Lepidoptera larvae (Noctuidae and Tortricidae), and Gravenhorstiini, which parasitize Lepidoptera larvae. They are distinguished from other ichneumonid subfamilies by a highly reticulated propodeum, slender metasoma, fore wing with vein 3 rs-m absent, and swollen hind tarsi in males.
Anomalonini
Anomalonini is a tribe of parasitoid wasps in the family Ichneumonidae, subfamily Anomaloninae. The tribe is dominated by the large genus *Anomalon*, with *Neogreeneia* sometimes recognized as a second genus. Members are koinobiont endoparasitoids of lepidopteran larvae. The tribe contains approximately 5,600+ documented observations, indicating moderate research attention.
Anthicidae
Ant-like Flower Beetles, Ant-like Beetles
Anthicidae is a family of beetles comprising over 3,500 species in approximately 100 genera. Members are commonly known as ant-like flower beetles due to their striking morphological resemblance to ants. The family is characterized by a constricted head forming a distinct neck and slender appendages that enhance the ant-mimicry. Adults are omnivorous, feeding on small arthropods, pollen, fungi, and other available materials. The family has fossil representatives dating to the Early Cretaceous (Barremian) from Lebanese amber.
Anthocoris
minute pirate bugs
Anthocoris is a genus of minute pirate bugs in the family Anthocoridae, containing at least 30 described species. These small predatory true bugs are important biological control agents in orchards and natural ecosystems, feeding on aphids, psyllids, mites, and other small arthropods. Several species have been studied extensively for their potential in integrated pest management, particularly in pear orchards and pistachio cultivation. The genus exhibits host-specific associations with particular tree species and prey types, with life cycles often synchronized to exploit seasonal aphid or psyllid populations.
Anthonomus
Anthonomus is a large genus of weevils (Curculionidae) described by Germar in 1817. The genus includes numerous species, many of significant economic importance. Several members are major agricultural pests, including the cotton boll weevil (A. grandis), strawberry blossom weevil (A. signatus), and pepper weevil (A. eugenii). Conversely, some species such as A. santacruzi are utilized as biological control agents against invasive plants. The taxonomy of the genus remains under revision, particularly regarding the status of Cnemocyllus as either a distinct genus or subgenus.
Anthonomus eugenii
Pepper Weevil
Anthonomus eugenii, the pepper weevil, is a small curculionid beetle and major agricultural pest of cultivated peppers (Capsicum spp.). Native to Mexico, it has spread throughout Central America, the Caribbean, the southern United States, and has been intercepted in more northern regions including Canada and Europe through imported infested fruit. The species completes its entire immature development within pepper fruits, making it particularly difficult to control with conventional insecticides. Females exhibit host-marking behavior, depositing pheromones that deter subsequent oviposition by conspecifics.
Anthrax georgicus
Tiger Bee Fly
Anthrax georgicus is a bee fly species in the family Bombyliidae, distributed across eastern and central North America. Adults are active in spring and are known for their parasitoid relationship with tiger beetle larvae (Cicindelidae), specifically attaching externally to host larvae and consuming them as they develop. The species was previously known as Anthrax analis but is now recognized as A. georgicus based on taxonomic revision.
Anthrax irroratus
Spotted Bee Fly
Anthrax irroratus is a bee fly (family Bombyliidae) widespread across North America, ranging from Alaska and northern Canada to Central America and the Caribbean. Adults are pollinators that frequent flowers, while larvae are parasitoids of solitary bees and wasps in at least five families. The species is one of numerous bee flies that exploit ground-nesting Hymenoptera as hosts.
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whirligig mites
Anystidae is a family of predatory mites comprising over 100 species, commonly known as whirligig mites. They are active generalist predators found in soils and on vegetation worldwide. The family includes notable genera such as Anystis, Tarsotomus, and Chaussieria. Several species have been investigated for biological control potential against agricultural pests including thrips, aphids, and spider mites.
Anystinae
Anystinae is a subfamily of prostigmatic mites in the family Anystidae. These fast-moving mites are commonly known as whirligig mites due to their rapid, erratic movement patterns. Members of this subfamily are primarily predatory and are frequently observed in terrestrial habitats. The subfamily includes the well-known genus *Anystis*, which contains species that are commercially used for biological pest control.
Anystis
Whirligig Mites
Anystis is a genus of predatory mites in the family Anystidae, commonly known as whirligig mites. Species in this genus are generalist predators of small arthropods, including mites, aphids, thrips, and other soft-bodied insects. They are frequently red in coloration with long legs, ranging from 500–1500 μm in size. The genus has gained attention for biological control applications, particularly Anystis baccarum, which has been documented in agricultural systems including apple orchards, raspberry crops, vineyards, and greenhouses.
Anystis baccarum
Whirligig mite
Anystis baccarum is a cosmopolitan predatory mite in the family Anystidae, commonly known as the whirligig mite. It is a generalist predator of small, soft-bodied arthropods including aphids, spider mites, thrips, and other pests in agricultural systems. Populations consist entirely of females that reproduce by thelytokous parthenogenesis. The species has been identified as a promising biological control agent in crops such as raspberry, apple, sweet pepper, and tea, though its cannibalistic tendencies and long generation time limit its effectiveness as a sole control method.
Aonidiella
Aonidiella is a genus of armored scale insects (family Diaspididae) containing approximately 30 described species. Several species, notably A. aurantii (California red scale) and A. citrina (yellow scale), are significant agricultural pests of citrus and other fruit crops. The genus exhibits a global distribution with highest diversity in tropical and subtropical regions where host plants occur.
Aonidiella aurantii
California red scale, red scale
Aonidiella aurantii, commonly known as California red scale or red scale, is an armored scale insect and major agricultural pest of citrus and other crops. Native to South China, it has spread globally through human-mediated transport of infected plant material, first appearing in California between 1868 and 1875. The species is now established across six continents and infests over 77 plant families, with citrus remaining its primary economic host. Heavy infestations cause fruit pitting, shoot distortion, leaf drop, and tree death.
Apanteles
Apanteles is a large genus of braconid wasps in the subfamily Microgastrinae, containing over 600 described species worldwide. A 2014 study from Costa Rica's Area de Conservación Guanacaste alone documented 186 new species, suggesting the genus is far more diverse than previously recognized. These small wasps (1–5 mm) are koinobiont endoparasitoids of caterpillars, with most species showing high host specificity.
Apanteles megathymi
Apanteles megathymi is a species of parasitoid wasp in the family Braconidae, first described by Riley in 1881. It belongs to the genus Apanteles, a large and diverse group of microgastrine braconids that are important biological control agents. The species name 'megathymi' suggests a historical association with the genus Megathymus (giant skippers), though current host records require verification. Like other Apanteles species, it is likely a larval parasitoid of Lepidoptera.
Apertochrysa
Apertochrysa is a genus of green lacewings in the family Chrysopidae, comprising 183 described species. The genus has undergone substantial taxonomic revision, having been previously classified under the names Navasius, Dichochrysa, and Pseudomallada before being synonymized under Apertochrysa in 2021. It is a subcosmopolitan genus with the majority of species distributed in the Palearctic region, particularly Asia and Africa. Species in this genus are predators of soft-bodied insects and some exhibit debris-carrying behavior as larvae.
Aphaereta
Aphaereta is a cosmopolitan genus of braconid wasps in the tribe Alysiini, comprising approximately 48 described species worldwide. Members are endoparasitoids of Diptera, with documented associations including dung-breeding flies, blow flies, shore flies, and fruit flies. The genus has been recorded across multiple continents including Europe, Asia, North America, South America, and Oceania. Several species have been studied for their potential in biological control programs targeting pest flies.
Aphalara
jumping plant lice, psyllids
Aphalara is a genus of jumping plant lice (psyllids) in the family Aphalaridae and tribe Aphalarini. The genus contains approximately 37 recognized valid species distributed across the Palearctic, Nearctic, and Neotropical regions. Many species are specialized herbivores of Polygonaceae, particularly Polygonum and Rumex, with some groups showing strict host associations. The genus includes A. itadori, a widely studied biological control agent for invasive knotweeds (Reynoutria/Fallopia spp.) in Europe and North America. Species exhibit diverse biologies including gall induction on host plants and vibrational communication during mate search.
Aphelinidae
Aphelinidae is a family of minute parasitic wasps containing approximately 1,100-1,160 described species across 34-35 genera. These tiny insects are among the most important biological control agents used in agriculture, particularly for managing scale insects, whiteflies, and other Hemiptera. The family was formerly paraphyletic, leading to the recent elevation of Azotidae, Calesidae, and Eriaporidae to family rank. Specimens deteriorate rapidly after death unless preserved in ethanol, making museum identification challenging.
Aphelininae
Aphelininae is a subfamily of minute parasitic wasps in the family Aphelinidae (order Hymenoptera). Members are primarily parasitoids of scale insects, whiteflies, and other Hemiptera, with some species serving as important biological control agents. The subfamily was established by Thomson in 1876. Some genera historically placed in Aphelininae have been subject to taxonomic revision, with certain species formerly classified here now reassigned to other families such as Eulophidae.
Aphelinoidea
Aphelinoidea is a genus of minute wasps in the family Trichogrammatidae, described by A.A. Girault in 1911. Species are egg parasitoids of leafhoppers (Hemiptera: Cicadellidae), with several species of economic importance as biological control agents. The genus has an almost cosmopolitan distribution. Notable hosts include the beet leafhopper (Circulifer tenellus), a vector of beet curly top virus affecting crops such as tomatoes, sugar beets, and peppers. Two species, A. anatolica and A. turanica, were introduced from Iran into California for biological control and became established. The genus reproduces by arrhenotoky, where unmated females produce only male offspring.
Aphelinus mali
woolly aphid parasite
Aphelinus mali is a tiny parasitoid wasp in the family Aphelinidae that specializes in attacking the woolly apple aphid (Eriosoma lanigerum), a major pest of apple orchards worldwide. Native to the northeastern United States, it has been introduced to numerous apple-growing regions as a biological control agent since 1928. The wasp's life cycle is tightly coupled with its aphid host, with females laying eggs inside living aphids where larvae develop before emerging as adults. It is considered one of the earliest and most successful examples of classical biological control in agriculture.
Aphelopus
Aphelopus is a genus of dryinid wasps described by Dalman in 1823. Species are parasitoids of leafhoppers (Cicadellidae), particularly typhlocybine leafhoppers, and are considered important natural enemies with potential for biological control in agricultural systems. The genus is relatively diverse, with 39 species recorded from China alone and a cosmopolitan distribution. Identification relies on wing venation characteristics and head morphology including clypeal structures.