Asia

Guides

Scrobipalpa artemisiella
thyme moth
Scrobipalpa artemisiella, commonly known as the thyme moth, is a small gelechiid moth with a wingspan of 10–12 mm. The species is widely distributed across Europe, Asia, and has unconfirmed records from North America. Adults are active in summer, and larvae are specialized feeders on thyme and related aromatic herbs.
moth Gelechiidae thyme herbivore Europe Asia Lamiaceae-feeder
Scythropiodes
Scythropiodes is a genus of small moths in the family Peleopodidae (subfamily Oditinae), established by Matsumura in 1931. The genus is distributed across Asia, with the majority of described species occurring in China. Taxonomic research has focused primarily on morphological characterization and species-level identification using genitalia features. The genus contains at least 15 recognized species, with ongoing revisions adding new taxa.
moth Asia taxonomic-revision-needed genitalia-based-identification
Scytonotus
Scytonotus is a genus of flat-backed millipedes in the family Polydesmidae, containing approximately 17 described species. The genus was established by C.L. Koch in 1847. Members of this genus are part of the diverse order Polydesmida, which includes millipedes with flattened bodies and lateral extensions of their dorsal plates (paranota). Species in this genus are poorly documented in scientific literature, with limited ecological and behavioral data available.
millipede flat-backed-millipede Polydesmidae Polydesmida soil-dwelling decomposer myriapod arthropod terrestrial North-America Europe Asia
Sogatella
white-backed planthopper (for S. furcifera)
Sogatella is a genus of delphacid planthoppers in the family Delphacidae, comprising more than 20 described species. The genus was established by Fennah in 1956. The best-studied species, Sogatella furcifera (white-backed planthopper), is a major agricultural pest of rice across Asia. Members of this genus are characterized by their association with rice and grass hosts, with S. furcifera serving as a vector for plant viruses including southern rice black-streaked dwarf virus.
planthopper rice-pest agricultural-pest vector migration Asia Delphacidae Hemiptera phloem-feeder virus-vector
Sophonia
Sophonia is a genus of leafhoppers in the family Cicadellidae, subfamily Evacanthinae. Species in this genus are native to Asia and have been introduced to multiple regions including Hawaii, the continental United States, and Europe. At least one species, Sophonia rufofascia, has become an established pest in Hawaii since its detection in 1987. The genus is characterized by features typical of the Evacanthinae subfamily.
leafhopper Cicadellidae Evacanthinae pest invasive Asia Hawaii Hemiptera
Synchroidae
synchroa bark beetles
Synchroidae is a small family of tenebrionoid beetles containing three extant genera: Mallodrya (monotypic, North America), Synchroa (five species, North America and Asia), and Synchroina (two species, Oriental region). The family has been historically confused with Melandryidae but was established as independent based on larval morphology and adult characters. Members are elongate, slightly flattened beetles with brownish to black coloration, ranging 10–16 mm in length.
beetles Coleoptera Tenebrionoidea nocturnal wood-associated Asia North-America
Tautoneura polymitusa
Tautoneura polymitusa is a small leafhopper (2.4–2.7 mm) in the family Cicadellidae, first described from South Korea in 2016. It represents the first European record of both the species and the genus Tautoneura, having been detected in Hungary in 2012 and subsequently found in Italy, Spain, Slovenia, Ukraine, Russia, and Moldova. The species is associated with Ulmaceae (elm family) and has established stable, locally abundant populations in parts of Europe. It hibernates as adults and is attracted to light traps.
invasive-species leafhopper Ulmaceae adventive Europe Asia Typhlocybinae Erythroneurini
Telamoptilia
Telamoptilia is a genus of small moths in the family Gracillariidae, established by Kumata & Kuroko in 1988. The genus name derives from Greek 'telamon' (belt) and 'ptilia' (small wing), likely referring to wing pattern or structure. It contains six described species distributed across Africa and Asia, with some species known as leaf miners on specific host plants.
Gracillariidae leaf-miners microlepidoptera Asia Africa
Thereuonema
Thereuonema is a genus of scutigeromorph centipedes in the family Scutigeridae. The genus contains at least one well-documented species, Thereuonema tuberculata, which is native to Asia and has been introduced to Europe and North America. Members of this genus are associated with human-modified environments and have established populations outside their native range.
centipede scutigeromorph synanthropic invasive Asia introduced-species
Thereuonema tuberculata
Japanese House Centipede
Thereuonema tuberculata is a scutigeromorph centipede in the family Scutigeridae. Native to Asia, it has been documented from China, Japan, the Korean peninsula, and Taiwan. The species has established introduced populations in North America, particularly in the eastern United States, with confirmed outdoor populations in forested and lakeside habitats in Nebraska. It is commonly confused with the related Scutigera coleoptrata in its introduced range.
centipede house-centipede introduced-species Scutigeridae Scutigeromorpha Chilopoda Asia North-America anthropophilic
Trachelas
Trachelas is a genus of hunting spiders in the family Trachelidae, established by Ludwig Carl Christian Koch in 1872. The genus contains approximately 99 species with worldwide distribution across tropical, subtropical, and temperate regions. Trachelas sensu stricto is restricted to the Old World, while New World species represent a distinct lineage only distantly related to the core group. The genus has undergone significant taxonomic revision, with species previously assigned to Corinnidae now placed in Trachelidae.
spider hunting-spider ground-spider indoor-spider Old-World New-World Trachelidae Corinnidae nocturnal scavenger autumn-activity North-America Asia Africa Europe broad-faced-sac-spider
Trachyuropodidae
Trachyuropodidae is a family of mites in the order Mesostigmata, classified within the suborder Monogynaspida and infraorder Uropodina. The family contains genera such as Leonardiella, with species documented from European and Asian localities including Turkey. Members are found in association with woody vegetation, particularly oak and willow habitats. The group belongs to the diverse assemblage of soil-dwelling and litter-inhabiting mites that contribute to decomposition processes.
mites Mesostigmata Uropodina soil-fauna Turkey Europe Asia
Tremecinae
woodwasps, horntails
Tremecinae is a subfamily of woodwasps (Siricidae) distinguished from Siricinae by its association with the white-rot fungus Cerrena unicolor rather than Amylostereum. Unlike Siricinae, which carry symbiotic fungi in specialized mycangia, at least some Tremecinae species lack these structures and do not internally transport fungal symbionts. Females locate host wood already infected with sapwood-decaying fungi using volatile compounds. The subfamily includes forest pests that interact with fungal communities in decaying wood.
Siricidae woodwasp horntail saproxylic forest-pest Cerrena-unicolor white-rot-fungus mycangia-absent olfactory-host-location Tremex-apicalis Ibalia-japonica parasitoid symbiosis Japan Asia
Trichonephila clavata
Jorō spider, Joro Spider, Parachute spider
Trichonephila clavata, commonly known as the Jorō spider, is a large orb-weaving spider native to East Asia that has become established as an invasive species in the southeastern United States since approximately 2010. First confirmed in Georgia in 2014, it has expanded rapidly across multiple states through a combination of ballooning dispersal and human-mediated transport. The species is notable for its substantial size, striking coloration, and extensive golden webs, but poses minimal risk to humans due to small fangs and docile behavior. Its physiological adaptations—including higher metabolic rate, faster heart rate, and greater cold tolerance than its congener Trichonephila clavipes—suggest potential for continued northward range expansion.
invasive-species orb-weaver ballooning urban-wildlife biological-control cold-tolerance startle-response cannibalism kleptoparasitism Japanese-folklore Nephilidae Trichonephila spider-silk range-expansion physiological-adaptation urban-ecology agricultural-pest-predator iNaturalist Georgia Maryland East-Asia United-States spider-bite venom docile non-aggressive web-architecture egg-sac overwintering sexual-dimorphism metabolic-rate heart-rate freeze-tolerance road-ecology prey-capture competition native-spider-impacts public-perception media-sensationalism management pesticide-alternatives broom-removal ornamental structural-pest ornithology cardinal web-strength trophic-interactions DNA-metabarcoding diet-analysis species-distribution-model physiological-entomology behavioral-ecology arachnology ballooning-dispersal human-mediated-dispersal hitchhiker cargo-stowaway port-of-entry climate-change warming northward-expansion DMV Howard-County Elkridge Ilchester West-Elkridge beech-forest secondary-forest suburban roadside telephone-pole streetlamp forest-edge disturbance-tolerance prey-detection vibrational-noise auditory-disturbance motionless freezing shyness Hannibal-Lecter stink-bug lanternfly evolutionary-history ancient-association ecosystem-service biocontrol natural-enemy invasive-pest brown-marmorated-stink-bug spotted-lanternfly Halyomorpha-halys Lycorma-delicatula competitive-exclusion dominant-orb-weaver abundant population-growth rapid-spread range-span 120,000-km2 physiological-plasticity metabolism twice-as-high 77%-higher-heart-rate 74%-freeze-survival development-rate seasonal-window life-cycle-completion thermal-limitation congener-comparison natural-dispersal human-transport iNaturalist-record citizen-science confirmed-sighting breeding-population male-and-female gravid-female egg-case stowaway cargo-container port-introduction local-port vehicle-transport hitchhiking accelerated-arrival several-years natural-means prognostication prediction model suitable-habitat abundant-suitable-habitat eastern-North-America western-North-America potential-range invasive-range potential-impact environmental-impact ecological-impact ecosystem-impact monitor cautious evidence-based reasonable-journalism sensationalism exaggeration uncritical-acceptance harmless carefully-monitored next-steps research-opportunity call-for-research knowledge-gap little-known novel-ecosystem spreading rapidly-expanding invasive-predator invasive-orb-weaver non-native introduced established colonized thriving doing-fine happy merely-20-minutes-away road-trip visit access-spread beech-leaf-disease BLD state-forest remarkable-colony amongst-beech-trees confirmed-prognostication Davis-and-Frick escape-relative-warmth expand-range-northward eastern-seaboard merely-20-minutes-away-from-home resist-opportunity amazing-predators tales-of-arrival previous-episodes 2022 2024 reduce-angst large-non-native-spider establishing-in-DMV facts-presented past-episode venomous-terrible-painful Nah expert-Rick-Hoebeke risks-small puny-fangs unlikely-pierce-skin video completely-non-aggressive eastern-Howard-County haphazard-webs littered-with-remains former-victims leaves shed-exoskeletons much-larger-female dwarfs-mate positioned-just-above strand-of-silk spinnerets underside-of-abdomen near-red-mound relative-to-hand large-and-docile wait-and-see what-Jorō-means-to-ecosystems help-other-spiders put-a-beat-down-on-invasive-pests stink-bugs spotted-lanternflies passive-hunters enormous-webs larger-than-a-meter-in-diameter capture-prey-snared-in-silk arachnophobes-scary arachnophiles-beautiful important-ecosystem-services crop-pests native-range Asia having-an-old-friend-over-for-dinner reunite first-time large-spiders juicy-prey-items feathered-and-non-feathered-reptiles poses-no-known-threat difficult-to-predict-impact non-native-species experts-suggest beyond-scary-mien give-indigenous-large-orb-weavers-run-for-their-money black-and-yellow-garden-spider marbled-orb-weaver spotted-orb-weaver other-parts-of-world most-abundant-and-dominant-orb-weaver what-will-it-mean only-time-will-tell final-tidbit Jorō-shapeshifter Jorō-gumo beautiful-woman seduces-men binds-with-silk devours bad-date acknowledgements Rick-Hoebeke identifying-Jorō insights David-Coyle Bob-Bellinger great-images knowledge studies Veni-vidi-vici Nelsen-et-al.Physiological-evaluation Nephila-clavata-newly-recorded Hoebeke-et-al.Life-Cycle-Habitat-and-Variation Moore Startle-responses Davis-and-Anerao How-Urban-Tolerant-Are-They Davis-et-al.inspiration details stars Dave-Clement Miri-Talabac Maddie-Potter hooked-up-with-colony differentiate resources.ipmcenters.org link Jorō-guru David-Coyle's-take YouTube calm-your-fears Journal-of-Economic-Entomology Journal-of-Medical-Entomology chemical-management-strategies effective-management-methods spider-control-products safe-and-effective unlikely-to-bite generally-harmless big-spiders colorful-spiders invasive-spiders in-your-face-spiders webs-on-homes landscape-plants structures middle-of-where-people-live-work-and-play northern-Georgia-2014 presumed-arrived-few-years-prior 15-years increased-dramatically much-of-Southeast disjunct-populations northeastern-U.S.firestorm-of-interest rapidly-expanding-range southeastern-states many-counties Carolinas Tennessee Oklahoma more-than-a-century parts-of-Florida rarely-Pennsylvania tropics thermal-limitations higher-metabolism faster-heart-rate better-ability-to-tolerate-freezing complete-life-cycle-rapidly chilly-temperatures suite-of-adaptations expand-northward warm-loving-cousin findings-support-potential much-yet-to-be-learned successfully-survive-northern-winters nation-and-world-warm southern-species-expand higher-latitudes-and-altitudes rapidly-range-expansion-will-occur typical-mode-of-dispersal aerial-dispersal ballooning-on-strands-of-silk Charlotte's-babies Charlotte's-Web spectacular-moniker parachute-spider rain-down-from-airplanes long-distance-transit human-assistance good-hitchhikers inseminated-gravid-female egg-case-stowaway arrival-in-DMV several-years-by-natural-means new-introduction-at-local-port human-assist-in-vehicles accelerate-arrival terrible-and-painful small-fangs visited-cousin golden-silk-spider Costa-Rica exceeding-meter-diameter indigenous-large-orb-weavers run-for-their-money minimal-direct-impact firestorm Andrew-Davis Benjamin-Frick University-of-Georgia past-week native-to-eastern-Asia Japan China Korea Taiwan known-in-Georgia-since-2013 spreading-rapidly joined-its-cousin Trichonephila-clavipes established-over-160-years better-tolerate-freezing survive-northern-winters rapidly-range-expansion typical-mode long-distance new-introduction human-assist arachnophobes arachnophiles brown-marmorated-stink-bugs difficult-to-predict scary-mien indigenous-orb-weavers Mary-Nouri Sarah-Morgan WAMU relatively-harmless spider-season fall leaves-changing-color cool-crisp-autumn-air pumpkin-spice extension-entomologist Southeast not-just-spiders 20-feet-wide homes people-live-work-and-play 15-years-or-so influx-of-calls public entomology-extension-programs media-frenzy two-questions get-rid-of-Jorō-spiders are-Jorō-spiders-dangerous Clemson-University Southern-Adventist-University Washington-College Union-College two-new-studies test-methods-for-eliminating how-likely-to-bite harm scoured-internet recommended-treatment-options spider-specific-management-recommendations medically-relevant widow-spiders brown-recluses do-not-enter-homes relatively-new recommended-products labelled-spider-control-products tested-products several-products-eliminate not-legal-pesticides machine-lubricant not-recommended commercially-available strongly-encourage labelled-products easiest-and-cheapest without-chemicals broom-or-stick walked-into-spider-web rode-bike took-spider-to-face not-great-feeling wonder-if-bitey give-bitten-superpowers unique-studies evaluated-behaviors react-to-people rather-drop-off-web hang-around-and-bite really-provoke volunteered-to-get-bitten yes-you-read-correctly akin-to-mosquito-bite little-redness little-swelling largely-gone-next-day no-acquired-superpowers take-home-message twofold simplest-and-most-effective without-pesticides feel-you-need-pesticides product-labeled-for-spider-control try-not-to-freak-out doesn't-want-to-be-on-you odds-of-getting-bitten-extremely-low enjoy-glorious-fall-weather drink-pumpkin-spiced-beverage watch-where-you-walk Keep-calm-and-carry-on bites-unlikely cause-minimal-discomfort associate-professor Department-of-Forestry-and-Environmental-Conservation socials email extension featured pest-management Entomology-Today latest-posts email-subscription March-2022 wondered make-its-way-to-DMV September-2022 two-observations three-years 40-sightings happy-and-doing-just-fine several-Howard-County-locations week-or-so-ago team-of-scientists University mission killing-ultra-valuable-beech-trees thriving-amongst-beech-trees visit-amazing-predators mature-females both-Trichonephila-species three-locally-common-orb-weaving-species time-spent-immobile mild-disturbance brief-puff-of-air air-puff-response-data five-other-North-American-species published-literature 453-observations freezing-behavior 10-spider-species remained-immobile-under-a-minute both-Trichonephila-spiders over-an-hour unprecedented shyest-ever-documented tolerate-urban-environments remaining-motionless fleeing nonsexual-cannibalism mating-activity females-cannibalize-males food-availability territorial-aggression Southeastern-United-States expanding-range shy behavioral-reactions descriptive-observations photo-documentation anecdotal-observations controlled-pairings lab natural-webs field one-female-biting-and-killing-another short-fight similar-size container 25-trials fights-ensued-40%different-sizes 27-trials fights-happened-18%larger-females-not-always-aggressor 52-lab-trials six-bouts 9%direct-killing field-trials empty-web 14-trials one-fight 7%aggressor-killing-and-wrapping silk away-from-any-web not-necessarily-territoriality intraspecific-aggression provoked-or-stressed deserves-more-study newly-invasive biology physiology new-range closely-related-species successfully-established 160-years investigation compare online-records iNaturalist.org seasonal-distributions timing females physiological-traits environmental-tolerance metabolic-rates cardiac-functioning cold-exposure survival brief-below-freezing-temperature shorter-season complete-lifecycle narrow-period suitable-weather inherently-higher-metabolism low-temperature survive-better 74%-compared-to-50%brief-freeze colder-climatic-region southeastern-USA useful-information planning urban-landscapes busy-roads frequent-disturbances auditory-and-vibrational physiological-or-behavioral-changes northeast-Georgia varying-levels-of-road-traffic prey-capture-behavior simulated-prey tuning-fork 128-hz-frequency touched-to-web attacked 357-total-trials 20-different-roads attacked-59%high-variability some-roadsides-over-80%others-less-than-30%small-but-significant-negative-correlation daily-road-traffic spider-attack-rates moderate--to-heavy-traffic-roads slightly-less-likely-to-attack 51%-vs-65%able-to-live-near-roads cost-in-terms-of-prey-capture did-not-weigh-less compensate-for-disturbance accumulating-evidence human-dominated-landscapes aid-spread introduced-range Eriostethus-rufus polysphinctine-ectoparasitoid araneid-spiders Neoscona-spp.endemic-to-Japan Yamagata-Prefecture 38º46'-N northern-Japan northernmost-record genus identification-confirmed morphology DNA-barcoding cocoon large-modified-web unique inverted-triangle over-50-cm hanging-from-ill-defined-part no-organized-structure host-spider circumstantial-evidences structure-of-modified-web pre-existing-web-partly-reused orb-web-completely-removed sustaining-threads-newly-moored parasitises-host-spiders two-subfamilies unusual-for-group invasion-potential known-invasive-range review dispersal-abilities potential-impacts management-strategies October-2022 at-least-120,000-km2 South-Carolina North-Carolina Alabama West-Virginia pattern-of-spread natural-dispersal-mechanisms human-mediated-transport large-bodied-orb-weavers flying-insects small-animals thirteen-co-occurring-spider-species monitored-for-competition resources web-building-sites natural-and-urban-habitats management-options-limited advise-journalists-and-experts against-exaggerating-potential-environmental-impact ecologically-harmless rapid-spread-carefully-monitored cautious-evidence-based-approach determining-next-steps three-material-types web-capture introduced-species alter-established-trophic-interactions molecular-analysis resolve-changes community-structure foraging-links east-coast fecal-samples prey-remains-from-webs dissected-guts compare-diet-composition arthropod-targeted-COI-primers Illumina-MiSeq highest-diversity highest-richness highest-proportion-prey-reads recovery-of-prey-reads dissected-gut-content low overwhelmed-by-Jorō-spider-DNA high-proportions-Jorō-spider-reads higher-prey-diversity-and-richness detected-prey-DNA several-days-after-capture initial-gut-retention-time-estimates first-glimpse complexity-of-trophic-associations introduced-web-building-spider viable-material source-of-prey-DNA estimates-of-biodiversity Jammu-and-Kashmir northernmost-shifting document-presence behavior habitat-adaptation colder-temperature-environment Goha-Tehsil Doda-district 1556.74-meters road-bridge nearby-vegetation brightly-colored-abdomens black green yellow red long-legs yellow-and-black-bandings predominantly-seen smaller fewer unique-behaviors remaining-motionless-after-disturbances 5-10-meters-of-roads adaptation-to-human-settlements field-observations image-documentation expert-confirmation thrive-in-colder-conditions 15-20-degrees-Celsius October-November no-spider-found higher-elevations significantly-lower-temperatures adaptation-limited-to-milder-cold-regions new-insight adaptation need-for-further-research non-native-habitats eggs overwinter harsh-weather-conditions no-protection compact-silk-case milky-coating chorionic-microspheres CM fine-structural-characteristics ecological-importance uneven-size-distribution outer-eggmass evenly-covered single-layer diameter-2.3-µm short-papillary-projections segmental-adhesion mucous-components insoluble-in-water partially-soluble-in-absolute-ethanol spherical-structure HFIP strong-organic-solvent not-derived-from-vitellogenic-or-choriogenetic-processes CM-adhesive-coatings ovipositional-process equivalent-to-cocoon-silk protective-functions silken-eggcase Northern-Cardinal Cardinalis-cardinalis perches steals-food southeast-of-the-United-States questions native-fauna-affected consuming-prey-items example native-species-deriving-benefit manner-of-kleptoparasitism perched-directly-on-web supported-its-weight 42-48-g first-documented-case spider-web-supporting-perching-bird measurements force-gauge typical-webs support-masses-up-to-70-g small-but-growing-body-of-knowledge non-native-spider species-distribution-models ecological-niche-modeling predict-areas establish-if-introduced three-distinct-genetic-lineages mitochondrial-COI genome-wide-SNPs Korean-Peninsula
Trimium brevicorne
Trimium brevicorne is a small rove beetle in the subfamily Pselaphinae, commonly known as ant-loving beetles. The species occurs across Europe, Northern Asia (excluding China), and has been introduced to North America. It belongs to a group of beetles that are typically found in association with ant colonies.
ant-loving-beetle rove-beetle Pselaphinae introduced-species Europe Asia North-America
Tvetenia
non-biting midges
Tvetenia is a genus of non-biting midges in the family Chironomidae, subfamily Orthocladiinae. Species occur across northern and eastern Europe, extending into Russia and Central Asia. The genus was established by Kieffer in 1922. Taxonomic revisions have clarified species boundaries, including the synonymy of T. vitracies with T. tshernovskii.
Chironomidae Orthocladiinae non-biting-midges aquatic-insects freshwater Europe Asia Kieffer-1922 Tvetenia-tshernovskii Tvetenia-vitracies taxonomic-revision
Unaspis
Unaspis is a genus of armored scale insects (family Diaspididae) containing economically significant agricultural and horticultural pests. The genus includes at least 19 described species, with several species infesting citrus crops (U. citri, U. yanonensis) and ornamental plants (U. euonymi). Species in this genus are characterized by their protective waxy coverings and phytophagous feeding habits on host plant tissues.
armored-scale Diaspididae agricultural-pest horticultural-pest citrus-pest biological-control invasive-species Asia global-distribution
Valgus
Valgus is a genus of small scarab beetles in the family Scarabaeidae, subfamily Cetoniinae. Most described species occur in Asia, with some extending into northern Africa and Europe. Three species are native to the New World, and one species is found in South Africa. Adults are characterized by their strongly flattened bodies and distinctive scale-like setae covering both dorsal and ventral surfaces.
scarab flower-visiting termite-associated flattened-body scale-setae Cetoniinae Valgini Asia New-World South-Africa
Xyleborus xylographus
Xyleborus xylographus is a species of bark beetle in the family Curculionidae, first described by Thomas Say in 1826. It belongs to the tribe Xyleborini, which includes ambrosia beetles that cultivate symbiotic fungi in wood galleries. The species has been recorded in North America, the Caribbean, and Southern Asia. Like other members of its genus, it likely engages in fungus-farming behavior, though specific ecological details for this species remain limited in published literature.
bark-beetle ambrosia-beetle Curculionidae Scolytinae Xyleborini fungus-farming wood-boring-beetle invasive-potential forest-pest Coleoptera Xyleborus North-America Caribbean Asia Say-1826 Bostrichus-xylographus
Xylocopa appendiculata
Appendiculate Carpenter Bee, Asian Carpenter Bee
Xylocopa appendiculata is a large carpenter bee native to eastern Asia, first reported in the United States in 2013 with established populations in northern California. It belongs to the subgenus Alloxylocopa. The species has been studied using distribution modeling to predict its potential spread across North America. Like other carpenter bees, it nests in wood and serves as a pollinator, though specific ecological details for this species remain limited.
carpenter-bee introduced-species invasive-potential pollinator wood-nesting-bee Alloxylocopa California Asia distribution-modeling Maxent
Zaitzevia
riffle beetles
Zaitzevia is a genus of riffle beetles (family Elmidae) comprising approximately 19–29 described species, depending on taxonomic treatment. The genus was established by George Charles Champion in 1923 and is named after Russian entomologist Filipp Zaitsev. Species diversity is concentrated in East, Southeast, and Central Asia, with additional representation in North America. Chinese species are particularly well-documented, with multiple new species described from Sichuan, Yunnan, Hunan, Hubei, and Guizhou provinces.
riffle-beetle aquatic-insect stream China Asia North-America Elmidae Macronychini