Tortricidae
Guides
Eucosma kiscana
Eucosma kiscana is a small moth species in the family Tortricidae, first described by Kearfott in 1907 under the basionym Thiodia kiscana. It belongs to the large genus Eucosma, a group of tortricid moths commonly known as phaneta or eucosma moths. The species has been documented in the northeastern United States, with confirmed records from Vermont. Like many Eucosma species, it is likely associated with herbaceous or woody plants, though specific biological details remain poorly documented.
Eucosma knudsoni
Eucosma knudsoni is a species of tortricid moth described from Texas in 2015. It belongs to a large genus of small moths commonly known as bell moths or eucosmas, many of which are associated with Asteraceae host plants. The species is known from very few observations, reflecting both its recent description and likely restricted distribution or cryptic habits. As a member of the Olethreutinae subfamily, it possesses the characteristic tortricid resting posture with wings held rooflike over the body.
Eucosma modernana
Eucosma modernana is a species of tortricid moth in the family Tortricidae, subfamily Olethreutinae. It was described by James Halliday McDunnough in 1925. The genus Eucosma comprises numerous small to medium-sized moths, many of which are associated with Asteraceae host plants. Specific ecological details for E. modernana remain poorly documented.
Eucosma montanana
Eucosma montanana is a species of tortricid moth first described by Walsingham in 1884. It belongs to the family Tortricidae (leafroller moths), a large and economically important group of moths. The species is placed in the subfamily Olethreutinae and tribe Eucosmini. As a member of the genus Eucosma, it is part of a diverse group of small moths, many of which are associated with specific host plants.
Eucosma ochrocephala
Pale-headed Eucosma Moth
Eucosma ochrocephala is a species of tortricid moth in the family Tortricidae, subfamily Olethreutinae. The species was first described by Walsingham in 1895 under the basionym Semasia ochrocephala. It is part of the diverse genus Eucosma, which contains numerous small moth species, many of which are associated with Asteraceae host plants. The species is recognized in major taxonomic databases including Catalogue of Life, GBIF, and NCBI Taxonomy.
Eucosma pallidarcis
Eucosma pallidarcis is a small tortricid moth described by Heinrich in 1923. As a member of the genus Eucosma, it belongs to a diverse group of moths commonly known as bell moths or shoot moths. The species is documented in North America, with observations primarily from the United States. Like other Eucosma species, adults are nocturnal and attracted to light. Larval biology remains poorly documented for this particular species.
Eucosma parmatana
Eucosma parmatana is a species of olethreutine leafroller moth in the family Tortricidae, first described by Clemens in 1860. It is native to North America and has been recorded from Vermont and other parts of the United States. The species is assigned MONA/Hodges number 2937.
Eucosma patagoniana
Eucosma patagoniana is a species of tortricid moth described by Wright in 2014. The species epithet refers to Patagonia, the region of southern Argentina and Chile where the type specimens were collected. As a member of the large genus Eucosma, it belongs to a group of small moths commonly known as phaneta moths or tortricid leafrollers. The species is known from a limited number of observations, reflecting the generally understudied nature of Neotropical tortricid diversity.
Eucosma rupestrana
Eucosma rupestrana is a species of tortricid moth in the subfamily Olethreutinae, described by James Halliday McDunnough in 1925. It belongs to the large genus Eucosma, which comprises numerous small moths commonly known as phaneta or eucosma moths. Very little species-specific information has been published for this taxon.
Eucosma striatana
Striated Eucosma Moth
Eucosma striatana is a small tortricid moth in the tribe Eucosmini. The species was described from North America by Clemens in 1860, originally placed in the genus Anchylopera. It is part of a species complex within Eucosma that is challenging to distinguish morphologically. The common name refers to the striated or lined appearance of the forewings.
Eucosma tarandana
Eucosma tarandana is a species of tortricid moth in the subfamily Olethreutinae, first described by Heinrich Benno Möschler in 1874. It belongs to a large genus of small moths commonly known as bell moths or tortrix moths. The species is part of the Eucosmini tribe, whose larvae are predominantly associated with coniferous and deciduous trees.
Eucosma tenuiana
Eucosma tenuiana is a species of tortricid moth in the subfamily Olethreutinae, described by Walsingham in 1879. It belongs to the large genus Eucosma, which contains numerous small moths often associated with composite plants. The species is rarely encountered, with only 8 observations recorded on iNaturalist, suggesting it is either genuinely uncommon or underreported.
Eucosma viridis
Eucosma viridis is a species of tortricid moth described in 2010 by Wright and Gilligan. It belongs to the large genus Eucosma, which comprises numerous small moths commonly known as bell moths or leafroller moths. As a recently described species, detailed biological information remains limited.
Eugnosta
Eugnosta is a genus of tortricid moths in the tribe Cochylini. The genus was first reported from Chile in 2015 with the description of Eugnosta azapaensis, a gall-inducing species associated with Baccharis salicifolia in the northern Atacama Desert. Species in this genus are characterized by their association with host plants, with at least some species inducing galls on Asteraceae.
Eugnosta bimaculana
narrow-patch carolella moth, two-spotted carolella
Eugnosta bimaculana is a small tortricid moth with a wingspan of 13–14 mm. It occurs across the southeastern and south-central United States, from Maryland to Florida and west to Texas and Oklahoma. Adults are active year-round in the southern portion of its range, with more restricted flight periods in northern populations.
Eugnosta deceptana
Deceptive Cochylid
Eugnosta deceptana is a small tortricid moth known from southern Texas. Adults are active across much of the year with records from November through January and again in March, May, June, and August. The species was described by August Busck in 1907 and remains poorly documented beyond basic collection data.
Eugnosta willettana
Eugnosta willettana is a tortricid moth described by Comstock in 1939. The species is known from a limited geographic range in southwestern North America, with records from southern California, southern Arizona, and Mexico. Adults are active during winter months, with flight records from January to February. The wingspan measures 24–27 mm.
Eulia ministrana
Ferruginous Eulia Moth
Eulia ministrana is a tortricid moth with a broad Palearctic and Nearctic distribution. Adults are active in late spring and early summer, with a single annual generation. Larvae develop as leaf-rollers on various deciduous trees and shrubs. The species has been studied for its population response to industrial pollution, showing that abundance decline may occur without detectable morphological stress indicators.
Eumarozia
Eumarozia is a genus of tortricid moths in the subfamily Olethreutinae, established by Heinrich in 1926. The genus contains at least three described species: E. beckeri, E. elaeanthes, and E. malachitana. These moths are small to medium-sized with the characteristic bell-shaped wing posture typical of the family Tortricidae. The genus has been recorded in North America, with distribution records from the United States including Vermont.
Evora
Evora is a genus of tortricid moths described by Heinrich in 1926. It belongs to the subfamily Olethreutinae, a group of small to medium-sized moths commonly known as tortrix moths or leafroller moths. The genus contains multiple species, though specific details about individual species remain limited in published literature.
Evora hemidesma
Evora hemidesma is a species of tortricid moth described by Zeller in 1875. It belongs to the subfamily Olethreutinae and is one of few species in the genus Evora. The species has been recorded from prairie and northern regions of central North America, with specimens documented from Canadian provinces and the northeastern United States.
Grapholita delineana
Eurasian hemp moth, hemp moth, hemp borer
Grapholita delineana, commonly known as the Eurasian hemp moth or hemp borer, is a tortricid moth species in the family Tortricidae. It has emerged as a significant economic pest of hemp (Cannabis sativa) in the United States following the crop's legalization for commercial production. The species is native to Eurasia but has established populations in North America where it can cause substantial damage to hemp crops, particularly those grown for cannabidiol (CBD) production.
Grapholita edwardsiana
San Francisco Tree Lupine Moth
Grapholita edwardsiana, known as the San Francisco Tree Lupine Moth, is a species of tortricid moth in the family Tortricidae, subfamily Olethreutinae. The species was described by Kearfott in 1907 and is placed in the genus Grapholita, a diverse group of small moths commonly referred to as fruitworms or bud moths. Many Grapholita species are associated with leguminous plants, and the specific epithet "edwardsiana" suggests a possible association with lupines (Lupinus), particularly tree lupine. The species is part of the tribe Grapholitini, which includes numerous economically important pests of fruits and seeds.
Grapholita interstinctana
Clover Head Caterpillar Moth
Grapholita interstinctana is a tortricid moth species first described by Clemens in 1860. It is commonly known as the Clover Head Caterpillar Moth, reflecting its association with clover as a host plant. The species belongs to the Olethreutinae subfamily within the Tortricidae family, a group known for many economically significant agricultural pests. Records indicate presence in the northeastern United States, particularly Vermont.
Grapholitini
Grapholitine moths, Leafroller moth tribe
Grapholitini is a tribe of tortricid moths comprising approximately 1200 described species. The tribe contains numerous economically significant agricultural pests that damage fruits and seeds. Molecular phylogenetic analyses have resolved the tribe into two major lineages—a Dichrorampha clade and a Cydia clade—with the genus Grapholita found to be polyphyletic. The tribe likely originated in the Nearctic, Afrotropical and Neotropical regions during the middle Eocene (ca. 44.3 Ma).
Gretchena amatana
Gretchena amatana is a species of tortricid moth in the family Tortricidae, described by Carl Heinrich in 1923. It belongs to the subfamily Olethreutinae and tribe Eucosmini. The species is known from the northeastern United States, with distribution records from Vermont and surrounding areas. As with many small tortricid moths, detailed ecological information remains limited in published literature.
Gretchena concubitana
Gretchena concubitana is a species of tortrix moth in the family Tortricidae, described by Carl Heinrich in 1923. It belongs to the subfamily Olethreutinae and tribe Eucosmini. The genus Gretchena comprises small moths primarily distributed in North America. Records from iNaturalist indicate this species has been documented in at least 37 observations, suggesting it is not extremely rare, though detailed biological studies remain limited.
Gretchena delicatana
ironwood fruitworm
Gretchena delicatana is a species of tortricid moth described by Heinrich in 1923. It is commonly known as the ironwood fruitworm, reflecting its association with ironwood trees (Ostrya virginiana). The species belongs to the subfamily Olethreutinae and tribe Eucosmini. It has been documented in the northeastern United States, particularly Vermont.
Gretchena deludana
arrowhead moth
Gretchena deludana, known as the arrowhead moth, is a species of tortricid moth in the family Tortricidae. It was first described by Clemens in 1864 under the basionym Hedya deludana. The species has been assigned MONA/Hodges number 3259, a standard identifier used in North American lepidopteran taxonomy. It belongs to the subfamily Olethreutinae and tribe Eucosmini.
Gretchena nymphana
Gretchena nymphana is a species of tortricid moth described by Blanchard and Knudson in 1983. It belongs to the subfamily Olethreutinae within the family Tortricidae, a large group of small to medium-sized moths commonly known as leafroller moths. The genus Gretchena is part of the tribe Eucosmini, which contains numerous species associated with various host plants. Information on the biology and ecology of this particular species remains limited in published literature.
Gretchena watchungana
Gretchena watchungana is a species of tortricid moth in the family Tortricidae, originally described as Epinotia watchungana by Kearfott in 1907. The species belongs to the subfamily Olethreutinae and tribe Eucosmini. It is part of a genus of small moths whose larvae are associated with various host plants. The specific epithet "watchungana" refers to the Watchung Mountains in New Jersey, indicating the type locality of the original specimen.
Gymnandrosoma
Gymnandrosoma is a genus of tortricid moths in the tribe Grapholitini. The genus includes at least eight described species distributed primarily in the Americas. Gymnandrosoma aurantianum, the most economically significant species, is a major agricultural pest known as the citrus fruit borer in Brazil and the macadamia nut borer in Central America. Other species in the genus have been described from North and South America, with limited biological information available.
Gynnidomorpha
Gynnidomorpha is a genus of tortricid moths in the tribe Cochylini, established by Turner in 1916. The genus comprises at least 16 described species distributed across the Palearctic and Nearctic regions. Species within this genus are generally small moths, though specific morphological and biological characteristics vary considerably between species. The genus has been subject to taxonomic revision, particularly for Chinese species.
Gypsonoma aceriana
poplar shoot-borer, Rosy Cloaked Shoot
Gypsonoma aceriana is a small tortricid moth whose larvae bore into poplar and maple shoots, causing economic damage in plantations and nurseries. Native to Europe, it ranges eastward through Russia to Turkey, Iraq, and North Africa. The species has been introduced to Japan, where it exhibits two to three generations annually. It is a significant pest of ornamental poplars, with older larvae destroying terminal shoots and causing bushy, unmarketable growth in young trees.
Gypsonoma adjuncta
Gypsonoma adjuncta is a species of tortricid moth in the tribe Eucosmini, described by Carl Heinrich in 1924. It is a small moth distributed across the Canadian Prairie provinces. The species belongs to a genus whose larvae are typically associated with woody plants, though specific host associations for this species remain poorly documented.
Gypsonoma fasciolana
Willow-and-poplar Leafroller
Gypsonoma fasciolana is a tortricid moth species known as the Willow-and-poplar Leafroller. The species is associated with willow and poplar host plants, as indicated by its common name. It has been documented across parts of Canada and the northern United States.
Gypsonoma n-sp-one
Gypsonoma n-sp-one is a species of tortrix moth in the family Tortricidae, subfamily Olethreutinae. The genus Gypsonoma comprises small moths whose larvae are commonly associated with woody plants, particularly in the family Salicaceae. As a newly described or undescribed species (indicated by the provisional designation 'n-sp-one'), formal documentation of its biology and distribution remains limited. The species follows the general morphological pattern of Gypsonoma, with characteristic wing patterns and genitalia structures typical of the genus.
Gypsonoma substitutionis
Gypsonoma substitutionis is a small moth in the family Tortricidae, described by Heinrich in 1923. The species belongs to the subfamily Olethreutinae and tribe Eucosmini. It has been documented across parts of northern North America, with records from the Canadian prairies and the northeastern United States. As a member of the genus Gypsonoma, it is part of a group of tortricid moths whose larvae are generally associated with woody plants.
Hedya
Hedya Moths
Hedya is a genus of tortricid moths in the subfamily Olethreutinae. The genus was established by Jacob Hübner in 1825. Species in this genus are found across the Holarctic region, with records from Europe and North America. The genus contains multiple species, though specific species-level information is limited in the provided sources.
Hedya nubiferana
marbled orchard tortrix, green budworm moth
Hedya nubiferana is a small tortricid moth with a wingspan of 15–21 mm, characterized by distinctive forewing markings including a square basal black spot and a prominent white cross-band at the wing tip. The species occurs across the Palearctic and Nearctic realms. Adults are active from May to August in western Europe, with timing varying across the range. Larvae develop on hawthorn (Crataegus) and blackthorn (Prunus spinosa).
Hedya ochroleucana
Hedya ochroleucana is a small moth in the family Tortricidae (leafroller moths). The name 'of-authors' indicates this is a name used in literature that does not correspond to a validly described species, representing a misidentification or unresolved taxonomic entity. Specimens under this name have been reported from certain European regions, but the true identity remains unclear. The genus Hedya contains numerous small, often poorly differentiated species that require genital dissection for reliable identification.
Hedya salicella
White-backed Marble
Hedya salicella is a small tortricid moth commonly known as the White-backed Marble. It is a single-brooded species with adults active from June through September. The species is widely distributed across Europe and has been introduced to North America. Its larvae are specialized feeders on willow and poplar species.
Hendecaneura
Hendecaneura is a genus of tortricid moths in the subfamily Olethreutinae, established by Walsingham in 1900. The genus contains seven described species distributed primarily in Asia and North America. At least one species, H. shawiana, is a documented agricultural pest of blueberry. Most species were described by Walsingham in 1900 from material collected in Asia.
Hendecaneura shawiana
Blueberry Tip Borer Moth
Hendecaneura shawiana is a tortricid moth commonly known as the blueberry tip borer. It is a univoltine species with one generation per year. The larvae are specialized feeders that bore into the shoot tips of blueberry plants, causing characteristic dieback damage. It is recognized as a pest of cultivated blueberry in northeastern North America.
Henricus cognata
Henricus cognata is a species of tortricid moth described by Walsingham in 1914. The species occurs in the southern United States and Mexico (Veracruz). As a member of the tribe Cochylini within the subfamily Tortricinae, it belongs to a group of moths commonly known as tortrix or leafroller moths. The genus Henricus comprises multiple species, most of which are distributed in the Americas.
Henricus edwardsiana
Contrasting Henricus Moth
Henricus edwardsiana is a small tortricid moth species described by Walsingham in 1884. Originally placed in the genus Conchylis, it is now classified in Henricus. The species is found in the southwestern United States, with confirmed records from Arizona and California. It belongs to the tribe Cochylini within the subfamily Tortricinae. Limited ecological information is available for this species.
Henricus fuscodorsana
cone cochylid moth, Cone Cochylid
Henricus fuscodorsana, known as the cone cochylid moth, is a small tortricid moth found in western North America. The species is notable for its specialized larval ecology: larvae are cone miners that develop within the cones of coniferous trees across multiple genera. Adults are active across a broad seasonal window from spring through autumn.
Henricus umbrabasana
Brown-shouldered Henricus
Henricus umbrabasana is a small tortricid moth known from the Pacific Coast of the United States. The species was described by Kearfott in 1908 and is characterized by its modest forewing size of 7.5–9 mm. It is one of approximately 40 described species in the genus Henricus, a group of small tortricine moths predominantly distributed in the Nearctic region.
Hulda
Hulda is a genus of tortricid moths in the family Tortricidae, subfamily Olethreutinae. The genus was described by Heinrich in 1926. It belongs to the tribe Olethreutini, a group of small moths commonly known as leafroller moths or tortrix moths. Very little specific information is documented about the biology or ecology of this genus.
Hystrichophora
Hystrichophora is a genus of small moths in the family Tortricidae, first described by Walsingham in 1879. These moths belong to the subfamily Olethreutinae and tribe Enarmoniini. Members of this genus are leafroller moths, with larvae that roll or tie leaves to create shelters. The genus is part of the diverse Tortricidae family, which contains many economically significant agricultural pests.