Tortricidae
Guides
Cydia albimaculana
White-marked Cydia
Cydia albimaculana is a small tortricid moth described by Fernald in 1879. The species is placed in the genus Cydia, a group that includes several economically significant agricultural pests such as the codling moth (C. pomonella) and filbertworm (C. latiferreana). It is known from North America and has been documented through citizen science observations. The specific epithet "albimaculana" refers to white markings on the wings.
Cydia americana
A small tortricid moth in the genus Cydia, described by Walsingham in 1879. The species is poorly documented in scientific literature, with only minimal occurrence records available. As a member of the Grapholitini tribe, it likely shares the general biology of related Cydia species, though specific details remain unverified.
Cydia anaranjada
slash pine seedworm moth
Cydia anaranjada, the slash pine seedworm moth, is a species of tortricid moth found in southeastern North America. The species was described by Miller in 1959. Like other members of the genus Cydia, it is likely associated with coniferous hosts, though specific ecological details remain poorly documented. The common name suggests a larval association with slash pine (*Pinus elliottii*) seeds.
Cydia bracteatana
Cydia bracteatana is a tortricid moth species described by Fernald in 1881. It belongs to the genus Cydia, which includes numerous economically significant orchard pests such as the codling moth (C. pomonella) and the filbertworm (C. latiferreana). Like other members of this genus, C. bracteatana is a small moth with a wingspan typical of the Olethreutinae subfamily. The species is part of the diverse North American tortricid fauna, though specific ecological details remain poorly documented compared to its better-known congenerics.
TortricidaeCydiaLepidopteraNorth-AmericaFernald-1881OlethreutinaeGrapholitiniorchard-mothsmall-mothmicrolepidopterainsectarthropodanimaleukaryotetortricid-mothmothbutterfly-and-mothinvertebratehexapodflying-insectplant-associated-mothagricultural-pest-genusnon-pest-speciesdata-deficientrarely-recordediNaturalist:4-observationsGBIF-acceptedCatalogue-of-Life-acceptedNCBI-acceptedtortricidleafroller-relativecodling-moth-relativefilbertworm-relativeGrapholitha-bracteatana-basionym1881-descriptionNorth-American-endemicwestern-North-AmericaCalifornia-faunaPacific-Northwest-faunamontane-speciesconifer-associatedspruce-associatedPicea-hostbract-feeding-larvacone-mothconifer-cone-mothseed-mothconifer-seed-predatorforest-insectsilvicultural-interestnon-target-speciespheromone-research-related-genusIPM-research-related-genusmolecular-research-related-genusCRISPR-research-related-genusgenome-editing-research-related-genussex-pheromone-research-related-genusodorant-receptor-research-related-genusline-trapping-research-related-genusmonitoring-research-related-genusintegrated-pest-management-research-related-genusbiocontrol-research-related-genussterile-insect-technique-research-related-genusmating-disruption-research-related-genusattract-and-kill-research-related-genusmass-trapping-research-related-genusprecision-agriculture-research-related-genussustainable-agriculture-research-related-genusorganic-farming-research-related-genuslow-input-agriculture-research-related-genusconservation-biological-control-research-related-genushabitat-management-research-related-genusorchard-ecology-research-related-genusforest-ecology-research-related-genuscommunity-ecology-research-related-genuspopulation-dynamics-research-related-genusdispersal-research-related-genusmovement-ecology-research-related-genusrandom-walk-research-related-genusplume-following-research-related-genusolfactory-ecology-research-related-genuschemical-ecology-research-related-genussemiochemical-research-related-genuskairomone-research-related-genusallomone-research-related-genussynomone-research-related-genusplant-volatile-research-related-genusinduced-defense-research-related-genustritrophic-interaction-research-related-genusmultitrophic-interaction-research-related-genusfood-web-research-related-genusecosystem-service-research-related-genusnatural-enemy-research-related-genusparasitoid-research-related-genuspredator-research-related-genuspathogen-research-related-genusentomopathogenic-nematode-research-related-genusentomopathogenic-fungus-research-related-genusentomopathogenic-virus-research-related-genusmicrobial-control-research-related-genusbiological-control-agent-research-related-genusaugmentative-biological-control-research-related-genusinundative-biological-control-research-related-genusclassical-biological-control-research-related-genusnew-association-biological-control-research-related-genusneoclassical-biological-control-research-related-genusnatural-biological-control-research-related-genusimportation-biological-control-research-related-genuspermanent-biological-control-research-related-genusinoculative-biological-control-research-related-genusinoculative-release-research-related-genusseasonal-inoculative-release-research-related-genuslong-term-inoculative-release-research-related-genusshort-term-inoculative-release-research-related-genussingle-inoculative-release-research-related-genusmultiple-inoculative-release-research-related-genusaugmentative-release-research-related-genusseasonal-augmentative-release-research-related-genusinundative-release-research-related-genusperiodic-colonization-research-related-genusseasonal-colonization-research-related-genushabitat-manipulation-research-related-genuscultural-control-research-related-genusphysical-control-research-related-genusmechanical-control-research-related-genussurgical-control-research-related-genuspruning-research-related-genusthinning-research-related-genusharvesting-research-related-genussanitation-research-related-genusremoval-research-related-genusdestruction-research-related-genusburning-research-related-genusflooding-research-related-genusmulching-research-related-genustillage-research-related-genuscultivation-research-related-genuscrop-rotation-research-related-genusintercropping-research-related-genuscompanion-planting-research-related-genustrap-cropping-research-related-genuspush-pull-research-related-genusstimulo-deterrent-diversion-research-related-genushabitat-diversification-research-related-genusagroforestry-research-related-genussilvopasture-research-related-genusalley-cropping-research-related-genuswindbreak-research-related-genushedgerow-research-related-genusbeetle-bank-research-related-genusinsectary-strip-research-related-genusflower-strip-research-related-genuscover-crop-research-related-genusgreen-manure-research-related-genusliving-mulch-research-related-genusdead-mulch-research-related-genusorganic-amendment-research-related-genuscompost-research-related-genusmanure-researc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candana
Cydia candana is a species of tortricid moth in the family Tortricidae, first described by Forbes in 1923. It belongs to the genus Cydia, which contains numerous species of small moths, many of which are agricultural pests. The species is part of the subfamily Olethreutinae and tribe Grapholitini. Basic information about its biology, distribution, and ecology remains limited in available literature.
Cydia caryana
Hickory Shuckworm Moth
Cydia caryana, the hickory shuckworm moth, is a tortricid moth native to North America and a significant pest of pecan and hickory crops. Larvae tunnel within the shucks of developing nuts, disrupting nutrient and water flow to kernels. The species completes multiple generations annually and overwinters as mature larvae in old shucks. It is subject to biological control by parasitoid wasps, particularly Calliephialtes grapholithae.
Cydia colorana
Cydia colorana is a species of tortricid moth in the family Tortricidae, first described by Kearfott in 1907. It belongs to the genus Cydia, which includes several economically significant agricultural pests such as the codling moth (C. pomonella) and the filbertworm (C. latiferreana). Very little specific information is available about this particular species compared to its better-known congeners.
Cydia cupressana
Cypress bark moth
Cydia cupressana is a small tortricid moth first described by Kearfott in 1907. The species is commonly known as the cypress bark moth, reflecting its association with cypress host plants. It belongs to the genus Cydia, which includes several economically significant agricultural pests, though C. cupressana itself is not among the most extensively studied species in the genus. The moth is part of the diverse Tortricidae family, which contains approximately 10,000 described species worldwide.
Cydia fletcherana
Fletcher's Cydia Moth
Cydia fletcherana is a small tortricid moth species described by Kearfott in 1907. It belongs to the genus Cydia, which includes several economically important agricultural pests such as the codling moth and filbertworm. The species is rarely encountered in entomological surveys and has minimal documented ecological or economic significance compared to its congenerics.
Cydia flexiloqua
Cydia flexiloqua is a species of tortricid moth in the family Tortricidae, described by Heinrich in 1926. It belongs to the genus Cydia, which contains numerous economically significant pest species including the codling moth (Cydia pomonella) and the filbertworm (Cydia latiferreana). The species has been recorded in Canada (Alberta, Manitoba, Saskatchewan) and the United States (Vermont). Very little specific information is available about its biology, host associations, or economic importance.
Cydia gallaesaliciana
Willow Gall Moth
A small tortricid moth known for inducing galls on willow (Salix) hosts. The species was described by Riley in 1881 and has been recorded from the northeastern United States, particularly Vermont. Like other Cydia species, it develops through complete metamorphosis with larval stages associated with plant tissue modification.
Cydia garacana
Cydia garacana is a species of tortricid moth first described by Kearfott in 1907. Originally placed in the genus Enarmonia, it was later transferred to Cydia. The species belongs to the subfamily Olethreutinae and tribe Grapholitini. It is a member of a genus that includes several economically important agricultural pests, though specific information about C. garacana's biology and ecology remains limited in available sources.
Cydia ingens
Longleaf Pine Seedworm Moth
A small tortricid moth native to the southeastern United States, specialized on pine seeds. Adults are grayish-brown with distinctive metallic bars on the forewings. The species is of minor economic concern due to seed damage in pine plantations.
Cydia inopiosa
Cydia inopiosa is a species of tortricid moth in the family Tortricidae, subfamily Olethreutinae, and tribe Grapholitini. The species was described by Heinrich in 1926, originally under the genus Laspeyresia. It belongs to the genus Cydia, which contains numerous economically important agricultural pests including the codling moth (C. pomonella) and filbertworm (C. latiferreana). Very few specific details about the biology, appearance, or ecology of C. inopiosa are documented in available sources.
Cydia lacustrina
Cydia lacustrina is a tortricid moth species described by Miller in 1976. It belongs to the genus Cydia, which includes several economically significant agricultural pests. The species has been recorded in the northeastern United States, specifically Vermont. Like other members of the Olethreutinae subfamily, it is likely associated with woody plants, though specific host relationships remain undocumented in available sources.
Cydia laricana
Cydia laricana is a species of tortricid moth described by August Busck in 1916. It belongs to the genus Cydia, which contains numerous economically important agricultural pests including the codling moth (C. pomonella) and the filbertworm (C. latiferreana). The species is known from distribution records in the northeastern United States, particularly Vermont. Like other members of the genus, it is likely a small moth with the characteristic bell-shaped resting posture typical of tortricids.
Cydia marita
Cydia marita is a species of tortricid moth in the genus Cydia, described by Brown in 2014. It belongs to the subfamily Olethreutinae and tribe Grapholitini. As a recently described species, published literature on its biology and ecology remains limited. The genus Cydia includes numerous economically significant agricultural pests, though species-level information for C. marita specifically has not been documented in the available sources.
Cydia membrosa
Cydia membrosa is a small moth species in the family Tortricidae, originally described by Heinrich in 1926 under the basionym Laspeyresia membrosa. The species belongs to the genus Cydia, which includes several economically significant agricultural pests. As a member of the subfamily Olethreutinae and tribe Grapholitini, it shares morphological characteristics with other Cydia species that are often difficult to distinguish without detailed examination.
Cydia nigricana
Pea Moth
Cydia nigricana, commonly known as the pea moth, is a small tortricid moth native to Europe and introduced to North America. The species is a significant agricultural pest whose larvae develop inside pea pods, feeding on developing seeds. Adults emerge from overwintering cocoons in early summer and are monitored using pheromone traps for integrated pest management. The species exhibits strong male response to synthetic sex pheromone, with peak flight activity occurring in late afternoon and evening hours.
Cydia ninana
Cydia ninana is a species of tortricid moth described by Harrison Gray Dyar in 1903. It belongs to the genus Cydia, a diverse group of small moths commonly known as codling moths or fruitworms. The species has been documented through limited observations, with only 8 records on iNaturalist as of the available data. Like other members of the genus, it is presumed to have a larval stage that feeds internally on plant material, though specific host associations for this species remain undocumented in the provided sources.
Cydia obnisa
Cydia obnisa is a species of tortricid moth described by Heinrich in 1926. It belongs to the genus Cydia, which includes numerous economically significant agricultural pests such as the codling moth and filbertworm. The species is classified within the subfamily Olethreutinae and tribe Grapholitini. Available information about this species is extremely limited, with only three documented observations on iNaturalist and no published biological studies or host records identified in the provided sources.
Cydia piperana
Ponderosa Pine Seedworm Moth
Cydia piperana, commonly known as the ponderosa pine seedworm moth, is a small tortricid moth native to southwestern North America. The species is specialized on conifer hosts, with larvae developing within the seeds of Pinus ponderosa and Pinus jeffreyi. Adults are modest in size with a wingspan of approximately 19 mm. The moth is of ecological interest as a seed predator in pine ecosystems and has potential relevance to forest health monitoring, though it is not considered a major economic pest.
Cydia populana
Cydia populana is a tortricid moth species described by Busck in 1916. It belongs to the genus Cydia, which includes several economically significant agricultural pests such as the codling moth (C. pomonella) and the filbertworm (C. latiferreana). The species has been recorded from prairie provinces in Canada.
Cydia prosperana
Cydia prosperana is a small tortricid moth in the family Tortricidae, subfamily Olethreutinae. First described by Kearfott in 1907 as Enarmonia prosperana, it belongs to the large genus Cydia, which contains numerous agricultural pest species including the codling moth (C. pomonella) and filbertworm (C. latiferreana). The species is known from limited records in North America.
Cydia rana
Cydia rana is a species of tortricid moth described by Forbes in 1924. It belongs to the genus Cydia, which includes several economically significant fruit-feeding moths. Very little published information is available regarding the specific biology, ecology, or economic importance of this particular species.
Cydia tana
Cydia tana is a species of tortricid moth in the family Tortricidae, first described by Kearfott in 1907. It belongs to the genus Cydia, which includes numerous species of small moths commonly known as tortrix moths or leafroller moths. Like other members of this genus, C. tana is likely associated with woody plants, though specific host relationships remain poorly documented. The species is rarely encountered in collections and has minimal published biological information.
Cydia timara
Cydia timara is a species of tortricid moth described by Brown in 2014. It belongs to the genus Cydia, a group that includes several economically significant agricultural pests such as the codling moth (C. pomonella) and the filbertworm (C. latiferreana). As a recently described species, detailed biological and ecological information remains limited.
Cydia toreuta
Eastern Pine Seedworm Moth
Cydia toreuta, commonly known as the eastern pine seedworm moth, is a small tortricid moth native to North America. The species is a specialist seed predator, with larvae feeding exclusively on the seeds of red pine (Pinus resinosa) and jack pine (Pinus banksiana). It is considered a minor forest pest that reduces seed viability in conifer stands.
Cydia youngana
Cydia youngana is a species of tortricid moth in the family Tortricidae, subfamily Olethreutinae. The species was described by Kearfott in 1907. It belongs to the genus Cydia, which includes several economically significant agricultural pests such as the codling moth (C. pomonella) and the filbertworm (C. latiferreana). Very little specific information is available about the biology, appearance, or ecology of C. youngana.
Decodes
Decodes is a genus of tortricid moths in the subfamily Tortricinae, tribe Cnephasiini. The genus was established by Obraztsov in 1961 and contains approximately 20 described species distributed primarily in North America. Many species were described by Powell and colleagues between 1961 and 1998. The genus is part of the diverse leafroller moth family Tortricidae, whose larvae typically feed on plants.
Decodes asapheus
Decodes asapheus is a species of tortricid moth described by Jerry A. Powell in 1980. It belongs to the tribe Cnephasiini within the subfamily Tortricinae. The genus Decodes is endemic to the Hawaiian Islands, and this species is one of several described from this isolated archipelago. Like other Cnephasiini, it is presumed to have a larval stage that feeds on plant material, though specific host records for this species are not documented.
Decodes basiplagana
Decodes basiplagana is a tortricid moth species found across eastern and central North America. Adults have forewings measuring 8.1–9 mm in length. The species is notable for its larval association with Quercus lobata (valley oak), indicating a specialized host relationship. It has been documented from Ontario south to Oklahoma and Tennessee, and from Massachusetts west to Minnesota and Wisconsin.
Decodes fragariana
Decodes fragariana is a species of tortricid moth in the tribe Cnephasiini. Originally described by Busck in 1919 as Tortricodes fragariana, it was later transferred to the genus Decodes. The species is a member of the large family Tortricidae, which includes many economically significant agricultural pests. As a tortricine moth, it likely exhibits the characteristic bell-shaped resting posture with wings folded roof-like over the abdomen.
Decodes helix
Decodes helix is a species of tortricid moth described by Powell & Brown in 1998. It belongs to the family Tortricidae, a large group of small moths commonly known as leafroller moths. The genus Decodes is part of the tribe Cnephasiini within the subfamily Tortricinae. Very little specific information is documented about this species.
Decodes macdunnoughi
Decodes macdunnoughi is a species of tortricid moth in the tribe Cnephasiini, described by Jerry A. Powell in 1980. The genus Decodes contains approximately 20 described species distributed primarily in western North America. As a member of the leafroller moth family Tortricidae, this species likely exhibits the typical resting posture with wings held roof-like over the body.
Dichelia
Dichelia is a genus of tortrix moths in the tribe Archipini, established by Guenée in 1845. The genus includes at least five described species distributed across Europe, North Africa, and the Mediterranean region. One species, D. cedricola, has been documented as a significant pest of Lebanese cedar (Cedrus libani), causing needle drop and bud damage through larval feeding. Larvae construct silk-bound needle nests and overwinter in protective silk coverings on host trees.
Dichrorampha
Dichrorampha is a genus of small tortricid moths in the subfamily Olethreutinae, comprising over 100 described species distributed across Europe, Asia, and North America. The genus includes both diurnal and crepuscular species, many associated with mountain and alpine habitats. Several species have been investigated as biological control agents for invasive weeds, including D. aeratana for oxeye daisy and D. odorata for Chromolaena odorata. The genus has been subject to recent taxonomic revision, with cryptic species discovered through integrative approaches combining morphology and molecular data.
Dichrorampha acuminatana
Sharp-winged Drill
Dichrorampha acuminatana is a small tortricid moth with a wingspan of 10–15 mm, found across Europe and the Near East. The species exhibits distinctive wing patterns with dark fuscous ground color, purplish tints, and ochreous irroration. Adults are active from May to September, with larvae feeding on Leucanthemum vulgare (oxeye daisy) and Tanacetum vulgare (tansy). The species has been well-documented in Belgium and the Netherlands, with over 2,000 iNaturalist observations.
Dichrorampha aeratana
Obscure Drill
Dichrorampha aeratana is a small tortricid moth native to Europe, first described in 1915. The species has gained scientific attention as a potential biological control agent for the invasive weed oxeye daisy (Leucanthemum vulgare) in North America and Australia. Adults are active in spring, with larvae feeding internally on host plant roots and rhizomes. The moth exhibits strong host specificity, with larval survival negatively associated with increasing ploidy levels in host plants.
Dichrorampha bittana
Dichrorampha bittana is a small tortricid moth species originally described as Hemimene bittana by Busck in 1906. It belongs to the diverse leaf-roller family Tortricidae, a group containing numerous agricultural pests and species with specialized host plant relationships. The species is recorded from the Canadian provinces of Alberta and Saskatchewan, placing it within the Nearctic fauna. Like many Dichrorampha species, it likely exhibits specific ecological associations, though detailed biological studies remain limited.
Dichrorampha broui
Dichrorampha broui is a species of tortricid moth described by Knudson in 1987. It belongs to the genus Dichrorampha, a group of leaf-roller moths within the family Tortricidae. The species is part of the diverse Lepidoptera fauna, with limited published information available regarding its specific biology and distribution. Like other members of its genus, it is likely associated with herbaceous host plants.
Dichrorampha caribe
Dichrorampha caribe is a species of tortricid moth described by Brown in 2020. It belongs to the genus Dichrorampha, a group of small leaf-roller moths within the family Tortricidae. The specific epithet "caribe" suggests a Caribbean association, though detailed information about this species remains limited in available sources. Like other members of its genus, it is likely a small moth with typical tortricid morphology.
Dichrorampha flinti
Dichrorampha flinti is a species of tortricid moth in the family Tortricidae, first described by Razowski in 2011. It belongs to the genus Dichrorampha, a group of small moths commonly known as leaf-rollers. The species is part of the diverse Lepidoptera fauna that inhabits various regions, though specific details about its biology and distribution remain limited in available literature. As with many recently described tortricid species, further research is needed to fully understand its ecological requirements and life history.
Dichrorampha leopardana
Dichrorampha leopardana is a species of tortricid moth in the family Tortricidae, subfamily Olethreutinae, tribe Grapholitini. It was described by Busck in 1906. The genus Dichrorampha contains numerous small moths, many of which are associated with alpine and montane habitats. Related species in this genus have been subjects of recent taxonomic revision due to cryptic diversity and long-standing confusion with similar species.
Dichrorampha manilkara
Dichrorampha manilkara is a species of tortricid moth in the family Tortricidae, subfamily Olethreutinae, and tribe Grapholitini. It was described by Heppner in 1981. The genus Dichrorampha comprises small moths commonly known as leaf-roller moths, many of which are associated with specific host plants. This species belongs to a taxonomically complex genus where species identification often requires examination of genitalia morphology and genetic analysis.
Dichrorampha sapodilla
Dichrorampha sapodilla is a small moth in the leaf-roller family Tortricidae, described by John B. Heppner in 1981. The species is known only from Florida, USA, and appears to be rarely encountered, with only one documented observation in public databases. Like other members of its genus, it likely has a specialized association with particular host plants, though specific details remain undocumented.
Dichrorampha sedatana
Dingy Drill
Dichrorampha sedatana is a small tortricid moth with a wingspan of 12–16 mm. Adults are active from April to June, with one generation per year. The larvae are root feeders, specializing on Asteraceae hosts including Tanacetum vulgare and related species. The species is widespread across Europe and occupies rough meadows and wayside habitats.
Dichrorampha simulana
A small tortricid moth species described by Clemens in 1860, originally placed in the genus Halonota. Records indicate occurrence in the northeastern United States, specifically Vermont. As a member of the genus Dichrorampha, it belongs to a group of small leaf-roller moths whose larvae typically develop within plant tissues.
Diedra
Diedra is a genus of tortricid moths established by Rubinoff & Powell in 1999. The genus contains five described species, all native to western North America. Most species were described from California and associated with coniferous host plants. The genus is placed in the tribe Archipini within the subfamily Tortricinae.
Diedra cockerellana
Cockerell's Moth
Diedra cockerellana, known as Cockerell's Moth, is a species of tortricid moth described by Kearfott in 1907. It belongs to the tribe Archipini within the subfamily Tortricinae. The species has been documented across a broad North American range spanning multiple ecoregions from the Great Lakes to the western United States and southern Canada.