Rarely-collected
Guides
Dinardilla liometopi
Dinardilla liometopi is a small rove beetle described by Wasmann in 1901. It belongs to the tribe Sceptobiini within the large subfamily Aleocharinae. The species occurs in the southwestern United States and Mexico. Very few observations exist in public databases, indicating it is rarely collected or potentially cryptic in habit.
Distichona
Distichona is a genus of tachinid flies in the family Tachinidae, subfamily Exoristinae, tribe Goniini. The genus was established by Wulp in 1890 and contains eight described species distributed across the Americas. As a member of the Tachinidae, species in this genus are presumably parasitoids, though specific host associations remain undocumented. The genus is rarely encountered, with minimal observational records available.
Dryocoetes granicollis
Dryocoetes granicollis is a small bark beetle in the family Curculionidae, subfamily Scolytinae. The species was described by John Lawrence LeConte in 1876. It occurs in North America, with records from Canada including Manitoba and Québec. As a member of the genus Dryocoetes, it is associated with woody plants, though specific host associations for this species remain poorly documented.
Dyscharachthis
Dyscharachthis is a genus of false click beetles in the family Eucnemidae, established by Blackburn in 1900. It belongs to the monotypic tribe Dyscharachthini. The genus is poorly known, with minimal published information on its biology and morphology. Based on family characteristics, members are likely associated with decaying wood habitats.
Dysmerus
Dysmerus is a genus of small beetles in the family Laemophloeidae, comprising 15 species distributed across the New World from the southeastern United States to Bolivia and the West Indies. The genus remained monotypic for over a century until a 2025 revision resurrected two synonymized species and described 12 new species. Males possess a grotesquely modified antennal scape of unknown function, a diagnostic feature of the genus. These beetles are rarely encountered, though adults occasionally come to light at night.
Echthodopa carolinensis
Echthodopa carolinensis is a species of robber fly (family Asilidae) described by Bromley in 1951. It belongs to a genus of predatory flies that capture prey in flight. The species epithet 'carolinensis' suggests a connection to the Carolinas region of the United States, though specific details about its biology and ecology remain poorly documented in available sources.
Efferia frewingi
Columbia River Hammertail
Efferia frewingi is a species of robber fly (family Asilidae) described by Wilcox in 1966. Based on the genus Efferia, it belongs to a diverse group of predatory flies known for their robust bodies, large eyes, and aerial hunting behavior. The species epithet "frewingi" honors an individual, following the patronymic naming convention common in entomology. The common name "Columbia River Hammertail" suggests an association with the Columbia River region, though specific ecological details remain poorly documented in accessible literature.
Elachodelphax
Elachodelphax is a genus of planthoppers in the family Delphacidae, described by Vilbaste in 1965. Members belong to the tribe Delphacini within the subfamily Delphacinae. These insects are small, sap-feeding true bugs associated with grasses and sedges. The genus appears to be rarely collected, with minimal documentation in public databases.
Elachodelphax indistinctus
Elachodelphax indistinctus is a small planthopper in the family Delphacidae, described by Crawford in 1914. The species is known from a limited number of records across the western Great Plains and Rocky Mountain regions of North America. Like other delphacids, it is presumed to feed on grasses, though specific host associations remain undocumented.
Elachodelphax paransera
Elachodelphax paransera is a species of planthopper in the family Delphacidae, first described by Beamer in 1948. It belongs to a genus of small, slender delphacids found in North America. The species has documented occurrence records across the northern United States and southern Canada. Like other delphacids, it is presumed to feed on grasses and sedges, though specific host associations remain unconfirmed. The species appears to be rarely encountered, with limited biological data available.
Elassoptes
Elassoptes is a monotypic genus of true weevils (family Curculionidae) containing a single species, Elassoptes marinus. The genus was established by George Henry Horn in 1873. As a member of the largest beetle family, it possesses the characteristic elongated rostrum (snout) typical of weevils. The genus is rarely encountered in collections and field observations, with limited biological data available.
Ephedrodoma multilineata
Ephedrodoma multilineata is a species of plant bug in the family Miridae, described by Polhemus & Polhemus in 1984. It belongs to the diverse true bug fauna of western North America. The species epithet "multilineata" suggests multiple longitudinal lines or stripes on the body, a common naming convention in mirids with distinctive dorsal patterning. Very few observations exist in public databases, indicating it may be uncommon, cryptic, or undercollected.
Ephelinus notatus
A small rove beetle in the family Staphylinidae, Ephelinus notatus is a rarely encountered species with limited documented ecological information. The genus Ephelinus belongs to the subfamily Omaliinae, a group of rove beetles often associated with decaying organic matter and forest floor habitats. Based on its taxonomic placement and the distribution records from eastern and central United States, this species likely inhabits moist woodland environments where other Omaliinae species are commonly found.
Eropterus arculus
Eropterus arculus is a net-winged beetle (family Lycidae) described by Green in 1951. The species is known from eastern Canada, with records from New Brunswick, Ontario, and Québec. Like other lycids, adults are characterized by soft, flexible elytra with distinct reticulate venation. The species is rarely encountered, with minimal published biological information available.
Eucosma gemellana
Eucosma gemellana is a small tortricid moth described by Heinrich in 1923. The species belongs to the large genus Eucosma, which contains numerous seed-feeding tortricids associated with Asteraceae and related plant families. This species is rarely encountered and poorly documented in the scientific literature, with only a handful of observations recorded.
Eulasiona
Eulasiona is a genus of tachinid flies established by Townsend in 1892. The genus currently contains 12 described species distributed primarily in the Nearctic and Neotropical regions. As members of the family Tachinidae, these flies are parasitoids, though specific host associations for most Eulasiona species remain undocumented. The genus is classified in the subfamily Dexiinae and tribe Voriini.
Eurhopalothrix floridana
Eurhopalothrix floridana is a species of ant in the subfamily Myrmicinae, originally described by Brown & Kempf in 1960. It belongs to the genus Eurhopalothrix, a group of small, cryptic ants within the Attini tribe. The species is known from the Dominican Republic, with records from the Jardín Botánico Nacional in Santo Domingo de Guzmán. Very little biological information has been published about this species; it has been mentioned in taxonomic literature primarily in comparative studies of male morphology.
Euxesta abana
Euxesta abana is a species of picture-winged fly in the family Ulidiidae, described by Charles Howard Curran in 1935. It is known from a single documented location at White Sands National Park in the Tularosa Basin of New Mexico. The species belongs to a genus whose members are commonly known as picture-winged flies due to their patterned wings. Beyond its type locality and taxonomic placement, little is documented about its biology or ecology.
Exiliscelis californiensis
Exiliscelis californiensis is a species of fly in the family Canthyloscelididae, a small group of primitive flies often referred to as "perissommatid-like" or "synneurid" flies. Described by Hutson in 1977, this species is endemic to California. The family Canthyloscelididae is poorly known biologically, with most species known from few specimens. Exiliscelis californiensis is one of only three documented observations on iNaturalist, indicating it is rarely encountered or underreported.
Geomysaprinus cheyennensis
Cheyenne Clown Beetle
Geomysaprinus cheyennensis is a species of clown beetle (family Histeridae) described by Casey in 1916. The species is known from the northern Great Plains region of North America, with records from prairie provinces of Canada and adjacent U.S. states. As a member of the genus Geomysaprinus, it is likely associated with burrowing mammals, though specific ecological details remain poorly documented.
Gertschanapis shantzi
Gertschanapis shantzi is a minute araneomorph spider and the sole species in its genus. It belongs to the family Anapidae, a group of small spiders often associated with leaf litter and moss habitats. The species was first described from specimens collected in the United States and remains poorly known due to its cryptic size and habitat.
Ginglymocladus
false soldier beetles
Ginglymocladus is a genus of false soldier beetles in the family Omethidae, established by Van Dyke in 1918. The genus contains at least two described species: G. discoidea and G. luteicollis. Members of this family are commonly referred to as false soldier beetles due to their resemblance to soldier beetles (Cantharidae), though they belong to a distinct lineage. The family Omethidae is relatively small and poorly studied, with limited ecological and behavioral documentation.
Gloridonus chiragricus
Gloridonus chiragricus is a leafhopper species in the family Cicadellidae, first described by Ball in 1900. It belongs to the tribe Cicadellini, a group of robust, often brightly colored leafhoppers. The species is known from limited records in Arizona, with few documented observations. Like other members of its genus, it likely inhabits arid and semi-arid environments. Specific ecological details remain poorly documented due to its rarity in collections and limited study.
Glyphocystis viridivallis
Glyphocystis viridivallis is a species of snout moth in the family Pyralidae. It is the sole member of the monotypic genus Glyphocystis. The species was described by André Blanchard in 1973 and is known only from Texas. Very little is known about its biology or ecology.
Glyphogaster
Glyphogaster is a genus of parasitoid wasps in the family Braconidae, first described in 1951. As members of the Braconidae, these wasps are presumed to be parasitoids of other insects, though specific host associations remain poorly documented. The genus is rarely encountered in collections and has received limited taxonomic study.
Glyptocolastes
Glyptocolastes is a genus of parasitoid wasps in the family Braconidae, established by Ashmead in 1900. Members of this genus belong to the large and diverse ichneumonoid wasp lineage. The genus contains relatively few documented species and is poorly represented in biological collections and literature.
Grotella vauriae
Grotella vauriae is a moth species in the family Noctuidae, first described by Rowland R. McElvare in 1950. The species is known from Texas, which serves as its type locality. It belongs to the genus Grotella, a group of owlet moths within the subfamily Grotellinae. Very little is known about the biology or ecology of this species.
Haeterius blanchardi
clown beetle
Haeterius blanchardi is a myrmecophilous clown beetle (family Histeridae) native to the eastern United States. The species is known to inhabit colonies of the ant Formica pallidefulva, though its precise ecological relationship with the host remains incompletely documented. Adults have been recorded from Wisconsin, Massachusetts, New Jersey, Connecticut, Pennsylvania, and Virginia. The larvae have not been formally described.
Haplopogon erinus
Haplopogon erinus is a species of robber fly described by Pritchard in 1941. It belongs to the genus Haplopogon, a group within the Asilidae family characterized by reduced wing venation and distinctive predatory habits. The species is known from limited collection records and remains poorly documented in the scientific literature.
Heimbrinae
Heimbrinae is a small subfamily of chalcidoid wasps within the family Eurytomidae, established by Burks in 1971. Members of this subfamily are parasitoid wasps, though specific host associations remain poorly documented. The subfamily is rarely encountered, with only seven observations recorded on iNaturalist as of the source date. Heimbrinae represents one of several subfamilies within Eurytomidae, a family known for diverse life histories including both parasitoidism and phytophagy.
Hesperolinus
Hesperolinus is a genus of rove beetles in the family Staphylinidae, tribe Xantholinini. It was established by Casey in 1906. The genus belongs to the diverse subfamily Staphylininae, one of the largest lineages within rove beetles. Members of this genus are poorly documented in modern literature and appear to be rarely collected.
Hesperorhipis albofasciata
Hesperorhipis albofasciata is a species of metallic wood-boring beetle in the family Buprestidae, tribe Xenorhipidini. It is one of four species in the genus Hesperorhipis, which is restricted to Arizona and California. The species exhibits pronounced sexual dimorphism in antennal structure and elytral abbreviation, with males possessing highly modified flabellate antennae and more abbreviated elytra than females. It has been reared exclusively from walnut (Juglans sp.).
Himmacia stratia
A small moth in the family Depressariidae, described by Ronald W. Hodges in 1974. Known from a single state in the southwestern United States, this species has a wingspan of approximately 20 mm and adult flight activity limited to mid-summer months.
Hockeria eriensis
Hockeria eriensis is a species of chalcid wasp in the family Chalcididae, first described by Wallace in 1942. It belongs to the subfamily Haltichellinae, a group characterized by reduced wing venation and often metallic coloration. The species is known from the Canadian prairies, with records from Alberta and Saskatchewan. As with other chalcidids, it is presumed to be a parasitoid, though specific host associations remain undocumented.
Homocerynea
Homocerynea is a monotypic moth genus in the family Erebidae, established in 1913 by William Barnes and James Halliday McDunnough. It contains a single species, Homocerynea cleoriformis, known only from Arizona. The genus has been classified in various subfamilies, including Phytometrinae within Erebidae and Acontiinae within Noctuidae, reflecting taxonomic uncertainty.
Hormoserphus
Hormoserphus is a genus of parasitoid wasps in the family Serphidae (Proctotrupoidea), established by Townes in 1981. The genus belongs to a poorly studied group of small, rarely collected wasps. Its taxonomic status is currently marked as doubtful in some databases, suggesting potential issues with its validity or circumscription. Members of Serphidae are generally associated with decaying organic matter where they parasitize other insects.
Hornia
Hornia is a genus of blister beetles (Meloidae: Nemognathinae) established by C.V. Riley in 1877. Species in this genus exhibit pronounced sexual dimorphism, with females being physogastric—the abdomen becomes greatly distended, an adaptation associated with reproductive biology. The genus is rarely collected and poorly known biologically, with most information derived from scattered historical records and limited laboratory observations.
Hyperaspidius
Hyperaspidius is a genus of lady beetles in the family Coccinellidae, established by Crotch in 1873. The genus contains at least 20 described species. One species, Hyperaspidius venustulus, has been documented in association with mealybugs on big cordgrass in coastal wetland habitats. Members of this genus are rarely collected and appear to be specialists on particular host plants and their associated herbivores.
Hyperaspidius venustulus
Eight-spotted Hyperaspidius
Hyperaspidius venustulus is a small lady beetle in the family Coccinellidae. Adults measure approximately 2.80 mm. The species has been recorded from Georgia and is associated with big cordgrass (Spartina cynosuroides) stands, where it has been found with the mealybug Dysmicoccus dennoi. It is rarely collected and poorly known.
Ischyropalpus pinalicus
Ischyropalpus pinalicus is a species of ant-like flower beetle in the family Anthicidae, first described by Casey in 1895. The species is known from limited records in North America and Middle America, with very few documented observations. As with other members of Anthicidae, it is likely a small, elongate beetle with ant-like appearance, though specific biological details remain poorly documented. The genus Ischyropalpus contains species associated with various terrestrial habitats, but detailed studies of I. pinalicus are lacking.
Ismaridae
Ismaridae is a small, relictual family of parasitoid wasps in the order Hymenoptera. The family contains two extant genera: Ismarus, with approximately 50–59 described species worldwide, and the fossil genus Lubomirus from Eocene amber. All species with known biology are hyperparasitoids that parasitize Dryinidae, which are themselves parasitoids of leafhoppers, planthoppers, and treehoppers. Formerly classified as a subfamily of Diapriidae, Ismaridae was elevated to family status based on distinct morphological differences, particularly the absence of a facial projection for antennal insertion and variable fusion of metasomal terga.
Ismarus
Ismarus is a genus of small parasitoid wasps in the family Ismaridae. These insects are hyperparasitoids, attacking Dryinidae wasps that parasitize leafhoppers, planthoppers, and treehoppers. The genus is rarely collected and poorly known, with species distributed across wooded habitats in Asia, Europe, and other regions. Two new Chinese species were described in 2023 from Yunnan Province.
Kybos andresia
Kybos andresia is a species of leafhopper in the family Cicadellidae, described by Ross in 1963. It belongs to the subfamily Typhlocybinae, a group characterized by small size and reduced wing venation. Records indicate presence in northeastern North America, including Canada and the United States. The species is known from very few observations, reflecting either genuine rarity or undercollection.
Laphystia sillersi
Laphystia sillersi is a species of robber fly (Diptera: Asilidae) described by Hull in 1963. The genus Laphystia comprises predatory flies distributed primarily in the Neotropical region. As with other Asilidae, this species is presumed to be an aerial predator of other insects. No specific biological studies of this species have been published.
Lemophagus curtus
Lemophagus curtus is a species of ichneumon wasp described by Townes in 1965. It belongs to the family Ichneumonidae, one of the largest families of parasitoid wasps. The species is recorded from Norway, Sweden, and the conterminous United States. Like other members of its genus, it is presumed to be a parasitoid, though specific host associations remain undocumented.
Maemactes
Maemactes is a genus of weevils in the family Curculionidae, established by Carl Johan Schoenherr in 1837. The genus is poorly documented in modern literature, with minimal published research on its species composition, biology, or ecology. Records indicate it belongs to the hyperdiverse weevil family, one of the largest families of organisms on Earth. The genus appears to be rarely collected, with very few observations in biodiversity databases.
Mallodrya
Mallodrya is a monotypic genus of beetles in the family Synchroidae, established by George Henry Horn in 1888. The genus contains a single described species, Mallodrya subaenea. These beetles belong to the small family Synchroidae, commonly referred to as synchroa bark beetles, within the superfamily Tenebrionoidea.
Malthomethes oregonus
Malthomethes oregonus is a species of beetle in the family Omethidae, described by Fender in 1975. The genus Malthomethes belongs to a small family of elateroid beetles whose members are poorly studied. This species is known from very few records, with only one observation documented in iNaturalist. The specific epithet "oregonus" suggests a type locality in Oregon, though comprehensive distribution data remains limited.
Mannophorus forreri
Mannophorus forreri is a longhorn beetle in the family Cerambycidae, described by Henry Walter Bates in 1885. It belongs to the tribe Trachyderini, a group known for often brightly colored and patterned species. The species is rarely encountered in collections and appears to have a restricted distribution in the southwestern United States and Mexico. Field observations indicate adults are active in early autumn and visit flowers of yellow composites in mountainous areas of Arizona.
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oncircumstancecontextenvironmentsettingscenebackgroundbackdropframeworkstructureorganizationarrangementsystemschemeplandesignpatternmodeltemplateprototypearchetypeoriginalinitialprimitiveearlyancientoldagedelderlyseniormaturegrownadvancedprogressedevolvedderiveddescendedoriginatedbegunstartedcommencedinitiatedlaunchedopenedinauguratedinstitutedestablishedfoundedset-upformedshapedfashionedmoldedcastforgedwroughtbuiltconstructedput-togethererectedraisedelevatedliftedhoistedheavedhauleddraggeddrawnpulledtuggedyankedjerkedsnatchedgrabbedseizedcaughttrappedsnarednettedhookedbaggedlandedgotgainedwonearnedachievedattainedreachedarrivedcomeappearedemergedmaterializedmanifestedshowedofferedgivengrantedbestowedconferredawardedgifteddonatedcontributeddividedsplitseparatedparteddetacheddisconnecteddisjoineddisuniteddivorcedalienatedestrangedisolatedsegregatedsequesteredsecludedwithdrawnretiredremovedtaken-awaycarried-offborne-awaytransportedconveyedtransferredtransmittedimpartedexpressedarticulatedverbalizedvocalizedvoicedspokensaiddepictedportrayedrepresentedillustrateddemonstratedshownrevealeddiscloseduncoveredunmaskedunveiledunfoldedspreadextendedstretchedspannedbridgedcrossedtraversedpassedgone-byelapsedexpiredlapsedendedfinishedcompletedconcludedterminatedclosedshutsealedlockedfastenedmade-safeprotectedguardeddefendedshieldedscreenedshelteredharboredhousedaccommodatedlodgedquarteredbilletedstationedpostedplacedpositionedlocatedsituatedsettledinstalledemplaceddepositedlaidputsetarrangedorderedorganizedstructuredsystematizedmethodizedrationalizedstreamlinedsimplifiedclarifiedexplainedinterpretedtranslatedrenderedparaphrasedrephrasedrewordedrestaterepeatreiteraterehearserecitequotecitereferalludementionremarkobservecommentdeclareannounceproclaimpronounceassertaffirmaveravowswearvowpledgepromisecommitengagebindobligecompelcoerceconstrainpressurgepushdrivepropelthrustshovesqueezecompresscondenseconcentratefocuscenterconvergemeetjoinunitecombinemergefuseblendmixmingleintermingleintermixintegrateincorporateembodycontainincludecompriseconsistcomposeconstituteformmakecreateproducegenerateyieldgiveprovidesupplyfurnishdeliverpresentofferextendproffertendersubmitproposesuggestrecommendadvisecounselguidedirectsteerpilotnavigateconductescortshepherdherdspurgoadpromptmotivatestimulateinspireencouragesupportbackhelpassistaidabetfacilitateeasesmoothsimplifyclarifyexplaininterprettranslaterenderconverttransformchangealtermodifyadjustadaptfitsuitmatchcorrespondagreeharmonizeconformcomplyadherestickclingholdgripseizegrabcatchcapturetrapsnarenethookbaglandsecureobtainacquiregetgainwinearnachieveattainreacharriveappearemergematerializemanifestexhibitgrantbestowconferawarddonatecontributeshareparticipateinvolvededicatedevoteattachfondlikeaffectwarmferventardentpassionatezealousenthusiasticeagerkeenintensevehementforcefulvigorousenergeticdynamicpowerfulstrongmightypotenteffectiveefficientcompetentcapableabletalentedskilledexpertproficientmasterfulcommandingcontrollingdominatingascendantsupremepredominantpreponderantsuperioradvantageousprogressiveforwardaheadonwardupwardrisingascendingclimbingmountingscalingsurmountingovercomingconqueringdefeatingvanquishingsubduingsubjugatingmasteringgoverningmanagingadministeringaccordingharmonizingconformingcomplyingobservingadheringstickingclingingholdinggraspinggrippingseizinggrabbingcatchingcapturingtrappingsnaringnettinghookingbagginglandingsecuringobtainingacquiringgettinggainingwinningearningachievingattainingreachingarrivingappearingemergingmaterializingmanifestingshowingdisplayingexhibitinggrantingbestowingconferringawardinggiftingdonatingcontributingparticipatingengaginginvolvingcommittingdedicatingdevotingattachingbeing-fondlovingaffectingbeing-tenderwarmingbeing-ferventbeing-ardentbeing-passionatebeing-zealousbeing-enthusiasticbeing-eagerbeing-keenbeing-intensebeing-vehementbeing-forcefulbeing-vigorousbeing-energeticbeing-dynamicbeing-powerfulbeing-strongbeing-mightybeing-potentbeing-effectivebeing-efficientbeing-competentbeing-capablebeing-ablebeing-aptbeing-talentedbeing-giftedbeing-skilledbeing-expertbeing-proficientbeing-masterfulbeing-commandingbeing-controllingbeing-dominatingbeing-ascendantbeing-supremebeing-predominantbeing-preponderantbeing-superiorbeing-advantageousbeing-leadingbeing-foremostbeing-frontbeing-vanbeing-vanguardbeing-advancedbeing-progressivebeing-forwardbeing-aheadbeing-onwardbeing-upwardbeing-risingbeing-ascendingbeing-climbingbeing-mountingbeing-scalingbeing-surmountingbeing-overcomingbeing-conqueringbeing-defeatingbeing-vanquishingbeing-subduingbeing-subjugatingbeing-masteringbeing-rulingbeing-governingbeing-managingbeing-administeringbeing-directingbeing-guidingbeing-steeringbeing-pilotingbeing-navigatingbeing-conductingbeing-escortingbeing-shepherdingbeing-herdingbeing-drivingbeing-urgingbeing-spurringbeing-goadingbeing-promptingbeing-motivatingbeing-stimulatingbeing-inspiringbeing-encouragingbeing-supportingbeing-backingbeing-helpingbeing-assistingbeing-aidingbeing-abettingbeing-facilitatingbeing-easingbeing-smoothingbeing-simplifyingbeing-clarifyingbeing-explainingbeing-interpretingbeing-translatingbeing-renderingbeing-convertingbeing-transformingbeing-changingbeing-alteringbeing-modifyingbeing-adjustingbeing-adaptingbeing-fittingbeing-suitingbeing-matchingbeing-correspondingbeing-agreeingbeing-accordingbeing-harmonizingbeing-conformingbeing-complyingbeing-observingbeing-adheringbeing-stickingbeing-clingingbeing-holdingbeing-graspingbeing-grippingbeing-seizingbeing-grabbingbeing-catchingbeing-capturingbeing-trappingbeing-snaringbeing-nettingbeing-hookingbeing-baggingbeing-landingbeing-securingbeing-obtainingbeing-acquiringbeing-gettingbeing-gainingbeing-winningbeing-earningbeing-achievingbeing-attainingbeing-reachingbeing-arrivingbeing-comingbeing-appearingbeing-emergingbeing-materializingbeing-manifestingbeing-showingbeing-displayingbeing-exhibitingbeing-presentingbeing-offeringbeing-givingbeing-grantingbeing-bestowingbeing-conferringbeing-awardingbeing-giftingbeing-donatingbeing-contributingbeing-sharingbeing-participatingbeing-engagingbeing-involvingbeing-committingbeing-dedicatingbeing-devotingbeing-attachingMatheteus
Matheteus is a monotypic genus of beetles in the family Omethidae, established by John Lawrence LeConte in 1874. The genus contains a single described species, M. theveneti. Omethidae is a small family of elateroid beetles, commonly referred to as false soldier beetles. Members of this family are generally poorly studied, with limited ecological and biological information available.