Olethreutinae
Guides
Epinotia subviridis
Epinotia subviridis is a species of tortricid moth in the family Tortricidae, subfamily Olethreutinae, described by Heinrich in 1929. The species belongs to the genus Epinotia, a diverse group of small moths commonly known as tortricids or leafroller moths. As with many Epinotia species, detailed biological information remains limited in published literature.
Epinotia trigonella
Birch Epinotia Moth
A small tortricid moth with a wingspan of 16–21 mm, found across Europe, the eastern Palearctic, and North America. Adults are active in late summer. The species is tightly associated with birch, with larvae feeding within folded or spun leaves.
Epinotia xandana
Epinotia xandana is a species of tortricid moth in the family Tortricidae, subfamily Olethreutinae. It was described by Kearfott in 1907. The species belongs to the large genus Epinotia, which contains numerous small moths commonly known as epinotias. Records indicate occurrence in Alberta, Canada.
Epinotia zandana
Epinotia zandana is a species of tortricid moth in the family Tortricidae, subfamily Olethreutinae. Originally described by Kearfott in 1907 as Eucosma zandana, it was later transferred to the genus Epinotia. Like other members of this genus, it is likely a small moth associated with coniferous hosts, though specific biological details remain poorly documented in published literature.
Episimus
Episimus is a genus of tortricid moths in the subfamily Olethreutinae, established by Walsingham in 1892. The genus contains approximately 40 described species distributed primarily in the Neotropical region, with some species ranging into North America. At least one species, E. unguiculus (syn. E. utilis), has been extensively studied as a candidate biological control agent for the invasive Brazilian peppertree (Schinus terebinthifolius) in Florida. The genus was taxonomically revised in 2008, with twelve new species described from the Neotropics and the synonymization of Episimoides.
Episimus argutana
Sumac Leaftier Moth
Episimus argutana, commonly known as the Sumac Leaftier Moth, is a species of tortricid moth in the family Tortricidae. The common name refers to the larval habit of tying leaves together with silk. The species is known from parts of eastern North America.
Episimus augmentana
Episimus augmentana is a species of tortricid moth in the subfamily Olethreutinae, first described by Zeller in 1877. It belongs to a genus whose members are primarily Neotropical in distribution. The species is documented in taxonomic databases but remains poorly studied in terms of natural history.
Episimus transferrana
Episimus transferrana is a species of tortricid moth in the family Tortricidae, described by Francis Walker in 1863. It belongs to the subfamily Olethreutinae, a group known for including many internal-feeding larvae. The species has been recorded from the Galápagos Islands and the Brazilian state of Amazonas (BR-AM). Records remain sparse, with limited biological data available.
Ethelgoda texanana
Ethelgoda texanana is a small tortricid moth in the subfamily Olethreutinae, tribe Grapholitini. The species was described from Texas in 1879 and belongs to a genus containing approximately five described species distributed in the Nearctic and Neotropical regions. As a member of the Grapholitini, it is likely associated with woody plants, though specific biological details remain poorly documented in published literature.
Eucopina cocana
Shortleaf Pine Coneborer Moth
Eucopina cocana, commonly known as the Shortleaf Pine Coneborer Moth, is a small tortricid moth in the subfamily Olethreutinae. The species is associated with pine cones, specifically those of shortleaf pine (Pinus echinata), and develops as a larval borer within cone tissues. Adults are small, nondescript moths with wingspans typical of the genus. The species is documented primarily from the southeastern United States where its host tree occurs.
Eucopina monitorana
Red Pine Coneborer Moth
Eucopina monitorana, commonly known as the Red Pine Coneborer Moth, is a species of tortricid moth first described by Heinrich in 1920. It belongs to the subfamily Olethreutinae and tribe Eucosmini. The species is associated with coniferous hosts, particularly pines, where larvae develop within cones. It has been documented across a limited number of observations, suggesting a more restricted or specialized distribution.
Eucopina ponderosa
Eucopina ponderosa is a species of tortricid moth in the subfamily Olethreutinae, described by Powell in 1968. The species epithet "ponderosa" suggests a likely association with ponderosa pine (Pinus ponderosa), a common pattern among Eucopina species which are generally conifer-feeding specialists. The genus Eucopina contains small moths whose larvae typically develop as shoot borers or cone feeders on Pinaceae.
Eucopina rescissoriana
Lodgepole Pine Coneborer Moth
A small tortricid moth whose larvae bore into the cones of lodgepole pine (Pinus contorta). The common name reflects this specialized association. Adults are rarely observed directly; records primarily derive from larval collections and cone dissections. The species appears restricted to regions where its host pine occurs.
Eucopina siskiyouana
Fir Coneborer Moth
Eucopina siskiyouana, commonly known as the Fir Coneborer Moth, is a species of tortricid moth in the tribe Eucosmini. First described by Kearfott in 1907, this species is associated with coniferous habitats, particularly fir trees. The common name suggests larval feeding within cones, a trait shared with related Eucopina species. It occurs in western North America.
Eucosma
Eucosma is a large genus of small tortricid moths with at least 670 described species worldwide. The genus has a Holarctic and Indomalayan distribution, with approximately 150 species in the Nearctic region and 53 in Europe. New species continue to be discovered even in well-studied regions. Species exhibit considerable variation in coloration, ranging from plain to boldly patterned.
Eucosma artemisiana
Eucosma artemisiana is a species of tortricid moth described by Walsingham in 1879. It belongs to the genus Eucosma, a large group of small moths commonly known as bell moths or gall moths. The species epithet 'artemisiana' suggests an association with Artemisia host plants. Like other members of the subfamily Olethreutinae, it likely has larvae that feed internally within plant tissues.
Eucosma awemeana
Eucosma awemeana is a species of tortricid moth in the family Tortricidae, described by William D. Kearfott in 1907. The species belongs to the large genus Eucosma, which contains numerous small moths commonly known as phaneta or eucosma moths. As with most members of Olethreutinae, the larvae are likely internal feeders within plant tissues, though specific host associations for this species remain poorly documented.
Eucosma bipunctella
Eucosma bipunctella is a small tortricid moth described by Francis Walker in 1863. As a member of the genus Eucosma, it belongs to a diverse group of moths commonly known as phaneta or eucosma moths. The specific epithet 'bipunctella' refers to two spots, likely describing a diagnostic wing pattern. Like other Olethreutinae, the larvae are likely internal feeders in plant tissues.
Eucosma browni
Eucosma browni is a species of tortricid moth described by Wright in 2010. It belongs to the genus Eucosma, a diverse group of small moths commonly known as phaneta moths or eucosma moths. The species is classified within the subfamily Olethreutinae and tribe Eucosmini. As a recently described species, detailed biological information remains limited.
Eucosma cana
Hoary Bell
Eucosma cana, commonly known as the hoary bell, is a small tortricid moth with a wingspan of 16–23 mm. It is widespread across Europe, parts of Asia including China and Japan, and Central Asia. The species is univoltine, with adults active from June to August. Larvae develop within the flowerheads of thistles and related plants, feeding on Cirsium, Carduus, and Centaurea nigra.
Eucosma canusana
Eucosma canusana is a small moth in the family Tortricidae, first described by Wright in 1997. As a member of the Olethreutinae subfamily, it belongs to a diverse group of tortrix moths commonly known as leafroller moths. The species is placed in the genus Eucosma, a large and taxonomically complex group within the tribe Eucosmini. Very little specific biological information has been published for this species.
Eucosma circulana
Folded-costa Eucosma Moth
Eucosma circulana is a species of tortricid moth in the subfamily Olethreutinae. It is one of approximately 300 described species in the genus Eucosma, a group commonly known as phaneta moths. The species was originally described by Jacob Hübner in 1823 under the basionym Tortrix circulana. Members of this genus are generally small moths with distinctive wing patterns and are frequently encountered in North American habitats.
Eucosma gemellana
Eucosma gemellana is a small tortricid moth described by Heinrich in 1923. The species belongs to the large genus Eucosma, which contains numerous seed-feeding tortricids associated with Asteraceae and related plant families. This species is rarely encountered and poorly documented in the scientific literature, with only a handful of observations recorded.
Eucosma glomerana
Eucosma glomerana is a small tortricid moth described by Walsingham in 1879. It belongs to the tribe Eucosmini within the subfamily Olethreutinae. Records indicate a restricted distribution in the Canadian prairie provinces.
Eucosma gomonana
Eucosma gomonana is a species of tortricid moth in the subfamily Olethreutinae, described by Kearfott in 1907. The species is provisionally accepted in current taxonomic databases and has been documented through limited observations. As a member of the genus Eucosma, it belongs to a group of small moths commonly known as bell moths or shoot moths, many of which are associated with specific host plants. Detailed biological information remains scarce due to limited study.
Eucosma grindeliana
Eucosma grindeliana is a tortricid moth species described by Busck in 1906, originally placed in the genus Cydia. It belongs to the Olethreutinae subfamily and is part of the diverse genus Eucosma, which contains numerous small moth species associated with Asteraceae hosts. The species has been documented in North America with 283 iNaturalist observations recorded. Specific biological details remain limited in published literature.
Eucosma indagatricana
Eucosma indagatricana is a species of tortricid moth described by Heinrich in 1923. It belongs to the subfamily Olethreutinae, a group commonly known as olethreutine moths. The species is part of the large genus Eucosma, which contains numerous small moths often associated with Asteraceae and other plant families. Limited published information exists regarding its biology and ecology.
Eucosma kiscana
Eucosma kiscana is a small moth species in the family Tortricidae, first described by Kearfott in 1907 under the basionym Thiodia kiscana. It belongs to the large genus Eucosma, a group of tortricid moths commonly known as phaneta or eucosma moths. The species has been documented in the northeastern United States, with confirmed records from Vermont. Like many Eucosma species, it is likely associated with herbaceous or woody plants, though specific biological details remain poorly documented.
Eucosma knudsoni
Eucosma knudsoni is a species of tortricid moth described from Texas in 2015. It belongs to a large genus of small moths commonly known as bell moths or eucosmas, many of which are associated with Asteraceae host plants. The species is known from very few observations, reflecting both its recent description and likely restricted distribution or cryptic habits. As a member of the Olethreutinae subfamily, it possesses the characteristic tortricid resting posture with wings held rooflike over the body.
Eucosma modernana
Eucosma modernana is a species of tortricid moth in the family Tortricidae, subfamily Olethreutinae. It was described by James Halliday McDunnough in 1925. The genus Eucosma comprises numerous small to medium-sized moths, many of which are associated with Asteraceae host plants. Specific ecological details for E. modernana remain poorly documented.
Eucosma monogrammana
Eucosma monogrammana is a species of tortricid moth in the subfamily Olethreutinae. It was described by Zeller in 1875. The species belongs to the large genus Eucosma, which comprises numerous small moths often associated with coniferous and deciduous trees. Specific biological details for this species remain poorly documented in published literature.
Eucosma montanana
Eucosma montanana is a species of tortricid moth first described by Walsingham in 1884. It belongs to the family Tortricidae (leafroller moths), a large and economically important group of moths. The species is placed in the subfamily Olethreutinae and tribe Eucosmini. As a member of the genus Eucosma, it is part of a diverse group of small moths, many of which are associated with specific host plants.
Eucosma ochrocephala
Pale-headed Eucosma Moth
Eucosma ochrocephala is a species of tortricid moth in the family Tortricidae, subfamily Olethreutinae. The species was first described by Walsingham in 1895 under the basionym Semasia ochrocephala. It is part of the diverse genus Eucosma, which contains numerous small moth species, many of which are associated with Asteraceae host plants. The species is recognized in major taxonomic databases including Catalogue of Life, GBIF, and NCBI Taxonomy.
Eucosma parmatana
Eucosma parmatana is a species of olethreutine leafroller moth in the family Tortricidae, first described by Clemens in 1860. It is native to North America and has been recorded from Vermont and other parts of the United States. The species is assigned MONA/Hodges number 2937.
Eucosma rupestrana
Eucosma rupestrana is a species of tortricid moth in the subfamily Olethreutinae, described by James Halliday McDunnough in 1925. It belongs to the large genus Eucosma, which comprises numerous small moths commonly known as phaneta or eucosma moths. Very little species-specific information has been published for this taxon.
Eucosma striatana
Striated Eucosma Moth
Eucosma striatana is a small tortricid moth in the tribe Eucosmini. The species was described from North America by Clemens in 1860, originally placed in the genus Anchylopera. It is part of a species complex within Eucosma that is challenging to distinguish morphologically. The common name refers to the striated or lined appearance of the forewings.
Eucosma tarandana
Eucosma tarandana is a species of tortricid moth in the subfamily Olethreutinae, first described by Heinrich Benno Möschler in 1874. It belongs to a large genus of small moths commonly known as bell moths or tortrix moths. The species is part of the Eucosmini tribe, whose larvae are predominantly associated with coniferous and deciduous trees.
Eucosma tenuiana
Eucosma tenuiana is a species of tortricid moth in the subfamily Olethreutinae, described by Walsingham in 1879. It belongs to the large genus Eucosma, which contains numerous small moths often associated with composite plants. The species is rarely encountered, with only 8 observations recorded on iNaturalist, suggesting it is either genuinely uncommon or underreported.
Eucosma viridis
Eucosma viridis is a species of tortricid moth described in 2010 by Wright and Gilligan. It belongs to the large genus Eucosma, which comprises numerous small moths commonly known as bell moths or leafroller moths. As a recently described species, detailed biological information remains limited.
Eumarozia
Eumarozia is a genus of tortricid moths in the subfamily Olethreutinae, established by Heinrich in 1926. The genus contains at least three described species: E. beckeri, E. elaeanthes, and E. malachitana. These moths are small to medium-sized with the characteristic bell-shaped wing posture typical of the family Tortricidae. The genus has been recorded in North America, with distribution records from the United States including Vermont.
Evora
Evora is a genus of tortricid moths described by Heinrich in 1926. It belongs to the subfamily Olethreutinae, a group of small to medium-sized moths commonly known as tortrix moths or leafroller moths. The genus contains multiple species, though specific details about individual species remain limited in published literature.
Evora hemidesma
Evora hemidesma is a species of tortricid moth described by Zeller in 1875. It belongs to the subfamily Olethreutinae and is one of few species in the genus Evora. The species has been recorded from prairie and northern regions of central North America, with specimens documented from Canadian provinces and the northeastern United States.
Grapholita edwardsiana
San Francisco Tree Lupine Moth
Grapholita edwardsiana, known as the San Francisco Tree Lupine Moth, is a species of tortricid moth in the family Tortricidae, subfamily Olethreutinae. The species was described by Kearfott in 1907 and is placed in the genus Grapholita, a diverse group of small moths commonly referred to as fruitworms or bud moths. Many Grapholita species are associated with leguminous plants, and the specific epithet "edwardsiana" suggests a possible association with lupines (Lupinus), particularly tree lupine. The species is part of the tribe Grapholitini, which includes numerous economically important pests of fruits and seeds.
Grapholita interstinctana
Clover Head Caterpillar Moth
Grapholita interstinctana is a tortricid moth species first described by Clemens in 1860. It is commonly known as the Clover Head Caterpillar Moth, reflecting its association with clover as a host plant. The species belongs to the Olethreutinae subfamily within the Tortricidae family, a group known for many economically significant agricultural pests. Records indicate presence in the northeastern United States, particularly Vermont.
Grapholitini
Grapholitine moths, Leafroller moth tribe
Grapholitini is a tribe of tortricid moths comprising approximately 1200 described species. The tribe contains numerous economically significant agricultural pests that damage fruits and seeds. Molecular phylogenetic analyses have resolved the tribe into two major lineages—a Dichrorampha clade and a Cydia clade—with the genus Grapholita found to be polyphyletic. The tribe likely originated in the Nearctic, Afrotropical and Neotropical regions during the middle Eocene (ca. 44.3 Ma).
Gretchena amatana
Gretchena amatana is a species of tortricid moth in the family Tortricidae, described by Carl Heinrich in 1923. It belongs to the subfamily Olethreutinae and tribe Eucosmini. The species is known from the northeastern United States, with distribution records from Vermont and surrounding areas. As with many small tortricid moths, detailed ecological information remains limited in published literature.
Gretchena concubitana
Gretchena concubitana is a species of tortrix moth in the family Tortricidae, described by Carl Heinrich in 1923. It belongs to the subfamily Olethreutinae and tribe Eucosmini. The genus Gretchena comprises small moths primarily distributed in North America. Records from iNaturalist indicate this species has been documented in at least 37 observations, suggesting it is not extremely rare, though detailed biological studies remain limited.
Gretchena delicatana
ironwood fruitworm
Gretchena delicatana is a species of tortricid moth described by Heinrich in 1923. It is commonly known as the ironwood fruitworm, reflecting its association with ironwood trees (Ostrya virginiana). The species belongs to the subfamily Olethreutinae and tribe Eucosmini. It has been documented in the northeastern United States, particularly Vermont.
Gretchena nymphana
Gretchena nymphana is a species of tortricid moth described by Blanchard and Knudson in 1983. It belongs to the subfamily Olethreutinae within the family Tortricidae, a large group of small to medium-sized moths commonly known as leafroller moths. The genus Gretchena is part of the tribe Eucosmini, which contains numerous species associated with various host plants. Information on the biology and ecology of this particular species remains limited in published literature.
Gretchena watchungana
Gretchena watchungana is a species of tortricid moth in the family Tortricidae, originally described as Epinotia watchungana by Kearfott in 1907. The species belongs to the subfamily Olethreutinae and tribe Eucosmini. It is part of a genus of small moths whose larvae are associated with various host plants. The specific epithet "watchungana" refers to the Watchung Mountains in New Jersey, indicating the type locality of the original specimen.