Tropical-diversity
Guides
Aderidae
ant-like leaf beetles
Aderidae is a family of small beetles commonly known as ant-like leaf beetles due to their resemblance to ants. The family contains approximately 1,000 species in about 40–50 genera, with greatest diversity in tropical regions but worldwide distribution. Adults are typically found on the undersides of leaves, while larvae inhabit rotting wood, leaf litter, and occasionally the nests of other insects. The oldest confirmed fossil member, Gryzmalia, dates to the mid-Cretaceous Burmese amber.
Brentidae
straight-snouted weevils, primitive weevils
Brentidae is a cosmopolitan family of weevils distinguished by their straight (non-elbowed) antennae and elongated, often flattened bodies. The family has undergone significant taxonomic expansion, now encompassing approximately 4,000 species across six subfamilies including groups formerly classified in Curculionidae (Apioninae, Cyladinae, Nanophyinae) and the previously separate family Ithycerinae. Members are primarily xylophagous, with larvae feeding on fungi in dead wood. The family exhibits its greatest diversity in tropical regions but extends into temperate zones worldwide.
Clusiidae
druid flies
A family of small acalyptrate flies (approximately 3.5 mm) commonly known as druid flies. The family comprises 636 species in 14 genera across three subfamilies, with highest diversity in tropical regions. Adults are typically found on tree trunks, while larvae inhabit decaying wood. Several genera exhibit lekking behavior and territorial defense by males.
Cremastinae
Cremastinae is a worldwide subfamily of parasitoid wasps within Ichneumonidae. Members are koinobiont endoparasitoids, primarily attacking concealed larvae of Lepidoptera. Several genera, including Pristomerus, have been used in biological control programs. The subfamily exhibits high diversity in tropical regions, with numerous undescribed species.
Diopsoidea
Diopsoidea is a small but diverse superfamily of acalyptrate muscoid flies with cosmopolitan distribution, particularly abundant in tropical regions. The group has experienced taxonomic revision, with Strongylophthalmyiidae and Tanypezidae now placed in Nerioidea. The superfamily includes families such as Diopsidae (stalk-eyed flies), Psilidae, and Syringogastridae.
Dipsocoromorpha
minute litter bugs
Dipsocoromorpha is an infraorder of minute true bugs within Heteroptera, containing approximately 300 species in one superfamily (Dipsocoroidea). Members are among the smallest adult true bugs, typically 0.5–4.0 mm in length. The group exhibits highest diversity in tropical regions, with many species remaining undescribed due to their cryptic habits and small size.
Emesinae
Thread-legged Bugs
Emesinae, commonly known as thread-legged bugs, are a subfamily of assassin bugs (Reduviidae) distinguished by their exceptionally slender body form. They are predatory insects that walk on their mid and hind legs while using their raptorial front legs to capture prey. The subfamily contains approximately 90 genera and 900 described species, with greatest diversity in tropical regions, particularly Africa. Some groups specialize on spiders as prey. Molecular phylogenetic analyses have questioned the monophyly of Emesinae, suggesting the group may be polyphyletic with respect to Saicinae and Visayanocorinae.
Helicopsyche
Speckled Peter, Spiral Caddisflies
Helicopsyche is a genus of caddisflies (Trichoptera: Helicopsychidae) containing more than 230 species with highest diversity in tropical and sub-tropical regions. The genus is represented on all major faunal regions worldwide. Larvae construct distinctive helical, snail-shell-like cases from sand grains and silk. Adults are known to fly fishers as the 'Speckled Peter'. The genus was first described by Siebold in 1856.
Hirtodrosophila
Hirtodrosophila is a genus of fruit flies in the family Drosophilidae, elevated from subgenus status within Drosophila by Grimaldi in 1990. The genus comprises approximately 150 described species divided into ten species groups, with highest diversity in tropical and subtropical regions. Most species are mycophagous, with larvae feeding on fungi, though at least one species has evolved predatory behavior on frog embryos. The genus belongs to the monophyletic Zygothrica genus group of mycophagous drosophilids, though Hirtodrosophila itself is paraphyletic.
Hydrometridae
marsh treaders, water measurers
Hydrometridae is a family of semiaquatic true bugs comprising over 147 species across seven genera and three subfamilies. Members are characterized by extremely elongated, slender bodies and heads that give them a measuring-stick appearance. They inhabit water surface margins of lakes, ponds, and wetlands worldwide, with greatest diversity in tropical regions. Some lineages have independently evolved terrestrial habits in insular Pacific habitats.
Lauxaniidae
Lauxaniid Flies
Lauxaniidae is a family of acalyptrate flies comprising approximately 1800 described species across 126 genera. Adults are small (2–7 mm), often with large, brightly colored compound eyes and variegated wing patterns. The family has a cosmopolitan distribution with highest diversity in tropical Asia and the Americas; species richness declines markedly toward temperate regions. Adults are typically sedentary, associated with shaded, humid forest environments where they feed on leaf-surface fungi. Larvae are primarily saprophagous, developing in decaying plant matter, leaf litter, rotting wood, and bird nests.
Lebiini
Lebiini is a tribe of ground beetles within the family Carabidae, containing over 250 genera and approximately 4,800 described species. Members exhibit considerable morphological diversity and occupy a wide range of habitats across tropical and subtropical regions worldwide. The tribe includes several subtribes such as Agrina, Dromiusina, Metallicina, Pericalina, and Physoderina.
Limnichidae
Minute Marsh-loving Beetles
Limnichidae is a family of small beetles in the superfamily Byrrhoidea, comprising at least 30 genera and 350 described species. The family exhibits considerable ecological diversity: while many species inhabit water-adjacent habitats such as riparian zones and coastal areas, others are fully terrestrial in leaf litter or arboreal environments. The subfamily Hyphalinae is uniquely specialized for intertidal marine habitats, with larvae capable of activity in seawater—a rarity among beetles. The oldest known fossils date to the mid-Cretaceous Burmese amber.
Micropezidae
stilt-legged flies
Micropezidae is a family of acalyptrate muscoid flies comprising approximately 500 species in 50 genera and five subfamilies. Commonly called stilt-legged flies, they are distinguished by extraordinarily elongated middle and hind legs, with markedly smaller fore legs. Adults range from 3–16 mm and exhibit remarkable mimicry of ants, wasps, and ichneumonid wasps. The family is most diverse in tropical and subtropical regions, particularly the Neotropics, and is absent from New Zealand and Macquarie Island. Larval biology remains poorly known, with development occurring in decaying vegetation, manure, fungi, and in some species, living plant roots.
Naucoridae
creeping water bugs, saucer bugs
Naucoridae is a family of aquatic predatory true bugs comprising approximately 400 species in 46 genera across five subfamilies. Members are commonly known as creeping water bugs or saucer bugs due to their dorsoventrally flattened, oval body shape. They inhabit freshwater environments worldwide with greatest diversity in tropical regions, ranging from still waters to torrential streams. Most species are lotic, occurring in flowing waters rather than lentic habitats. They are predatory, feeding on other small invertebrates using piercing-sucking mouthparts.
Oonopidae
Goblin spiders
Oonopidae, commonly known as goblin spiders, is a diverse family of minute spiders comprising approximately 2,000 described species across 115 genera worldwide, with total species diversity estimated at 2,000–2,500. These spiders are characterized by their tiny size (typically 1–3 mm), reduced eye number (usually six, though some species have four, two, or are completely eyeless), and frequently exhibit unusual morphological modifications in males. The family shows remarkable morphological diversity including hardened abdominal plates (scuta), modified mouthparts, sternal pouches, and elaborate pedipalp structures. Goblin spiders are predominantly ground-dwelling, inhabiting leaf litter, soil, and rock crevices, with some lineages adapted to canopy habitats, caves, or termite nests. The family has an extensive fossil record in amber deposits dating back over 100 million years, with Orchestina already widespread by the end of the Cretaceous.
Pangrapta
Pangrapta is a genus of moths in the family Erebidae, first described by Jacob Hübner in 1818. The genus contains approximately 80 described species distributed across tropical and subtropical regions worldwide. Members are classified in the subfamily Pangraptinae, a group of underwing and related moths. The genus shows notable diversity in Asia, with significant representation in Borneo, Japan, China, and the Indian subcontinent.
Phycitinae
Knot-horn Moths
Phycitinae is a subfamily of snout moths (Pyralidae) representing the most diverse lineage within its family, encompassing over 600 genera and more than 4,000 species—more than three-quarters of living snout moth diversity. Together with Epipaschiinae, they form the most advanced lineage of snout moths. Adults are typically small, slender-bodied moths with well-developed proboscises and often elongated labial palps forming a 'snout.' The subfamily exhibits remarkable ecological diversity, with larvae occupying roles as leaf rollers, seed feeders, gall inquilines, aquatic predators, and agricultural pests.
Platystomatidae
Signal Flies
Platystomatidae, commonly known as signal flies, is a family of acalyptrate Diptera in the superfamily Tephritoidea. The family comprises approximately 1200 species in 127 genera, with highest diversity in tropical regions, particularly the Australasian and Afrotropical realms. Adults are characterized by distinctive wing patterns and often metallic coloration. Many species exhibit elaborate sexual dimorphisms, including extreme head modifications in males used in agonistic interactions. The family is notable for convergent evolution of eyestalks with Diopsidae, though developed through different morphological pathways.
Psocidae
Common Barklice
Psocidae is a family of barklice in the order Psocodea, distinguished by diagnostic wing venation where the areola postica is fused to the M-vein, forming a discoidal cell. The family contains approximately 86 genera and is widespread globally, with documented diversity in East Africa, Southeast Asia, Melanesia, and the Americas. Psocidae is closely related to Myopsocidae. Many species are arboreal, though specific ecological roles remain understudied.
Scaphidiinae
Shining Fungus Beetles
Scaphidiinae is a subfamily of rove beetles (Staphylinidae) commonly known as shining fungus beetles. The subfamily comprises over 2000 described species across 46 genera, with greatest diversity concentrated in tropical and subtropical forests. Members are strongly associated with fungi and slime molds, on which they feed and complete their development. The Neotropical region remains particularly understudied, with recent work revealing substantially higher diversity than previously recorded.
Scarabaeoidea
Scarabs, Stag Beetles, Dung Beetles, Fruit and Flower Chafers
Scarabaeoidea is a superfamily of beetles comprising approximately 35,000 described species, with around 200 new species described annually. It is the sole superfamily within the infraorder Scarabaeiformia. The group includes diverse families such as Scarabaeidae (scarabs, dung beetles, chafers), Lucanidae (stag beetles), Geotrupidae (earth-boring scarabs), and Bolboceratidae. Many species exhibit distinctive circularly polarized light reflection from their cuticles, produced by helicoidal stacks of chitin microfibrils. The superfamily is currently undergoing taxonomic revision at the family level.
Sciapodinae
Sciapodinae is a subfamily of long-legged flies (Dolichopodidae) distinguished by ancestral wing venation features, including a branched vein M1+2. The group exhibits high diversity in tropical and subtropical regions of the Southern Hemisphere, particularly South America, sub-Saharan Africa, the Indian subcontinent, Southeast Asia, and Australasia. Phylogeographic evidence supports a Gondwanan origin during the Early Cretaceous. The subfamily contains three tribes—Mesorhagini, Sciapodini, and Chrysosomatini—with approximately 40 genera.
Spilomelinae
Pearl Moths
Spilomelinae is a species-rich subfamily of Crambidae (crambid snout moths) containing approximately 4,180 described species in 351 genera worldwide, making it the most diverse group among pyraloid moths. Formerly treated as tribe Spilomelini within Pyraustinae, it was elevated to subfamily status based on phylogenetic studies. The subfamily includes numerous agricultural pests as well as the invasive box tree moth (Cydalima perspectalis).
Sterrhinae
Waves and Mochas
Sterrhinae is a large subfamily of geometer moths (Geometridae) comprising approximately 3,000 described species, with exceptional diversity in the tropics decreasing toward higher latitudes and elevations. The subfamily was established by Edward Meyrick in 1892 and includes the taxonomically challenging genera Idaea and Scopula, which together account for more than half of all species. Members are commonly known as 'waves' due to their characteristic wing patterns. The group exhibits notable morphological synapomorphies including forewing areoles and specific venation patterns that distinguish it from other geometrid subfamilies.