Tephritoidea
Guides
Platystomatidae
Signal Flies
Platystomatidae, commonly known as signal flies, is a family of acalyptrate Diptera in the superfamily Tephritoidea. The family comprises approximately 1200 species in 127 genera, with highest diversity in tropical regions, particularly the Australasian and Afrotropical realms. Adults are characterized by distinctive wing patterns and often metallic coloration. Many species exhibit elaborate sexual dimorphisms, including extreme head modifications in males used in agonistic interactions. The family is notable for convergent evolution of eyestalks with Diopsidae, though developed through different morphological pathways.
Pyrgotidae
Scarab Pursuing Flies, Pyrgotid Flies
Pyrgotidae is a family of endoparasitoid flies within the superfamily Tephritoidea. Unusual among Cyclorrhapha, they lack ocelli entirely—one of only two families with this trait. Most species display patterned wings with bands or spots. Females pursue scarab beetles in flight to deposit eggs beneath the elytra, where larvae develop internally and eventually kill the host. The family has a worldwide distribution with approximately 67 species in Australia alone and 59 species in 13 genera in the Neotropics.
Richardiidae
Richardiidae is a small family of acalyptrate flies in the superfamily Tephritoidea, comprising over 30 genera and approximately 175 species. The family is predominantly Neotropical in distribution, with most species occurring in tropical regions of the Americas. General biology remains poorly known, though larvae of some species feed on living plant tissue, pollen, or decaying plant material. Adults are frequently observed on vegetation and are characterized by conspicuously patterned wings and often metallic body coloration.