Territorial-behavior
Guides
Protoxaea gloriosa
glorious protoxaea
Protoxaea gloriosa is a species of mining bee in the family Andrenidae, subfamily Oxaeinae. Males exhibit pronounced territorial behavior, establishing and defending flowering plants during morning hours to increase mating opportunities with virgin females. The species shows strong seasonality tied to summer rainfall patterns, with principal emergence triggered by first soaking rains. Females collect pollen primarily from Solanum and nectar from Asclepias, Verbesina, and Larrea. This bee is found in arid and semi-arid regions of the southwestern United States and Mexico.
Sphecius speciosus
Eastern cicada-killer wasp, cicada killer, cicada hawk
Sphecius speciosus is a large, solitary digger wasp and one of the largest wasps in eastern North America. Females hunt cicadas, paralyzing them with venom to provision underground nest cells for their larvae. Despite their formidable size and appearance, they pose minimal threat to humans—females rarely sting unless roughly handled, and males lack stingers entirely. The species exhibits pronounced sexual dimorphism, with females substantially larger to accommodate egg-laying and prey transport duties.
Stizus
sand wasps
Stizus is a genus of sand wasps in the family Crabronidae, containing over 100 species distributed across Europe, Africa, and North America. These solitary wasps are typically yellow and black, rarely reddish, and reach lengths of about 34 mm. Species nest in sandy soils and provision burrows with prey, primarily grasshoppers, to feed their larvae. Some species form nesting aggregations where males defend territories and engage in contest behavior to gain mating access to females.
Strymon
Scrub-Hairstreaks
Strymon is a genus of scrub hairstreak butterflies in the family Lycaenidae, comprising small, delicate butterflies found primarily in the Nearctic and Neotropical realms. The genus was historically treated as its own monotypic tribe (Strymonini) due to its distinct evolutionary lineage within the Eumaeini. Species such as the Gray Hairstreak (Strymon melinus) are among the most widespread and commonly encountered members of this group in North America.
Tachytes aurulentus
square-headed wasp, green-eyed wasp, sand-loving wasp
Tachytes aurulentus is a species of solitary square-headed wasp in the family Crabronidae. Like other members of its genus, it is characterized by notably large green eyes, particularly in males. The species is part of a diverse North American assemblage of approximately 35 Tachytes species. These wasps are ground-nesting predators that provision their burrows with paralyzed orthopteran prey for their offspring.
Tachytes sayi
Tachytes sayi is a solitary wasp in the family Crabronidae, tribe Larrini. It is one of approximately 35 North American species in the genus Tachytes. These wasps are characterized by large, often green eyes in males, and are commonly referred to as 'green-eyed wasps' or 'sand-loving wasps.' Females excavate burrows in soil to provision with paralyzed orthopteran prey for their larvae.
Tetramorium immigrans
Immigrant Pavement Ant, Pavement Ant, Sugar Ant
Tetramorium immigrans is a highly successful invasive ant native to Europe that has become one of the most abundant ant species in urban North America. The species thrives in disturbed, paved environments and exhibits remarkable thermal tolerance, with workers surviving temperatures up to 46°C. Colonies are typically large, with thousands to over 10,000 workers, and are highly territorial, engaging in conspicuous inter-colony battles. The species was previously misidentified as Tetramorium caespitum until a 2017 taxonomic revision clarified its status as a distinct species within the T. caespitum complex.
Tetramorium tsushimae
Japanese pavement ant
Tetramorium tsushimae, commonly known as the Japanese pavement ant, is a small myrmicine ant native to Asia that has become invasive in North America. Workers are approximately 2.5 mm in length. The species exhibits both monogyne and polygyne colony structures and is known for its territorial behavior, with colonies competing for high-temperature nesting sites to rear reproductives. It has been extensively used as a laboratory surrogate for the red imported fire ant (Solenopsis invicta) in toxicity and control studies. The species displays notable cadaveric behaviors on animal remains, creating scratched scars, mounds, and soil-covered nests that have forensic implications.
Toxomerus
Calligrapher Flies
Toxomerus is a large genus of hoverflies (family Syrphidae) distributed across the Americas, with over 130 described Neotropical species and additional Nearctic endemics. Species are small (typically 6–9 mm), characterized by large eyes with posterior indentation and diagnostic abdominal patterns. Most larvae are predatory on soft-bodied insects, particularly aphids, though at least three species are known pollen-feeders. Adults are pollinivorous, feeding on nectar and pollen from diverse flowering plants. The genus includes notable ecological specialists such as Toxomerus basalis, a kleptoparasite of sundews (Drosera), and several species introduced to the Afrotropics.
hoverflyflower-flyaphid-predatorpollinatorbiological-controlHymenoptera-mimickleptoparasiteintroduced-speciesNeotropicalNearcticAfrotropical-introductionpolyvoltineSyrphiniSyrphinaeDipteraSyrphidaeToxomerus-marginatusToxomerus-disparToxomerus-pulchellusToxomerus-basalisToxomerus-floralisToxomerus-politusToxomerus-apegiensisDroserasundewpollenivorypredatory-larvaeurban-adapteragricultural-beneficialcryptic-larvaeeye-dimorphismterritorial-behaviorDNA-barcodingcitizen-sciencemuseum-collectionsrange-expansionconservation-concernToxotrypana curvicauda
papaya fruit fly
Toxotrypana curvicauda, commonly known as the papaya fruit fly, is a tephritid fruit fly native to the Americas and now established in Florida and other parts of the southern United States. Males are highly territorial, defending papaya fruit surfaces through aerial combat using specialized midleg armature. Females are attracted to male-produced sex pheromones and oviposit directly into papaya fruits, where larvae develop within the seeds. The species is a significant agricultural pest of papaya (Carica papaya) and has been recorded on alternative hosts including Jacaratia mexicana and Morrenia odorata.
Xylocopa micans
Southern Carpenter Bee
Xylocopa micans, the southern carpenter bee, is a large solitary bee in the subgenus Schonnherria. It excavates nests in dead wood but, unlike the sympatric X. virginica, has not been documented nesting in structural timbers. The species exhibits a unique polymorphic mating strategy, shifting from resource defense polygyny in early spring to lek polygyny in mid-summer. It is an important generalist pollinator capable of buzz pollination.
Xylocopa virginica virginica
Virginia Carpenter Bee, Eastern Carpenter Bee
Xylocopa virginica virginica is a large carpenter bee native to eastern North America. It is facultatively social, forming small colonies with linear dominance hierarchies where one female monopolizes both reproduction and foraging. Females excavate tunnels in wood to create brood galleries, provisioning cells with bee bread (pollen and nectar mixture). Males defend nest-side territories and mate in early spring, then die; females may live two years. The subspecies is distinguished by the nominate form's distribution and morphology.