Ephemerellidae
Guides
Attenella margarita
spiny crawler mayfly
Attenella margarita is a species of spiny crawler mayfly in the family Ephemerellidae. It exhibits a disjunct east-west distribution pattern across North America, which is rare among mayflies. The species has been documented from the southern half of Canada and the continental United States, with a southeastern range extension to North Carolina representing approximately 1,300 km beyond previously known western localities.
Attenella soquele
Attenella soquele is a species of spiny crawler mayfly in the family Ephemerellidae. The species was described by Day in 1954 and is recognized as valid in current taxonomic databases. Members of the genus Attenella are characterized by their distinctive spiny abdominal tubercles and are primarily distributed in North America.
Caudatella
Caudatella is a genus of mayflies in the family Ephemerellidae, established by Edmunds in 1959. Species within this genus are primarily distributed in western North America, with documented occurrences in Montana and other regions. The genus has been subject to taxonomic revision, particularly regarding species complexes such as C. heterocaudata and C. hystrix. Larval stages are aquatic and associated with stream habitats.
Caudatella heterocaudata
A species of spiny crawler mayfly in the family Ephemerellidae, originally described by McDunnough in 1929. The species has undergone taxonomic revision, with Caudatella circia previously treated as distinct but now recognized as a synonym. Larvae possess diagnostic morphological characters that allow separation from the related C. hystrix complex.
Drunella cornutella
spiny crawler mayfly
Drunella cornutella is a species of spiny crawler mayfly in the family Ephemerellidae, originally described by McDunnough in 1931 as Ephemerella cornutella. It is found in North America, with records from the Nearctic region including the northeastern United States. Like other members of the genus Drunella, it is an aquatic insect with immature stages living in freshwater habitats.
Drunella doddsii
Western Green Drake
Drunella doddsii is a species of spiny crawler mayfly in the family Ephemerellidae, commonly known as the Western Green Drake. Originally described as Ephemerella doddsi by Needham in 1927, this species is significant to fly fishing communities in western North America. The species is part of the diverse Drunella genus, which includes other notable green drake species such as D. grandis.
Drunella flavilinea
Flav
Drunella flavilinea is a species of spiny crawler mayfly in the family Ephemerellidae. It is native to Central America and North America, with a range spanning southwestern Canada, the western United States, and northern Mexico. The species is known to fly fishermen by the common name "Flav" and is an important component of trout diets in western streams.
Drunella lata
Drunella lata is a species of spiny crawler mayfly in the family Ephemerellidae. It was first described by Morgan in 1911. The species is found in North America and belongs to a genus commonly known as "green drakes" among fly fishermen. Like other members of Ephemerellidae, it has aquatic nymphs and winged adults.
Drunella spinifera
Western Slate Olive
Drunella spinifera is a species of spiny crawler mayfly in the family Ephemerellidae. It is found in southwestern and northern Canada, the western United States, and Alaska. The common name "Western Slate Olive" reflects its regional distribution and likely coloration. Like other members of the genus Drunella, it belongs to a group of mayflies commonly targeted by fly fishermen, though specific angling references for this species are limited.
Drunella tuberculata
spiny crawler mayfly
Drunella tuberculata is a species of spiny crawler mayfly in the family Ephemerellidae, described by Morgan in 1911. It is found in North America and is closely related to D. allegheniensis, from which it can be distinguished by specific morphological characters. The species is part of a group of mayflies important to fly fishermen, where nymphs and adults serve as significant food sources for trout and other sport fish.
Drunella walkeri
Small Blue-winged Olive
Drunella walkeri is a species of spiny crawler mayfly in the family Ephemerellidae, commonly known as the Small Blue-winged Olive among anglers. It is found in North America and is one of the mayfly species frequently imitated by fly fishermen. The species was first described by Eaton in 1884.
Ephemerella aurivillii
spiny crawler mayfly
Ephemerella aurivillii is a spiny crawler mayfly in the family Ephemerellidae with a strict one-year life cycle. The species occurs across northern regions of the Northern Hemisphere, including Europe, northern Asia, and North America. Research in Norway demonstrated that its distribution and growth rates are strongly temperature-dependent, with populations expanding following hydropower-induced warming of previously cold streams.
Ephemerella dorothea
Pale Dun
Ephemerella dorothea is a species of spiny crawler mayfly in the family Ephemerellidae, commonly known as the Pale Dun. It is native to North America, with documented occurrence in southwestern and northern Canada, the western United States, and Alaska. The species was first described by Needham in 1908 and includes two recognized subspecies: E. d. dorothea and E. d. infrequens.
Ephemerella subvaria
red quill, Dark Hendrickson
Ephemerella subvaria is a spiny crawler mayfly in the family Ephemerellidae. The species was described by McDunnough in 1931. It is widely distributed in the eastern United States with limited occurrence in the upper Midwest. A 2019 record from Ohio represents the first state documentation, with specimens reared from nymphs collected in March to adult emergence in late April.
Ephemerella tibialis
Little Western Dark Hendrickson
Ephemerella tibialis is a species of spiny crawler mayfly described by McDunnough in 1924. It belongs to the family Ephemerellidae, a group known for their distinctive spiny or tuberculate body ornamentation. The species is found in North America and is recognized among anglers by the common name Little Western Dark Hendrickson, indicating its importance as a model organism for fly-fishing.
Ephemerella verruca
Ephemerella verruca is a species of mayfly in the family Ephemerellidae, described by Allen and Edmunds in 1965. It belongs to the order Ephemeroptera, a group of aquatic insects commonly known as mayflies. The species is recorded from the Nearctic region, specifically North America. Like other members of its genus, it has an aquatic larval stage and terrestrial adult stage.
Eurylophella aestiva
spiny crawler mayfly
Eurylophella aestiva is a species of spiny crawler mayfly in the family Ephemerellidae. It was originally described by McDunnough in 1931 as Ephemerella aestiva. The species is known from North America and belongs to a genus characterized by distinctive spiny projections on the body. Like other members of Ephemerellidae, it exhibits the subimago stage typical of mayflies.
Eurylophella bicolor
bicolor spiny crawler mayfly
Eurylophella bicolor is a species of spiny crawler mayfly in the family Ephemerellidae, originally described as Ephemerella bicolor by Clemens in 1913. The species is distributed across the Nearctic region, particularly in North America. Like other members of the genus Eurylophella, it is associated with freshwater aquatic habitats during its nymphal stages.
Eurylophella funeralis
Eurylophella funeralis is a mayfly species in the family Ephemerellidae, described by McDunnough in 1925. Originally placed in the genus Ephemerella, it was later transferred to Eurylophella. The species is known from the Nearctic region, with records from North America including Vermont. As a member of the order Ephemeroptera, it has an aquatic larval stage and terrestrial adult stage, though specific ecological details for this species remain limited in available literature.
Eurylophella lodi
Eurylophella lodi is a species of spiny crawler mayfly in the family Ephemerellidae, described by Mayo in 1952. It belongs to a genus of mayflies found in North America. The species is known from limited observations, with only two documented occurrences in iNaturalist. Like other members of Ephemerellidae, it is likely associated with freshwater aquatic habitats during its immature stages.
Matriella
Matriella is a genus of mayflies in the family Ephemerellidae, established by Jacobus & McCafferty in 2008. As a relatively recently described genus within the spiny crawler mayfly family, it represents part of the ongoing taxonomic refinement of Ephemerellidae. The genus belongs to the order Ephemeroptera, an ancient group of aquatic insects with aquatic immature stages and terrestrial adults. Specific details about species diversity and biology within Matriella remain limited in available sources.
Matriella teresa
Matriella teresa is a species of spiny crawler mayfly in the family Ephemerellidae, described by Traver in 1934. It belongs to a genus of mayflies found in North America. As with other members of Ephemerellidae, it likely inhabits freshwater streams and rivers during its aquatic nymphal stage.
Penelomax
Penelomax is a genus of spiny crawler mayflies in the family Ephemerellidae, established by Jacobus and McCafferty in 2008. The genus contains a single described species, Penelomax septentrionalis. These mayflies belong to the order Ephemeroptera, a group characterized by short adult lifespans and aquatic larval stages.
Serratella levis
spiny crawler mayfly
Serratella levis is a species of spiny crawler mayfly in the family Ephemerellidae. It was described by Day in 1954 and is found in North America, specifically in the western United States. As a member of the spiny crawler mayflies, it belongs to a group characterized by distinctive morphological features and ecological adaptations to freshwater habitats.
Serratella serrata
little sooty olive
Serratella serrata is a species of spiny crawler mayfly in the family Ephemerellidae. It is found in the south half of Canada and the eastern United States. The species is commonly known as the little sooty olive.
Serratella serratoides
spot-bellied spiny crawler mayfly
Serratella serratoides is a species of spiny crawler mayfly in the family Ephemerellidae. It was originally described as Ephemerella serratoides by McDunnough in 1931 before being transferred to the genus Serratella. The species is known from North America.
Teloganopsis
spiny crawler mayfly
Teloganopsis is a genus of spiny crawler mayflies (Ephemeroptera: Ephemerellidae) comprising approximately 17 described species. The genus was established by Ulmer in 1939 and occurs primarily in the Oriental region. Recent taxonomic work in Thailand has used integrative approaches combining morphology and molecular data (COI and 28S rDNA) to distinguish species, with abdominal color patterns serving as consistent diagnostic characters across developmental stages. Species in this genus have been documented from Thailand and other parts of Asia, though many remain poorly known due to limited collection and study.
Teloganopsis deficiens
little black quill
Teloganopsis deficiens, commonly known as the little black quill, is a species of spiny crawler mayfly in the family Ephemerellidae. It was first described by Morgan in 1911. The species occurs in North America and is part of a mayfly group characterized by crawling behavior and spiny body features. Like other Ephemerellidae, it has an aquatic larval stage and short-lived adult form.
Timpanoga
Timpanoga is a genus of spiny crawler mayflies in the family Ephemerellidae, established by Needham in 1927. The genus contains a single described species, Timpanoga hecuba, commonly known to fly fishermen as the great red quill. This species is significant in angling communities as a model for artificial fly patterns imitating adult and subimago (dun) forms.
Timpanoga hecuba
Great Red Quill
Timpanoga hecuba, commonly known as the Great Red Quill, is a species of mayfly in the family Ephemerellidae. This Nearctic species is significant in fly fishing, where its adult and subimago (dun) forms are imitated by artificial flies. The species was originally described as Ephemerella hecuba by Eaton in 1884 and later transferred to the genus Timpanoga.
Tsalia
Tsalia is a genus of mayflies in the family Ephemerellidae, established by Jacobus and McCafferty in 2008. The genus is poorly documented in scientific literature, with minimal published information on its species composition, biology, or ecology. As a member of Ephemerellidae, it belongs to a family commonly known as spiny-crawler mayflies, though genus-specific traits remain largely uncharacterized.
Tsalia berneri
Tsalia berneri is a species of mayfly in the family Ephemerellidae, originally described as Ephemerella berneri by Allen & Edmunds in 1958. The genus Tsalia was later established to accommodate this and related species based on distinctive morphological features. This Nearctic species is known from limited records in North America. Like other members of Ephemerellidae, it is a small to medium-sized mayfly with aquatic nymphal stages.