Toxoneuron
Say, 1836
Species Guides
1Toxoneuron is a of koinobiont endoparasitoid in the Braconidae. The best-studied , Toxoneuron nigriceps, is a of the tobacco budworm (Heliothis virescens). Members of this genus develop internally within larvae, with a distinctive post-egression feeding phase where larvae feed externally on host tissues before . The genus exhibits sophisticated host manipulation mechanisms including -mediated immune suppression and teratocyte-derived physiological regulation.
Pronunciation
How to pronounce Toxoneuron: //tɒk.soʊˈnʊə.rɒn//
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Habitat
Tobacco agroecosystems and associated agricultural environments where noctuid occur. Laboratory rearing occurs at 29±1°C, 60±10% relative humidity, with 14:10 L:D .
Distribution
United States; historically documented from field studies in Georgia with high rates. Laboratory colonies maintained at Texas A&M University.
Host Associations
- Heliothis virescens - primary tobacco budworm (Lepidoptera: Noctuidae); sole developmentally suitable for T. nigriceps. Koinobiont endoparasitoid development with internal feeding on followed by external tissue feeding.
- Helicoverpa zea - unsuitable (Lepidoptera: Noctuidae); females may oviposit but development unsuccessful
- Manduca sexta - unsuitable (Lepidoptera: Sphingidae); females may oviposit but development unsuccessful
Life Cycle
Koinobiont endoparasitoid development: and larval instars 1-3 develop internally within . First through third instar larvae feed nondestructively on host . Late third instar egresses from host (typically after host to fifth instar), then undergoes obligate external tissue-feeding phase lasting 3-4 hours while anchored to host . occurs within cocoon spun in pupation chamber; larvae require enclosed space (host-formed or artificial chamber) to successfully spin cocoons.
Behavior
Females exhibit -searching mediated by herbivore-induced plant volatiles and direct host cues. Searching time and oviposition effort are disproportionately allocated to suitable hosts over non-hosts. Larvae display distinctive post-egression feeding: after exiting host, the larva anchors its in the egress hole, curves to form a second hole in host , and feeds externally on liquefied host tissues. During this phase, larvae undergo dramatic color change (green to whitish) and near-doubling in size. Larvae can feed on presented tissues from parasitized or unparasitized hosts when normal feeding is disrupted.
Ecological Role
agent of Heliothis virescens, a major pest of tobacco, cotton, soybean, and tomato. Historical field rates estimated at 50-100% in some .
Human Relevance
Studied as a agent for agricultural pest management. Laboratory rearing methods developed including artificial chambers (gelatin capsules and microtube rack wells) for mass production.
Sources and further reading
- BugGuide
- GBIF taxonomy match
- iNaturalist taxon
- NCBI Taxonomy
- Catalogue of Life
- Host regulation and the embryonic development of the endoparasitoid Toxoneuron nigriceps (Hymenoptera: Braconidae)
- Comparison of Searching Behavior of Parasitoid, Toxoneuron nigriceps Vierick, for Three Tobacco Herbivores
- Postegression Feeding Enhances Growth, Survival, and Nutrient Acquisition in the Endoparasitoid Toxoneuron nigriceps (Hymenoptera: Braconidae)
- Host regulation and release of parasitism-specific proteins in the system Toxoneuron nigriceps–Heliothis virescens
- Toxoneuron nigriceps polydnavirus encodes a putative aspartyl protease highly expressed in parasitized host larvae
- Structure and function of the extraembryonic membrane persisting around the larvae of the parasitoid Toxoneuron nigriceps
- Suitability of Various Artificial Pupation Chambers for use by Koinobiont Parasitoid Toxoneuron nigriceps (Hymenoptera: Braconidae)
- Beyond host specificity: the biotechnological exploitation of chitolectin from teratocytes of Toxoneuron nigriceps to control non-permissive hosts
- Identification and Functional Characterization of Toxoneuron nigriceps Ovarian Proteins Involved in the Early Suppression of Host Immune Response
- The probable significance of tracheal tufts in the 8th abdominal segment of Heliothis virescens (F.) on the development of its parasitoid, Toxoneuron nigriceps (Viereck)
- In vitro rearing of Toxoneuron nigriceps (Hymenoptera: Braconidae), a larval endoparasitoid of Heliothis virescens (Lepidoptera: Noctuidae) from early second instar to third instar larvae
- Post-Egression Host Tissue Feeding is Another Strategy of Host Regulation by the Koinobiont Wasp,Toxoneuron nigriceps
- Role of Ovarian Proteins Secreted by Toxoneuron nigriceps (Viereck) (Hymenoptera, Braconidae) in the Early Suppression of Host Immune Response
- Physiological and molecular interaction in the host–parasitoid system Heliothis virescens–Toxoneuron nigriceps: current status and future perspectives
- Characterization of a cDNA encoding a putative chitinase from teratocytes of the endoparasitoidToxoneuron nigriceps